188 lines
40 KiB
XML
188 lines
40 KiB
XML
<document id="A46C6D245889483014E12BCCA96347A5" ID-DOI="10.11646/zootaxa.4742.3.1" ID-GBIF-Dataset="84ef451f-d1a1-48f6-871e-f688204fd959" ID-ISSN="1175-5326" ID-Zenodo-Dep="3677869" ID-ZooBank="44ECBE3C-DEA4-4A6B-87D8-D7065591577C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582272866112" checkinUser="plazi" docAuthor="Nunes, Benedito Mendes, Campos, Lourival Dias, Mendonca, Maria Thayane Da Silva, Cunha, Eduardo Victor De Paiva & Fernandes, Jose Antonio Marin" docDate="2020" docId="720587AC3179D91944B26BF1FC4F19F5" docLanguage="en" docName="zootaxa.4742.3.1.pdf" docOrigin="Zootaxa 4742 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hypoxys quadriden s" docType="treatment" docVersion="10" lastPageNumber="407" masterDocId="8E3CFFD4317DD91F44256D6EFF9C1A3E" masterDocTitle="Revision of Hypoxys Amyot & Serville, 1843 stat. rest. (Heteroptera Pentatomidae)" masterLastPageNumber="441" masterPageNumber="401" pageNumber="405" updateTime="1698720852491" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="AFE679FBE82A5F26B27642371B8666EA">Revision of Hypoxys Amyot & Serville, 1843 stat. rest. (Heteroptera Pentatomidae)</mods:title>
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<mods:namePart id="99F0ECE6446A763A971AEFD379A4D10C">Nunes, Benedito Mendes</mods:namePart>
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<mods:namePart id="C3BA2546FFFC622B901476D38F6183BA">Campos, Lourival Dias</mods:namePart>
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<mods:namePart id="28646FC4CE7B5ACEDA4C8576C88DECC9">Mendonca, Maria Thayane Da Silva</mods:namePart>
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<mods:namePart id="66E8D7A49E09890D7D740FFD1C1F9F4E">Cunha, Eduardo Victor De Paiva</mods:namePart>
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<mods:namePart id="DF474DB2318ACD9DD1CFC3A5D012936E">Fernandes, Jose Antonio Marin</mods:namePart>
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<mods:title id="ACACE81A5FDBC8ADB30222B1323CB731">Zootaxa</mods:title>
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<mods:date id="50299A0329C616ADB5113103AEF266C8">2020</mods:date>
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<mods:number id="FC9DBB4F7811A297DFD6C965C62C5055">2020-02-21</mods:number>
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<treatment id="720587AC3179D91944B26BF1FC4F19F5" ID-DOI="http://doi.org/10.5281/zenodo.3684980" ID-GBIF-Taxon="162244891" ID-Zenodo-Dep="3684980" LSID="urn:lsid:plazi:treatment:720587AC3179D91944B26BF1FC4F19F5" httpUri="http://treatment.plazi.org/id/720587AC3179D91944B26BF1FC4F19F5" lastPageId="6" lastPageNumber="407" pageId="4" pageNumber="405">
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<subSubSection id="B2B665313179D91B44B26BF1FE011C84" box="[151,413,1695,1722]" pageId="4" pageNumber="405" type="nomenclature">
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<paragraph id="FA1336BA3179D91B44B26BF1FE011C84" blockId="4.[151,576,1695,1758]" box="[151,413,1695,1722]" pageId="4" pageNumber="405">
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<emphasis id="C8D8EAA83179D91B44B26BF1FE011C84" bold="true" box="[151,413,1695,1722]" pageId="4" pageNumber="405">
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Group
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<taxonomicName id="3DAC4D393179D91B44D46BCEFE011C84" box="[241,413,1695,1722]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="quadriden">
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<emphasis id="C8D8EAA83179D91B44D46BCEFE0F1C84" bold="true" box="[241,403,1695,1722]" italics="true" pageId="4" pageNumber="405">H. quadriden</emphasis>
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s
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="B2B665313179D91B44B26BADFDDC1CE0" box="[151,576,1731,1758]" pageId="4" pageNumber="405" type="description">
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<paragraph id="FA1336BA3179D91B44B26BADFDDC1CE0" blockId="4.[151,576,1695,1758]" box="[151,576,1731,1758]" pageId="4" pageNumber="405">
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(
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<figureCitation id="62972A3F3179D91B44BA6BADFE951CE0" box="[159,265,1731,1758]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="5.[151,250,1912,1937]" captionStartId-1="8.[151,250,1805,1830]" captionStartId-2="10.[151,250,1983,2008]" captionStartId-3="11.[151,250,1608,1633]" captionStartId-4="13.[151,250,1969,1994]" captionStartId-5="14.[151,250,1892,1917]" captionStartId-6="16.[151,250,1746,1771]" captionStartId-7="18.[151,250,1647,1672]" captionTargetBox-0="[225,1364,199,1866]" captionTargetBox-1="[171,1421,437,1746]" captionTargetBox-2="[210,1383,200,1948]" captionTargetBox-3="[162,1424,203,1547]" captionTargetBox-4="[166,1421,496,1904]" captionTargetBox-5="[166,1419,346,1824]" captionTargetBox-6="[169,1410,207,1693]" captionTargetBox-7="[170,1421,201,1594]" captionTargetId-0="figure@5.[214,1374,179,1884]" captionTargetId-1="figure@8.[151,1436,417,1780]" captionTargetId-2="figure@10.[190,1398,181,1957]" captionTargetId-3="figure@11.[151,1436,181,1583]" captionTargetId-4="figure@13.[151,1436,486,1944]" captionTargetId-5="figure@14.[151,1436,334,1866]" captionTargetId-6="figure@16.[151,1436,181,1721]" captionTargetId-7="figure@18.[151,1436,181,1621]" captionTargetPageId-0="5" captionTargetPageId-1="8" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="16" captionTargetPageId-7="18" captionText-0="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." captionText-1="FIGURE 2. H. boerneri. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-2="FIGURE 3. H. claricolor. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data (Scale = 1mm)." captionText-3="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-4="FIGURE 5. H. eburatulus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-5="FIGURE 6. H. necopinatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-6="FIGURE 7. H. offuscatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-7="FIGURE 8. H. trabeculus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi-0="http://doi.org/10.5281/zenodo.3677871" figureDoi-1="http://doi.org/10.5281/zenodo.3677873" figureDoi-2="http://doi.org/10.5281/zenodo.3677875" figureDoi-3="http://doi.org/10.5281/zenodo.3677877" figureDoi-4="http://doi.org/10.5281/zenodo.3677879" figureDoi-5="http://doi.org/10.5281/zenodo.3677881" figureDoi-6="http://doi.org/10.5281/zenodo.3677883" figureDoi-7="http://doi.org/10.5281/zenodo.3677885" httpUri-0="https://zenodo.org/record/3677871/files/figure.png" httpUri-1="https://zenodo.org/record/3677873/files/figure.png" httpUri-2="https://zenodo.org/record/3677875/files/figure.png" httpUri-3="https://zenodo.org/record/3677877/files/figure.png" httpUri-4="https://zenodo.org/record/3677879/files/figure.png" httpUri-5="https://zenodo.org/record/3677881/files/figure.png" httpUri-6="https://zenodo.org/record/3677883/files/figure.png" httpUri-7="https://zenodo.org/record/3677885/files/figure.png" pageId="4" pageNumber="405">Figs. 1–8</figureCitation>
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A–E; 18 A–L, 19 A–D; 21)
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</paragraph>
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</subSubSection>
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<subSubSection id="B2B665313179D91944B26A65FAA21B21" lastPageId="6" lastPageNumber="407" pageId="4" pageNumber="405" type="discussion">
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<paragraph id="FA1336BA3179D91B44B26A65FEB51DC0" blockId="4.[151,1436,1803,2046]" pageId="4" pageNumber="405">
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This group of species is by far the largest in
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<emphasis id="C8D8EAA83179D91B46AB6A62FD721D1B" box="[654,750,1804,1829]" italics="true" pageId="4" pageNumber="405">Hypoxys</emphasis>
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and was treated by
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<bibRefCitation id="9E3D4B4B3179D91B47F56A65FB1E1D18" author="Breddin, G." box="[976,1154,1803,1830]" pageId="4" pageNumber="405" pagination="135 - 153" refId="ref24439" refString="Breddin, G. (1904 a) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1904, 135 - 153." type="journal article" year="1904">Breddin (1904a</bibRefCitation>
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,
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<bibRefCitation id="9E3D4B4B3179D91B40AB6A62FB561D18" author="Breddin, G." box="[1166,1226,1804,1830]" pageId="4" pageNumber="405" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">1907</bibRefCitation>
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). His last paper on this group (1907) brings a curious statement: “Subsequent examinations done on abundant material showed that, the number of species placed under the old species name [
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<taxonomicName id="3DAC4D393179D91B470B6A3DFB981D53" authorityName="Fabricius" authorityYear="1803" box="[814,1028,1875,1901]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="quadridens">
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<emphasis id="C8D8EAA83179D91B470B6A3DFB981D53" box="[814,1028,1875,1901]" italics="true" pageId="4" pageNumber="405">Edessa quadridens</emphasis>
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</taxonomicName>
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] is wide larger than ever suspected, so big, that in the beginning I hardly found the courage to go for the solution of this chaos of close related forms.” More than one hundred years later we still have the same feeling about this group. Even with more than 50 new species in
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<emphasis id="C8D8EAA83179D91B452A6AAEFEF31DE7" box="[271,367,1984,2009]" italics="true" pageId="4" pageNumber="405">Hypoxys</emphasis>
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being recognized so far (in preparation), the sensation that the job is far from be done is still very present.
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</paragraph>
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<caption id="AED366323178D91A44B26A16FCED1DC3" ID-DOI="http://doi.org/10.5281/zenodo.3677871" ID-Zenodo-Dep="3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="5" pageNumber="406" startId="5.[151,250,1912,1937]" targetBox="[225,1364,199,1866]" targetPageId="5">
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<paragraph id="FA1336BA3178D91A44B26A16FCED1DC3" blockId="5.[151,1436,1912,2045]" pageId="5" pageNumber="406">
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<emphasis id="C8D8EAA83178D91A44B26A16FE341DAF" bold="true" box="[151,424,1912,1937]" pageId="5" pageNumber="406">
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FIGURE 1.
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<taxonomicName id="3DAC4D393178D91A453E6A17FE341DAF" box="[283,424,1912,1937]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="quadriden">
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<emphasis id="C8D8EAA83178D91A453E6A17FE341DAF" box="[283,424,1912,1937]" italics="true" pageId="5" pageNumber="406">H. quadriden</emphasis>
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</taxonomicName>
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</emphasis>
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s. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ven-tral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm).
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</paragraph>
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</caption>
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<paragraph id="FA1336BA317BD91944E26DF7FAA21B21" blockId="6.[151,1437,152,971]" pageId="6" pageNumber="407">
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<taxonomicName id="3DAC4D39317BD91944E26DF7FE001A8D" authorityName="Fabricius" authorityYear="1803" box="[199,412,153,179]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="quadridens">
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<emphasis id="C8D8EAA8317BD91944E26DF7FE001A8D" box="[199,412,153,179]" italics="true" pageId="6" pageNumber="407">Edessa quadridens</emphasis>
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</taxonomicName>
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is a name used for almost 150 years to identify a large number of species of
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<emphasis id="C8D8EAA8317BD91941276DF4FAFE1A8D" box="[1282,1378,154,179]" italics="true" pageId="6" pageNumber="407">Hypoxys</emphasis>
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with stripe of pronotum solid black, ventral surface with narrow, transversal, brown lines, and other characteristics presented below. This group includes:
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<emphasis id="C8D8EAA8317BD919463A6D8FFDDA1B21" italics="true" pageId="6" pageNumber="407">
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||
<taxonomicName id="3DAC4D39317BD919463A6D8FFD241AC5" box="[543,696,224,251]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="quadriden">
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<emphasis id="C8D8EAA8317BD919463A6D8FFD241AC5" bold="true" box="[543,696,224,251]" italics="true" pageId="6" pageNumber="407">H. quadriden</emphasis>
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</taxonomicName>
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s,
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<taxonomicName id="3DAC4D39317BD91946F46D8CFCC91AC5" authorityName="Nunes & Campos & Mendonca & Cunha & Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[721,853,225,251]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="boerneri">H. boerneri</taxonomicName>
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,
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<taxonomicName id="3DAC4D39317BD91947456D8CFC6E1AC5" authorityName="Nunes & Campos & Mendonca & Cunha & Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[864,1010,225,251]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="claricolor">H. claricolor</taxonomicName>
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,
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<taxonomicName id="3DAC4D39317BD91947D96D8CFBE91AC5" authorityName="Nunes & Campos & Mendonca & Cunha & Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[1020,1141,225,251]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="dolosus">H. dolosus</taxonomicName>
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,
|
||
<taxonomicName id="3DAC4D39317BD91940A56D8CFA811AC5" authorityName="Nunes & Campos & Mendonca & Cunha & Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[1152,1309,225,251]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="eburatulus">H. eburatulus</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DAC4D39317BD91941026D8CFF4A1B21" authorityName="Nunes & Campos & Mendonca & Cunha & Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="necopinatus">H. necopinatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DAC4D39317BD91944C76C68FEEA1B21" authorityName="Nunes & Campos & Mendonca & Cunha & Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[226,374,261,287]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="offuscatus">H. offuscatus</taxonomicName>
|
||
and
|
||
<taxonomicName id="3DAC4D39317BD919458B6C68FDDA1B21" authorityName="Nunes & Campos & Mendonca & Cunha & Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[430,582,261,287]" class="Insecta" family="Pentatomidae" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="trabeculus">H. trabeculus</taxonomicName>
|
||
</emphasis>
|
||
, and a lot of new species that will be described in upcoming works.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2B66531317BD91944E26C46FED71BED" pageId="6" pageNumber="407" type="diagnosis">
|
||
<paragraph id="FA1336BA317BD91944E26C46FED71BED" blockId="6.[151,1437,152,971]" pageId="6" pageNumber="407">
|
||
<emphasis id="C8D8EAA8317BD91944E26C46FEA61B7D" bold="true" box="[199,314,296,323]" pageId="6" pageNumber="407">Diagnosis</emphasis>
|
||
. Second antennomere longer than third. Posterior part of the pronotum with a transverse solid black stripe (
|
||
<figureCitation id="62972A3F317BD91944C06C23FE321B59" box="[229,430,333,359]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18A,C,G,I,K</figureCitation>
|
||
). Anterior part of scutellum with part of the punctures on dark spots (
|
||
<figureCitation id="62972A3F317BD919409D6C23FABD1B59" box="[1208,1313,333,359]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18A</figureCitation>
|
||
). Connexivum with anterior and posterior black spots on each segment extending ventrally (
|
||
<figureCitation id="62972A3F317BD91940676C1FFB421BB5" box="[1090,1246,369,395]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18A,C,E</figureCitation>
|
||
). Body ventrally yellow to greenish yellow with dark brown transverse stripes on intersegmental areas and pseudosutures (
|
||
<figureCitation id="62972A3F317BD919414A6CFBFEA31BED" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18B,D,F,H,J,L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2B66531317BD91944E26CB3FC4F19F5" pageId="6" pageNumber="407" type="description">
|
||
<paragraph id="FA1336BA317BD91944E26CB3FBCD1825" blockId="6.[151,1437,152,971]" pageId="6" pageNumber="407">Head. Apex of mandibular plates concolorous, sometimes with dark spot; lateral margin sometimes with yellow spot. Antennae brown with setae more densely distributed in antennomeres IV and V.</paragraph>
|
||
<paragraph id="FA1336BA317BD91944E26F4BFADB18F1" blockId="6.[151,1437,152,971]" pageId="6" pageNumber="407">
|
||
Thorax. Pronotum with punctures black to brown. Scutellum with punctures black or dark brown (
|
||
<figureCitation id="62972A3F317BD919414A6F4BFEDF185D" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18A,C,E,G,I,K</figureCitation>
|
||
); anterior margin with a yellow stripe; apex not punctured (
|
||
<figureCitation id="62972A3F317BD91947C26F27FBCD185D" box="[999,1105,585,611]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18A</figureCitation>
|
||
). Black spot of costal margin reaching between 1/3 and 1/4 of corial extension. Propleura with punctures brown, except in a rounded area adjacent to coxa (proepisternun and proepimeron). Evaporatorium concolorous and slightly whitish (
|
||
<figureCitation id="62972A3F317BD91940B86FFFFA9F1895" box="[1181,1283,657,683]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18B</figureCitation>
|
||
). Metasternal process (
|
||
<figureCitation id="62972A3F317BD91944DF6FDBFEEA18F1" box="[250,374,693,719]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="5.[151,250,1912,1937]" captionStartId-1="8.[151,250,1805,1830]" captionStartId-2="10.[151,250,1983,2008]" captionStartId-3="11.[151,250,1608,1633]" captionStartId-4="13.[151,250,1969,1994]" captionStartId-5="14.[151,250,1892,1917]" captionStartId-6="16.[151,250,1746,1771]" captionStartId-7="18.[151,250,1647,1672]" captionTargetBox-0="[225,1364,199,1866]" captionTargetBox-1="[171,1421,437,1746]" captionTargetBox-2="[210,1383,200,1948]" captionTargetBox-3="[162,1424,203,1547]" captionTargetBox-4="[166,1421,496,1904]" captionTargetBox-5="[166,1419,346,1824]" captionTargetBox-6="[169,1410,207,1693]" captionTargetBox-7="[170,1421,201,1594]" captionTargetId-0="figure@5.[214,1374,179,1884]" captionTargetId-1="figure@8.[151,1436,417,1780]" captionTargetId-2="figure@10.[190,1398,181,1957]" captionTargetId-3="figure@11.[151,1436,181,1583]" captionTargetId-4="figure@13.[151,1436,486,1944]" captionTargetId-5="figure@14.[151,1436,334,1866]" captionTargetId-6="figure@16.[151,1436,181,1721]" captionTargetId-7="figure@18.[151,1436,181,1621]" captionTargetPageId-0="5" captionTargetPageId-1="8" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="16" captionTargetPageId-7="18" captionText-0="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." captionText-1="FIGURE 2. H. boerneri. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-2="FIGURE 3. H. claricolor. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data (Scale = 1mm)." captionText-3="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-4="FIGURE 5. H. eburatulus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-5="FIGURE 6. H. necopinatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-6="FIGURE 7. H. offuscatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-7="FIGURE 8. H. trabeculus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi-0="http://doi.org/10.5281/zenodo.3677871" figureDoi-1="http://doi.org/10.5281/zenodo.3677873" figureDoi-2="http://doi.org/10.5281/zenodo.3677875" figureDoi-3="http://doi.org/10.5281/zenodo.3677877" figureDoi-4="http://doi.org/10.5281/zenodo.3677879" figureDoi-5="http://doi.org/10.5281/zenodo.3677881" figureDoi-6="http://doi.org/10.5281/zenodo.3677883" figureDoi-7="http://doi.org/10.5281/zenodo.3677885" httpUri-0="https://zenodo.org/record/3677871/files/figure.png" httpUri-1="https://zenodo.org/record/3677873/files/figure.png" httpUri-2="https://zenodo.org/record/3677875/files/figure.png" httpUri-3="https://zenodo.org/record/3677877/files/figure.png" httpUri-4="https://zenodo.org/record/3677879/files/figure.png" httpUri-5="https://zenodo.org/record/3677881/files/figure.png" httpUri-6="https://zenodo.org/record/3677883/files/figure.png" httpUri-7="https://zenodo.org/record/3677885/files/figure.png" pageId="6" pageNumber="407">Figs. 1–8E</figureCitation>
|
||
) with anterior bifurcation receiving fourth and at least half of the third rostral segment.
|
||
</paragraph>
|
||
<paragraph id="FA1336BA317BD91944E26FB7FC651905" blockId="6.[151,1437,152,971]" pageId="6" pageNumber="407">
|
||
Abdomen. Dorsal surface brown. Connexivum green with punctures concentrated in two excavated areas separated by median yellow callous (
|
||
<figureCitation id="62972A3F317BD91946616F93FD331929" box="[580,687,765,791]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18A</figureCitation>
|
||
). End of pseudosutures with rounded dark spots (
|
||
<figureCitation id="62972A3F317BD91940CB6F93FACA1929" box="[1262,1366,765,791]" captionStart="FIGURE 18" captionStartId="34.[151,250,1983,2008]" captionTargetBox="[244,1330,184,1951]" captionTargetId="figure@34.[190,1398,181,1957]" captionTargetPageId="34" captionText="FIGURE 18. Dorsal and ventral view. H. quadridens A–B; H. boerneri C–D; H. claricolor E–F; H. dolosus G–H; H. eburatulus I–J; H. necopinatus K–L (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677905" httpUri="https://zenodo.org/record/3677905/files/figure.png" pageId="6" pageNumber="407">Fig. 18B</figureCitation>
|
||
). One trichobothrium in line with spiracles and the other located laterad of this line.
|
||
</paragraph>
|
||
<paragraph id="FA1336BA317BD91944E26E2BFCAF1999" blockId="6.[151,1437,152,971]" pageId="6" pageNumber="407">
|
||
Male (
|
||
<figureCitation id="62972A3F317BD91945366E2BFEE11961" box="[275,381,837,863]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="5.[151,250,1912,1937]" captionStartId-1="8.[151,250,1805,1830]" captionStartId-2="10.[151,250,1983,2008]" captionStartId-3="11.[151,250,1608,1633]" captionStartId-4="13.[151,250,1969,1994]" captionStartId-5="14.[151,250,1892,1917]" captionStartId-6="16.[151,250,1746,1771]" captionStartId-7="18.[151,250,1647,1672]" captionTargetBox-0="[225,1364,199,1866]" captionTargetBox-1="[171,1421,437,1746]" captionTargetBox-2="[210,1383,200,1948]" captionTargetBox-3="[162,1424,203,1547]" captionTargetBox-4="[166,1421,496,1904]" captionTargetBox-5="[166,1419,346,1824]" captionTargetBox-6="[169,1410,207,1693]" captionTargetBox-7="[170,1421,201,1594]" captionTargetId-0="figure@5.[214,1374,179,1884]" captionTargetId-1="figure@8.[151,1436,417,1780]" captionTargetId-2="figure@10.[190,1398,181,1957]" captionTargetId-3="figure@11.[151,1436,181,1583]" captionTargetId-4="figure@13.[151,1436,486,1944]" captionTargetId-5="figure@14.[151,1436,334,1866]" captionTargetId-6="figure@16.[151,1436,181,1721]" captionTargetId-7="figure@18.[151,1436,181,1621]" captionTargetPageId-0="5" captionTargetPageId-1="8" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="16" captionTargetPageId-7="18" captionText-0="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." captionText-1="FIGURE 2. H. boerneri. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-2="FIGURE 3. H. claricolor. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data (Scale = 1mm)." captionText-3="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-4="FIGURE 5. H. eburatulus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-5="FIGURE 6. H. necopinatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-6="FIGURE 7. H. offuscatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-7="FIGURE 8. H. trabeculus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi-0="http://doi.org/10.5281/zenodo.3677871" figureDoi-1="http://doi.org/10.5281/zenodo.3677873" figureDoi-2="http://doi.org/10.5281/zenodo.3677875" figureDoi-3="http://doi.org/10.5281/zenodo.3677877" figureDoi-4="http://doi.org/10.5281/zenodo.3677879" figureDoi-5="http://doi.org/10.5281/zenodo.3677881" figureDoi-6="http://doi.org/10.5281/zenodo.3677883" figureDoi-7="http://doi.org/10.5281/zenodo.3677885" httpUri-0="https://zenodo.org/record/3677871/files/figure.png" httpUri-1="https://zenodo.org/record/3677873/files/figure.png" httpUri-2="https://zenodo.org/record/3677875/files/figure.png" httpUri-3="https://zenodo.org/record/3677877/files/figure.png" httpUri-4="https://zenodo.org/record/3677879/files/figure.png" httpUri-5="https://zenodo.org/record/3677881/files/figure.png" httpUri-6="https://zenodo.org/record/3677883/files/figure.png" httpUri-7="https://zenodo.org/record/3677885/files/figure.png" pageId="6" pageNumber="407">Figs. 1–8</figureCitation>
|
||
A–C). Pygophore with distal part of dorsal surface brown. Superior process of genital cup black. Paramere with distal margin black. Proctiger with tufts of setae covering lateral excavations; posterior face laterally expanded usually covering part of the tuft of setae.
|
||
</paragraph>
|
||
<paragraph id="FA1336BA317BD91944B26EDFFC4F19F5" blockId="6.[151,1437,152,971]" box="[151,979,945,971]" pageId="6" pageNumber="407">
|
||
Female (
|
||
<figureCitation id="62972A3F317BD91944DC6EDFFEE619F5" box="[249,378,945,971]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="5.[151,250,1912,1937]" captionStartId-1="8.[151,250,1805,1830]" captionStartId-2="10.[151,250,1983,2008]" captionStartId-3="11.[151,250,1608,1633]" captionStartId-4="13.[151,250,1969,1994]" captionStartId-5="14.[151,250,1892,1917]" captionStartId-6="16.[151,250,1746,1771]" captionStartId-7="18.[151,250,1647,1672]" captionTargetBox-0="[225,1364,199,1866]" captionTargetBox-1="[171,1421,437,1746]" captionTargetBox-2="[210,1383,200,1948]" captionTargetBox-3="[162,1424,203,1547]" captionTargetBox-4="[166,1421,496,1904]" captionTargetBox-5="[166,1419,346,1824]" captionTargetBox-6="[169,1410,207,1693]" captionTargetBox-7="[170,1421,201,1594]" captionTargetId-0="figure@5.[214,1374,179,1884]" captionTargetId-1="figure@8.[151,1436,417,1780]" captionTargetId-2="figure@10.[190,1398,181,1957]" captionTargetId-3="figure@11.[151,1436,181,1583]" captionTargetId-4="figure@13.[151,1436,486,1944]" captionTargetId-5="figure@14.[151,1436,334,1866]" captionTargetId-6="figure@16.[151,1436,181,1721]" captionTargetId-7="figure@18.[151,1436,181,1621]" captionTargetPageId-0="5" captionTargetPageId-1="8" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="16" captionTargetPageId-7="18" captionText-0="FIGURE 1. H. quadridens. A–C Male, pygophore; A—dorsal view; B–posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." captionText-1="FIGURE 2. H. boerneri. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-2="FIGURE 3. H. claricolor. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data (Scale = 1mm)." captionText-3="FIGURE 4. H. dolosus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-4="FIGURE 5. H. eburatulus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-5="FIGURE 6. H. necopinatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-6="FIGURE 7. H. offuscatus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." captionText-7="FIGURE 8. H. trabeculus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi-0="http://doi.org/10.5281/zenodo.3677871" figureDoi-1="http://doi.org/10.5281/zenodo.3677873" figureDoi-2="http://doi.org/10.5281/zenodo.3677875" figureDoi-3="http://doi.org/10.5281/zenodo.3677877" figureDoi-4="http://doi.org/10.5281/zenodo.3677879" figureDoi-5="http://doi.org/10.5281/zenodo.3677881" figureDoi-6="http://doi.org/10.5281/zenodo.3677883" figureDoi-7="http://doi.org/10.5281/zenodo.3677885" httpUri-0="https://zenodo.org/record/3677871/files/figure.png" httpUri-1="https://zenodo.org/record/3677873/files/figure.png" httpUri-2="https://zenodo.org/record/3677875/files/figure.png" httpUri-3="https://zenodo.org/record/3677877/files/figure.png" httpUri-4="https://zenodo.org/record/3677879/files/figure.png" httpUri-5="https://zenodo.org/record/3677881/files/figure.png" httpUri-6="https://zenodo.org/record/3677883/files/figure.png" httpUri-7="https://zenodo.org/record/3677885/files/figure.png" pageId="6" pageNumber="407">Figs. 1–8D</figureCitation>
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). Gonocoxites 9 small, narrow, distal margin straight.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |