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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1026.61554" ID-GBIF-Dataset="0a2d723c-d292-4f0c-9d00-6cdd176c49a6" ID-PMC="PMC8016817" ID-Pensoft-Pub="1313-2970-1026-45" ID-Pensoft-UUID="2EFE20EFF6705E34BFA20CF497608C4D" ID-PubMed="33850418" ID-ZooBank="A3212DAACEF24A1F93CC1823940C0108" ModsDocID="1313-2970-1026-45" checkinTime="1616697129133" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob &amp; Balke, Michael" docDate="2021" docId="B471A3682E38576C82998670D6AE89EA" docLanguage="en" docName="ZooKeys 1026: 45-67" docOrigin="ZooKeys 1026" docPubDate="2021-03-25" docSource="http://dx.doi.org/10.3897/zookeys.1026.61554" docTitle="Exocelina foja Shaverdo, Surbakti &amp; Balke 2021, sp. nov." docType="treatment" docUuid="464A09DF-E5D6-4D1D-9455-59676F42C60D" docUuidSource="ZooBank" docVersion="6" id="2EFE20EFF6705E34BFA20CF497608C4D" lastPageNumber="45" masterDocId="2EFE20EFF6705E34BFA20CF497608C4D" masterDocTitle="Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="67" masterPageNumber="45" pageNumber="45" updateTime="1668149974149" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaverdo, Helena</mods:namePart>
<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">shaverdo@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Surbakti, Suriani</mods:namePart>
<mods:affiliation>Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sumoked, Bob</mods:namePart>
<mods:affiliation>Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1026</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1026.61554</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1026-45</mods:identifier>
<mods:identifier type="ZooBank">A3212DAACEF24A1F93CC1823940C0108</mods:identifier>
<mods:identifier type="Pensoft-UUID">2EFE20EFF6705E34BFA20CF497608C4D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="180680993" LSID="urn:lsid:zoobank.org:act:464A09DF-E5D6-4D1D-9455-59676F42C60D" httpUri="http://treatment.plazi.org/id/B471A3682E38576C82998670D6AE89EA" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="http://zoobank.org464A09DF-E5D6-4D1D-9455-59676F42C60D/" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina foja" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="foja" status="sp. nov.">Exocelina foja Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">Figures 1</figureCitation>
<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">, 5</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="reference_group">
<paragraph pageId="0" pageNumber="45">
<taxonomicName authority="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke, 2021" authorityName="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudosoppi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pseudosoppi" status="sp. nov.">Exocelina nr. pseudosoppi</taxonomicName>
#7286:
<bibRefCitation DOI="https://doi.org/10.1186/s12862-021-01764-2" author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" refId="B10" refString="Toussaint, EFA, White, LT, Shaverdo, H, Lam, A, Surbakti, S, Panjaitan, R, Sumoked, B, von Rintelen, T, Sagata, K, Balke, M, 2021. [in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology. https://doi.org/10.1186/s12862-021-01764-2" title="[in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology." url="https://doi.org/10.1186/s12862-021-01764-2" year="2021">Toussaint et al. 2021</bibRefCitation>
: figs 3-6.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type locality">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138332" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" elevation="1700" latitude="-2.5718334" location="Foja Mts" longLatPrecision="1" longitude="138.71724" municipality="Sarmi Regency" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
,
<collectingMunicipality>Sarmi Regency</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:21D3633A7BAC2CF23C47FAC8065D398A" country="Indonesia" latitude="-2.5718334" longLatPrecision="1" longitude="138.71724" municipality="Sarmi Regency" name="Foja Mts" stateProvince="Papua Barat">Foja Mts</location>
,
<geoCoordinate degrees="02" direction="south" minutes="34" orientation="latitude" precision="1" seconds="18.6" value="-2.5718334">02°34'18.6&quot;S</geoCoordinate>
,
<geoCoordinate degrees="138" direction="east" minutes="43" orientation="longitude" precision="1" seconds="02.1" value="138.71724">138°43'02.1&quot;E</geoCoordinate>
,
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3062138314" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
male &quot;
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:563E0FA5FD394E70DADF78CFAEE8E80E" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot; (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138330" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MZB, KSP" specimenCount="45" specimenCount-female="19" specimenCount-male="26" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount type="male">26 males</specimenCount>
,
<specimenCount type="female">19 females</specimenCount>
with the same label as the
<typeStatus>holotype</typeStatus>
, three males with additional handwritten labels &quot;creek A&quot;, &quot;creek C&quot; and &quot;creek D&quot; (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3062138333" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="KSP" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">1 male</specimenCount>
,
<specimenCount type="female">6 females</specimenCount>
&quot;
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
(1700A):
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:6838DEAF27438B31517C99228A9E9BA1" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot; (
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3062138324" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">4 males</specimenCount>
,
<specimenCount type="female">5 females</specimenCount>
&quot;
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
(1700B):
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:F1036E62FA48332BFC68FAFCBEFFF844" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot; (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3062138308" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="KSP" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">2 males</specimenCount>
,
<specimenCount type="female">3 females</specimenCount>
&quot;
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
(1700D):
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:38026C8BB9DDE9397DA80D50519986DD" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot; (
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3062138311" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="23" specimenCount-female="10" specimenCount-male="13" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">13 males</specimenCount>
,
<specimenCount type="female">10 females</specimenCount>
&quot;
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:0DAE6C6B38184A1EFFEB2F94B837BA1B" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
,
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,&quot;, &quot;
<geoCoordinate degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot;, two females with additional green text labels
<normalizedToken originalValue="“7357”">&quot;7357&quot;</normalizedToken>
and
<normalizedToken originalValue="“7358”">&quot;7358&quot;</normalizedToken>
(
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3062138327" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="1600" latitude="-2.561006" location="Foja Mountains" longLatPrecision="1" longitude="138.71149" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">2 males</specimenCount>
,
<specimenCount type="female">4 females</specimenCount>
&quot;
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:9BA00C217927AE911629CA60C6BF64CA" country="Indonesia" latitude="-2.561006" longLatPrecision="1" longitude="138.71149" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, river camp,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate degrees="2.561006" direction="south" orientation="latitude" precision="1" value="-2.561006">-2.561006</geoCoordinate>
<geoCoordinate degrees="138.711487" direction="east" orientation="longitude" precision="1" value="138.71149">138.711487</geoCoordinate>
, Sumoked (Pap059)&quot;, one male with an additional handwritten label &quot;forest near bog camp&quot;, the other male with an additional green text label
<normalizedToken originalValue="“7286”">&quot;7286&quot;</normalizedToken>
(
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
:
</emphasis>
Beetle small: TL-H
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.575" metricValueMax="3.85" metricValueMin="3.3" unit="mm" value="3.575" valueMax="3.85" valueMin="3.3">3.30-3.85 mm</quantity>
, TL
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" unit="mm" value="4.0" valueMax="4.3" valueMin="3.7">3.70-4.30 mm</quantity>
, MW
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.80-2.10 mm</quantity>
(
<typeStatus>holotype</typeStatus>
: TL-H
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.85" unit="mm" value="3.85">3.85 mm</quantity>
, TL
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.30 mm</quantity>
,
<collectionCode country="Netherlands" name="Museum Wasmann">MW</collectionCode>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.1" unit="mm" value="2.1">2.10 mm</quantity>
), with oblong-oval habitus (Fig.
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">1</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
:
</emphasis>
Dorsally dark brown to piceous, with paler head and sides of pronotum (Fig.
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">1</figureCitation>
). Head more or less uniformly dark brown to reddish brown, darker around eyes, or slightly paler anteriorly; pronotum dark brown to piceous on disc and distinctly paler (to yellowish red) anteriorly, posteriorly, and especially laterally; dark area on disc sometimes represented just as median band; elytra dark brown to piceous, with reddish brown sutural lines; head appendages and legs yellowish red to reddish brown. Teneral specimens paler.
</paragraph>
<caption doi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45" start="Figures 1–4" startId="F1">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figures 1-4.</emphasis>
Habitus and colouration of
<typeStatus>holotype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="45">1</emphasis>
<taxonomicName authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina foja" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="foja">
<emphasis italics="true" pageId="0" pageNumber="45">Exocelina foja</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="45">2</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. waaf" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="waaf">
<emphasis italics="true" pageId="0" pageNumber="45">E. waaf</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="45">3</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. riberai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="45">E. riberai</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="45">4</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti">
<emphasis italics="true" pageId="0" pageNumber="45">E. apistefti</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
:
</emphasis>
Shiny dorsally, with fine punctation and microreticulation. Head with dense and coarse punctation (spaces between punctures 0-3 times size of punctures), distinctly finer and sparser anteriorly and posteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum with distinctly finer and sparser punctation than on head. Elytra with very sparse and fine punctation, almost invisible. Pronotum and elytra with weakly impressed microreticulation; head with stronger microreticulation. Metaventrite and metacoxa distinctly but weakly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with weak microreticulation, strioles, and fine sparse punctation, coarser and denser on two terminal abdominal ventrites.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Structures</emphasis>
:
</emphasis>
Pronotum with narrow lateral bead. Base of prosternum and neck of prosternal process with ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex medially, with distinct bead and few setae. Abdominal ventrite 6 broadly rounded apically.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Male</emphasis>
:
</emphasis>
Antenna simple. Pro- and mesotarsomeres 1-3 not dilated, narrow. Protarsomere 4 cylindrical, narrow, with medium-sized, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of eleven and posterior row of six short setae (Fig.
<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">5A</figureCitation>
). Median lobe with distinctly discontinuous outline; in lateral view, almost straight, with apex broad, curved downwards, and sharply pointed at tip; in ventral view, with distinct submedian constriction, distal part narrower than proximal one, apex truncate (Fig.
<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">5B, C</figureCitation>
). Paramere with strong notch on dorsal side and subdistal part relatively large and elongate; subdistal setae very few, dense and flattened: three upper longer, thinner, curved at apex and four lower shorter, almost straight, thicker; proximal setae hair-like, numerous, dense, but distinctly more inconspicuous than subdistal ones (Fig.
<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. 5 Exocelina foja sp. nov., holotype 6 E. waaf sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45">5D</figureCitation>
). Abdominal ventrite 6 broadly rounded, with 5-9 lateral striae on each side.
</paragraph>
<caption doi="10.3897/zookeys.1026.61554.figures5-6" httpUri="https://binary.pensoft.net/fig/524446" pageId="0" pageNumber="45" start="Figures 5, 6" startId="F2">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figures 5, 6.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="45">5</emphasis>
<taxonomicName authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina foja" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="foja">
<emphasis italics="true" pageId="0" pageNumber="45">Exocelina foja</emphasis>
</taxonomicName>
sp. nov.,
<typeStatus>holotype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="45">6</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. waaf" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="waaf">
<emphasis italics="true" pageId="0" pageNumber="45">E. waaf</emphasis>
</taxonomicName>
sp. nov.,
<typeStatus>holotype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
right protarsomeres 4 and
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
ventral view
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
median lobe in ventral view
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
median lobe in lateral view
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
right paramere in external view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Female</emphasis>
:
</emphasis>
Pro- and mesotarsi not modified. Abdominal ventrite 6 without striae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="affinities">
<paragraph pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="45">
The species evidently belongs to the
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
</taxonomicName>
group due to the discontinuous outline of its median lobe. Within the group, it can be placed close to
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. oceai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="oceai">
<emphasis italics="true" pageId="0" pageNumber="45">E. oceai</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al., 2012</bibRefCitation>
due to presence of the pronotal lateral bead. Shape of its median lobe is similar to that of
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. pseudosoppi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pseudosoppi">
<emphasis italics="true" pageId="0" pageNumber="45">E. pseudosoppi</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al., 2012</bibRefCitation>
; setation of the paramere is very characteristic.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138325" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" location="Foja Mountains" municipality="This" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
.
<collectingMunicipality>This</collectingMunicipality>
species is known only from
<location LSID="urn:lsid:plazi:treatment:B471A3682E38576C82998670D6AE89EA:BB3F48F11C533ADABE8DABBB08F53A26" country="Indonesia" municipality="This" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, from and near the
<typeStatus>type</typeStatus>
locality (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="habitat">
<paragraph pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="45">The specimens were collected from small, shallow forest creeks.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">The species is named after Foja Mountains. The name is a noun in the nominative singular standing in apposition.</paragraph>
</subSubSection>
</treatment>
</document>
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