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<document id="6C1AD53CDA89FAE7BF5A0A434FB9AB8F" ID-DOI="10.5281/zenodo.187023" ID-GBIF-Dataset="f47f2de5-ebee-4164-92d8-e3542a1cd334" ID-ISSN="1175-5326" ID-Zenodo-Dep="187023" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461060654237" checkinUser="plazi" docAuthor="Hsueh, Pan-Wen &amp; Kuo, Chia-Ming" docDate="2009" docId="3B3687EBBF11D15CFF35FBDC1EB15886" docLanguage="en" docName="zt02067p061.pdf" docOrigin="Zootaxa 2067" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Phascolosoma (Phascolosoma) arcuatum Gray" docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="C70FFF93BF16D15BFFA2FFD91D0C5F08" masterDocTitle="New records of sipunculan worms from Taiwan" masterLastPageNumber="61" masterPageNumber="51" pageNumber="58" updateTime="1698582104263" updateUser="plazi">
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<mods:title id="05E2E0243D655C879FD080D9AC1683F0">New records of sipunculan worms from Taiwan</mods:title>
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<mods:namePart id="B5417943C4EE217551BE5AA8F2631A26">Hsueh, Pan-Wen</mods:namePart>
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<mods:namePart id="145D4649E3DAD99693E90FBEBBAFA32D">Kuo, Chia-Ming</mods:namePart>
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<mods:date id="6F21B34865FF5B1CF025F2BA1525889A">2009</mods:date>
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<mods:number id="EB90189150869F35846A879EC56DE05A">2067</mods:number>
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<treatment id="3B3687EBBF11D15CFF35FBDC1EB15886" ID-DOI="http://doi.org/10.5281/zenodo.5671997" ID-GBIF-Taxon="119614116" ID-Zenodo-Dep="5671997" LSID="urn:lsid:plazi:treatment:3B3687EBBF11D15CFF35FBDC1EB15886" httpUri="http://treatment.plazi.org/id/3B3687EBBF11D15CFF35FBDC1EB15886" lastPageNumber="58" pageId="7" pageNumber="58">
<subSubSection id="FB856576BF11D15CFF35FBDC1C285B4A" pageId="7" pageNumber="58" type="nomenclature">
<paragraph id="B32036FDBF11D15CFF35FBDC1FEA5B17" blockId="7.[151,742,1029,1090]" box="[151,742,1029,1055]" pageId="7" pageNumber="58">
<heading id="E8688191BF11D15CFF35FBDC1FEA5B17" bold="true" box="[151,742,1029,1055]" fontSize="11" level="1" pageId="7" pageNumber="58" reason="1">
<taxonomicName id="749F4D7EBF11D15CFF35FBDC1FEA5B17" ID-CoL="7WCMR" authority="Gray" authorityName="Gray" box="[151,742,1029,1055]" class="Phascolosomatidea" family="Phascolosomatidae" genus="Phascolosoma" kingdom="Animalia" order="Phascolosomatiformes" pageId="7" pageNumber="58" phylum="Sipuncula" rank="species" species="arcuatum" subGenus="Phascolosoma">
<emphasis id="81EBEAEFBF11D15CFF35FBDC1FEA5B17" bold="true" box="[151,742,1029,1055]" pageId="7" pageNumber="58">
<emphasis id="81EBEAEFBF11D15CFF35FBDC1C495B17" bold="true" box="[151,325,1029,1055]" italics="true" pageId="7" pageNumber="58">Phascolosoma</emphasis>
(
<emphasis id="81EBEAEFBF11D15CFEF4FBDC1F085B17" bold="true" box="[342,516,1029,1055]" italics="true" pageId="7" pageNumber="58">Phascolosoma</emphasis>
)
<emphasis id="81EBEAEFBF11D15CFDB7FBDC1F865B17" bold="true" box="[533,650,1029,1055]" italics="true" pageId="7" pageNumber="58">arcuatum</emphasis>
(Gray)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="B32036FDBF11D15CFF35FBF11C285B4A" blockId="7.[151,742,1029,1090]" box="[151,292,1064,1090]" pageId="7" pageNumber="58">
<figureCitation id="2BA42A78BF11D15CFF35FBF11DEC5B4A" box="[151,224,1064,1090]" captionStart="FIGURE 2. A" captionStartId="3.[151,256,1873,1897]" captionTargetBox="[434,1154,185,1843]" captionTargetId="figure@3.[434,1156,185,1843]" captionTargetPageId="3" captionText="FIGURE 2. A, Sipunculus robustus Keferstein. B, Siphonosoma funafuti (Shipley). C, Siphonosoma vastum (Selenka and Bülow). D, Golfingia margaritacea margaritacea (Sars). E, Golfingia muricaudata (Southern, 1913). F, Thysanocardia nigra (Ikeda). Scalebars: A-E, 10 mm; F, 2.5 mm. G, Themiste minor minor (Ikeda), H, Phascolosoma arcuatum (Gray). I, Phascolosoma glabrum glabrum (Sluiter). J, Aspidosiphon (Aspidosiphon) muelleri Diesing. K, Aspidosiphon coyi de Quatrefages. Scalebars: G, 5 mm; H J, 10 mm; K, 2.5 mm." httpUri="https://zenodo.org/record/187025/files/figure.png" pageId="7" pageNumber="58">Figs 2</figureCitation>
H, 3C
</paragraph>
</subSubSection>
<subSubSection id="FB856576BF11D15CFF35FBAC18675B84" box="[151,1387,1141,1164]" pageId="7" pageNumber="58" type="reference_group">
<paragraph id="B32036FDBF11D15CFF35FBAC18675B84" blockId="7.[151,1387,1141,1164]" box="[151,1387,1141,1164]" pageId="7" pageNumber="58">
<treatmentCitationGroup id="938F11D3BF11D15CFF35FBAC18675B84" box="[151,1387,1141,1164]" pageId="7" pageNumber="58">
<taxonomicName id="749F4D7EBF11D15CFF35FBAC1E195B84" ID-CoL="7WCMR" authority="Cutler, 1994: 163" authorityName="Cutler" authorityPageNumber="163" authorityYear="1994" box="[151,789,1141,1164]" class="Phascolosomatidea" family="Phascolosomatidae" genus="Phascolosoma" kingdom="Animalia" order="Phascolosomatiformes" pageId="7" pageNumber="58" phylum="Sipuncula" rank="species" species="arcuatum" subGenus="Phascolosoma">
<emphasis id="81EBEAEFBF11D15CFF35FBAC1C3E5B84" box="[151,306,1141,1164]" italics="true" pageId="7" pageNumber="58">Phascolosoma</emphasis>
(
<emphasis id="81EBEAEFBF11D15CFEE0FBAC1CD25B84" box="[322,478,1141,1164]" italics="true" pageId="7" pageNumber="58">Phascolosoma</emphasis>
)
<emphasis id="81EBEAEFBF11D15CFE4FFBAC1F5E5B84" box="[493,594,1141,1164]" italics="true" pageId="7" pageNumber="58">arcuatum</emphasis>
<treatmentCitation id="323E10ECBF11D15CFDFBFBAC1E195B84" author="Cutler" box="[601,789,1141,1164]" page="163" pageId="7" pageNumber="58" year="1994">
<bibRefCitation id="D70E4B0CBF11D15CFDFBFBAC1FED5B84" author="Cutler" box="[601,737,1141,1164]" pageId="7" pageNumber="61" refString="Cutler, E. B. (1994) The Sipuncula: Their Systematics, Biology and Evolution. New York: Cornell University. 453 pp." type="book" year="1994">Cutler, 1994</bibRefCitation>
: 163
</treatmentCitation>
</taxonomicName>
, 167, 170, 172, 176, fig. 49D (for complete synonymy).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="FB856576BF11D15CFF35FB621EDF59AE" pageId="7" pageNumber="58" type="materials_examined">
<paragraph id="B32036FDBF11D15CFF35FB621EDF59AE" blockId="7.[151,1437,1211,1934]" pageId="7" pageNumber="58">
<emphasis id="81EBEAEFBF11D15CFF35FB621C995BDD" bold="true" box="[151,405,1211,1237]" pageId="7" pageNumber="58">Material examined.</emphasis>
Nine specimens (NMNS-5928-012), Kaomei tidal flat (
<geoCoordinate id="D6AB503ABF11D15CFBE4FB6519D35BDE" box="[1094,1247,1212,1238]" direction="north" orientation="latitude" pageId="7" pageNumber="58" precision="9" value="24.322334">24°19.34' N</geoCoordinate>
,
<geoCoordinate id="D6AB503ABF11D15CFB4EFB6518825BDE" box="[1260,1422,1212,1238]" direction="east" orientation="longitude" pageId="7" pageNumber="58" precision="9" value="120.551834">120°33.11' E</geoCoordinate>
), Taichung County, high intertidal sandy mud bottom, coll. P.-W. Hsueh,
<date id="C721103DBF11D15CFC49FB3B19DD5BF4" box="[1003,1233,1250,1276]" pageId="7" pageNumber="58" value="1992-12-22">22 December 1992</date>
; three specimens (NMNS-5928-013), Kaomei tidal flat (
<geoCoordinate id="D6AB503ABF11D15CFDCFFAD01E0F5A2B" box="[621,771,1289,1315]" direction="north" orientation="latitude" pageId="7" pageNumber="58" precision="9" value="24.322334">24°19.34' N</geoCoordinate>
,
<geoCoordinate id="D6AB503ABF11D15CFCADFAD01EA35A2B" box="[783,943,1289,1315]" direction="east" orientation="longitude" pageId="7" pageNumber="58" precision="9" value="120.551834">120°33.11' E</geoCoordinate>
), Taichung County, intertidal sandy mud bottom, coll. P.-W. Hsueh,
<date id="C721103DBF11D15CFE7EFAE91F845A42" box="[476,648,1328,1354]" pageId="7" pageNumber="58" value="1999-04-25">25 April 1999</date>
;
<specimenCount id="A599FD74BF11D15CFD37FAE91E245A42" box="[661,808,1328,1354]" pageId="7" pageNumber="58" type="generic">4 specimens</specimenCount>
(NMNS-5928-014), Kaomei tidal flat (
<geoCoordinate id="D6AB503ABF11D15CFAA1FAE9189B5A42" box="[1283,1431,1328,1354]" direction="north" orientation="latitude" pageId="7" pageNumber="58" precision="9" value="24.322334">24°19.34' N</geoCoordinate>
,
<geoCoordinate id="D6AB503ABF11D15CFF35FA8F1C3A5A78" box="[151,310,1366,1392]" direction="east" orientation="longitude" pageId="7" pageNumber="58" precision="9" value="120.551834">120°33.11' E</geoCoordinate>
), Taichung County, high intertidal sandy mud bottom, coll. P.-W. Hsueh,
<date id="C721103DBF11D15CFB12FA8F18515A78" box="[1200,1373,1366,1392]" pageId="7" pageNumber="58" value="2003-03-08">8 March 2003</date>
; One specimen (NMNS-5928-015), Hambao tidal flat (
<geoCoordinate id="D6AB503ABF11D15CFCAAFAA41EA85A9F" box="[776,932,1405,1431]" direction="north" orientation="latitude" pageId="7" pageNumber="58" precision="9" value="24.003834">24°00.23' N</geoCoordinate>
,
<geoCoordinate id="D6AB503ABF11D15CFC11FAA419575A9F" box="[947,1115,1405,1431]" direction="east" orientation="longitude" pageId="7" pageNumber="58" precision="9" value="120.351166">120°21.07' E</geoCoordinate>
), Changhua County, high intertidal sandy mud bottom, coll. H.-T. Hung,
<date id="C721103DBF11D15CFD6FFA7D1E675AB6" box="[717,875,1444,1470]" pageId="7" pageNumber="58" value="2001-07-11">11 July 2001</date>
; one specimen (NMNS-5928-016), Shenkuang tidal flat (
<geoCoordinate id="D6AB503ABF11D15CFEA8FA131C975AEC" box="[266,411,1482,1508]" direction="north" orientation="latitude" pageId="7" pageNumber="58" precision="9" value="24.158333">24°09.50' N</geoCoordinate>
,
<geoCoordinate id="D6AB503ABF11D15CFE0AFA131F4F5AEC" box="[424,579,1482,1508]" direction="east" orientation="longitude" pageId="7" pageNumber="58" precision="9" value="120.45533">120°27.32' E</geoCoordinate>
), Changhua County, high intertidal sandy mud bottom, coll. H.-T. Hung,
<date id="C721103DBF11D15CFF35FA281C3E5903" box="[151,306,1521,1547]" pageId="7" pageNumber="58" value="2001-07-11">11 July 2001</date>
;
<specimenCount id="A599FD74BF11D15CFE9DFA281CDC5903" box="[319,464,1521,1547]" pageId="7" pageNumber="58" type="generic">7 specimens</specimenCount>
(NMNS-5928-017), Bajhang tidal flat (
<geoCoordinate id="D6AB503ABF11D15CFC05FA2819355903" box="[935,1081,1521,1547]" direction="north" orientation="latitude" pageId="7" pageNumber="58" precision="9" value="24.003834">24°00.23' N</geoCoordinate>
,
<geoCoordinate id="D6AB503ABF11D15CFBE4FA2819EE5903" box="[1094,1250,1521,1547]" direction="east" orientation="longitude" pageId="7" pageNumber="58" precision="9" value="120.351166">120°21.07' E</geoCoordinate>
), Chiai County, high intertidal sandy mud bottom, coll. H.-T. Hung,
<date id="C721103DBF11D15CFCB8F9C11EEB593A" box="[794,999,1560,1586]" pageId="7" pageNumber="58" value="2007-01-17">17 January 2007</date>
; One specimen (NMNS-5928-018), Fubao tidal flat (
<geoCoordinate id="D6AB503ABF11D15CFEF9F9E71CE05950" box="[347,492,1598,1624]" direction="north" orientation="latitude" pageId="7" pageNumber="58" precision="9" value="23.988">23°59.28' N</geoCoordinate>
,
<geoCoordinate id="D6AB503ABF11D15CFE5BF9E71F985950" box="[505,660,1598,1624]" direction="east" orientation="longitude" pageId="7" pageNumber="58" precision="9" value="120.35083">120°21.05' E</geoCoordinate>
), Changhua County, high intertidal sandy mud bottom, coll. H.-T. Hung,
<date id="C721103DBF11D15CFF4AF9BC1C895977" box="[232,389,1637,1663]" pageId="7" pageNumber="58" value="2001-07-11">11 July 2001</date>
;
<specimenCount id="A599FD74BF11D15CFE33F9BC1F2F5977" box="[401,547,1637,1663]" pageId="7" pageNumber="58" type="generic">5 specimens</specimenCount>
(NMNS-3636-058), Yunsen (
<geoCoordinate id="D6AB503ABF11D15CFC24F9BC19155977" box="[902,1049,1637,1663]" direction="north" orientation="latitude" pageId="7" pageNumber="58" precision="9" value="24.489">24°29.34' N</geoCoordinate>
,
<geoCoordinate id="D6AB503ABF11D15CFB84F9BC19CE5977" box="[1062,1218,1637,1663]" direction="east" orientation="longitude" pageId="7" pageNumber="58" precision="9" value="118.540665">118°32.44' E</geoCoordinate>
), Kinmen County, high intertidal sandy mud bottom, coll. H.-T. Hung,
<date id="C721103DBF11D15CFD59F9551EDC59AE" box="[763,976,1676,1702]" pageId="7" pageNumber="58" value="2001-02-21">21 February 2001</date>
.
</paragraph>
</subSubSection>
<subSubSection id="FB856576BF11D15CFF64F96B1EB15886" pageId="7" pageNumber="58" type="discussion">
<paragraph id="B32036FDBF11D15CFF64F96B1EB15886" blockId="7.[151,1437,1211,1934]" pageId="7" pageNumber="58">
<emphasis id="81EBEAEFBF11D15CFF64F96B1C4C59C4" bold="true" box="[198,320,1714,1740]" pageId="7" pageNumber="58">Remarks.</emphasis>
The external morphology of the present specimens agrees well with descriptions of the species in
<bibRefCitation id="D70E4B0CBF11D15CFF14F9001C5559FB" author="Cutler" box="[182,345,1753,1779]" pageId="7" pageNumber="61" refString="Cutler, E. B. (1994) The Sipuncula: Their Systematics, Biology and Evolution. New York: Cornell University. 453 pp." type="book" year="1994">Cutler (1994)</bibRefCitation>
; the species has large and dark papillae set off from the skin, the introvert is up to twice the trunk length, there are more than 100 complete and incomplete rings of hooks (about 4070 um tall), basally with greatly expanded clear streak and it is without secondary teeth, warts, or toes (
<bibRefCitation id="D70E4B0CBF11D15CFB2FF8FF1CD2586F" author="Cutler" pageId="7" pageNumber="61" refString="Cutler, E. B. (1994) The Sipuncula: Their Systematics, Biology and Evolution. New York: Cornell University. 453 pp." type="book" year="1994">
Cutler, 1994: 170, fig. 49D; present study,
<figureCitation id="2BA42A78BF11D15CFE20F8941CCB586F" box="[386,455,1869,1895]" captionStart="FIGURE 3. A" captionStartId="8.[151,256,1175,1199]" captionTargetBox="[456,1134,198,1137]" captionTargetId="figure@8.[453,1134,197,1149]" captionTargetPageId="8" captionText="FIGURE 3. A, Introvert hooks of Siphonosoma vastum (Selenka and Bülow). B, Anterior close-up view of Themiste minor minor (Ikeda), showing posteriorly orientated hooks. C, Introvert hooks of Phascolosoma arcuatum (Gray). D, Introvert hooks of Phascolosoma glabrum glabrum (Sluiter). E, Anal shield of Aspidosiphon muelleri Diesing. F, Introvert unidentate hook of Aspidosiphon muelleri Diesing. G, Anal shield of Aspidosiphon coyi de Quatrefages, showing grooves on the surface of anal shield. H, Introvert hook of Aspidosiphon coyi de Quatrefages, showing a small secondary tooth on the hook. Scalebars: A, 100 μm; B, E, G, 1 mm; C D, 80 μm; F, 10 μm; H, 20 μm." httpUri="https://zenodo.org/record/187026/files/figure.png" pageId="7" pageNumber="58">Fig. 3</figureCitation>
C
</bibRefCitation>
). It is commonly seen in the Indo-West Pacific region (from northeastern
<collectingCountry id="CB88766DBF11D15CFAE3F8941871586F" box="[1345,1405,1869,1895]" name="India" pageId="7" pageNumber="58">India</collectingCountry>
to southern
<collectingCountry id="CB88766DBF11D15CFEA1F8AD1C475886" box="[259,331,1908,1934]" name="China" pageId="7" pageNumber="58">China</collectingCountry>
) and known to tolerate brackish-water environments.
</paragraph>
</subSubSection>
</treatment>
</document>