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<document id="7B43CA6B824123E6F7DE436C414F1F0A" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C6A9B3EFF67F7439C64FEBB" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus asitius Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="27" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="26" updateTime="1698455438235" updateUser="plazi">
<mods:mods id="654226CC0CC829ED12ADC89AB19FA85E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D760D0598CB4CF2BC48C7EA2423CCE5F">
<mods:title id="51FE123D07E43256A31D4338EEA627D5">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
</mods:titleInfo>
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<mods:role id="753D009A6C088F66016AB20E83339D9F">
<mods:roleTerm id="4A24635DDC60FE0A7300E0E0A50D8450">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BA4110B04FBCFC2F1ED70FBD6F50D7F2">Azevedo, Celso O.</mods:namePart>
</mods:name>
<mods:typeOfResource id="34222FF6DD4B6594911E6B227EC96B10">text</mods:typeOfResource>
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<mods:title id="5670064533417A31CF43E55E4F7085C1">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2FB35B3DE149CD368541281053628323">
<mods:date id="6CBCFD22DFCE1DC4286F611A7AAEFED7">2017</mods:date>
<mods:detail id="C42E8119D3CE50F261970FEFE9E346FC" type="pubDate">
<mods:number id="5BEDFFE899CE52DC5D07D9660AB9FC75">2017-10-13</mods:number>
</mods:detail>
<mods:detail id="8831284E1B492B5FDCBA9F546E76B146" type="volume">
<mods:number id="F48D418736AD020363D3F3C7D5216E6D">4335</mods:number>
</mods:detail>
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<mods:number id="C6793B6505AE6879D6DDBCF56CDEC148">1</mods:number>
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<mods:classification id="77645BAF539D2F6F51EC0BE48735357A">journal article</mods:classification>
<mods:identifier id="8FF3098257275294AF2B4BAF2F2E6932" type="CLB-Dataset">31852</mods:identifier>
<mods:identifier id="4FE5637A5E702AA5625F37E131D90471" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
<mods:identifier id="D3995938BB1FC209581ADC3E153DE1D7" type="GBIF-Dataset">f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf</mods:identifier>
<mods:identifier id="D5C68A2A634A399F1B1598B31F5D3D4A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="36F962B03B07555E90D14AD7DBA6CCAD" type="Zenodo-Dep">1010014</mods:identifier>
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<treatment id="3B3687C07C6A9B3EFF67F7439C64FEBB" ID-DOI="http://doi.org/10.5281/zenodo.5020346" ID-GBIF-Taxon="135435235" ID-Zenodo-Dep="5020346" LSID="urn:lsid:plazi:treatment:3B3687C07C6A9B3EFF67F7439C64FEBB" httpUri="http://treatment.plazi.org/id/3B3687C07C6A9B3EFF67F7439C64FEBB" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<subSubSection id="FB85655D7C6A9B3DFF67F7439ED6FA68" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="B32036D67C6A9B3DFF67F7439DB9FA49" blockId="25.[151,732,1404,1463]" box="[151,732,1404,1430]" pageId="25" pageNumber="26">
<heading id="E86881BA7C6A9B3DFF67F7439DB9FA49" bold="true" box="[151,732,1404,1430]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="81EBEAC47C6A9B3DFF67F7439DB9FA49" bold="true" box="[151,732,1404,1430]" pageId="25" pageNumber="26">
<taxonomicName id="749F4D557C6A9B3DFF67F7439D14FA49" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,625,1404,1430]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="asitius" status="sp. nov.">
<emphasis id="81EBEAC47C6A9B3DFF67F7439EF2FA49" bold="true" box="[151,407,1404,1430]" italics="true" pageId="25" pageNumber="26">Dissomphalus asitius</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="9AD857BF7C6A9B3DFD8FF7439DB9FA49" box="[639,732,1404,1430]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C6A9B3DFF67F7A19ED6FA68" blockId="25.[151,732,1404,1463]" box="[151,435,1438,1463]" pageId="25" pageNumber="26">
(
<figureCitation id="2BA42A537C6A9B3DFF6FF7A19F82FA68" box="[159,231,1438,1463]" captionStart="FIGURES 1 – 12" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[151,1436,195,1484]" captionTargetId="figure@10.[151,1436,195,1489]" captionTargetPageId="10" captionText="FIGURES 1 – 12. Dissomphalus spp., ♂, head, dorsal view, except 2 lateral view. 1 – 2. D. isteus sp. nov. 3. D. azagus sp. nov. 4. D. asfartus sp. nov. 5. D. iozus sp. nov. 6. D. asitius sp. nov. 7. D. ibocius sp. nov. 8. D. tisinus sp. nov. 9. D. ardonius sp. nov. 10. D. arteus sp. nov. 11. D. ettus sp. nov. 12. D. ibrastus sp. nov.. Scale bars = 20 µm, except fig. 10 = 50 µm and fig. 4 = 100 µm." httpUri="https://zenodo.org/record/1010016/files/figure.png" pageId="25" pageNumber="26">Figs 6</figureCitation>
,
<figureCitation id="2BA42A537C6A9B3DFF03F7A09E77FA68" box="[243,274,1439,1463]" captionStart="FIGURES 33 – 44" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[151,1436,553,1886]" captionTargetId="figure@13.[151,1436,553,1886]" captionTargetPageId="13" captionText="FIGURES 33 – 44. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 33. D. isteus sp. nov. 34. D. azagus sp. nov. 35. D. asfartus sp. nov. 36. D. iozus sp. nov. 37. D. asitius sp. nov. 38. D. ibocius sp. nov. 39. D. tisinus sp. nov. 40. D. ardonius sp. nov. 41. D. arteus sp. nov. 42. D. ettus sp. nov. 43. D. ibrastus sp. nov. 44. D. racarus sp. nov.. Scale bars = 20 µm, except fig. 35 = 200 µm." httpUri="https://zenodo.org/record/1010022/files/figure.png" pageId="25" pageNumber="26">37</figureCitation>
,
<figureCitation id="2BA42A537C6A9B3DFEEDF7A19E5EFA68" box="[285,315,1438,1463]" captionStart="FIGURES 64 – 75" captionStartId="16.[151,264,1938,1961]" captionTargetBox="[175,1408,606,1889]" captionTargetId="figure@16.[151,1436,569,1917]" captionTargetPageId="16" captionText="FIGURES 64 – 75. Dissomphalus spp., ♂, hypopygium, internal view. 64. D. isteus sp. nov. 65. D. azagus sp. nov. 66. D. asfartus sp. nov. 67. D. iozus sp. nov. 68. D. asitius sp. nov. 69. D. ibocius sp. nov. 70. D. tisinus sp. nov. 71. D. ardonius sp. nov. 72. D. arteus sp. nov. 73. D. ettus sp. nov. 74. D. ibrastus sp. nov. 75. D. racarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010028/files/figure.png" pageId="25" pageNumber="26">68</figureCitation>
,
<figureCitation id="2BA42A537C6A9B3DFEB7F7A19ECFFA68" box="[327,426,1438,1463]" captionStart="FIGURES 123 – 135" captionStartId="26.[151,264,1820,1843]" captionTargetBox="[170,1430,415,1784]" captionTargetId="figure@26.[151,1436,411,1799]" captionTargetPageId="26" captionText="FIGURES 123 – 135. Dissomphalus spp., ♂, genitália. 123 – 124. D. angulatus 123. Ventral view. 124. Lateral view. 125 – 126. D. asitius sp. nov. 125. Dorsal view. 126. Ventral view. 127 – 128. D. ibocius sp. nov. 127. Dorsal view. 128. Ventral view. 129 – 130. D. tisinus sp. nov. 129. Dorsal view. 130. Ventral view. 131 – 132. D. cervoides 131. Dorsal view. 132. Ventral view. 133 – 135. D. ardonius sp. nov. 133. Dorsal view. 134. Ventral view. 135. Lateral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010038/files/figure.png" pageId="25" pageNumber="26">125–126</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6A9B3DFF67F7D99E1DF854" pageId="25" pageNumber="26" type="description">
<paragraph id="B32036D67C6A9B3DFF67F7D99E1DF854" blockId="25.[151,1437,1510,2003]" pageId="25" pageNumber="26">
<emphasis id="81EBEAC47C6A9B3DFF67F7D99E4CFA20" bold="true" box="[151,297,1510,1535]" pageId="25" pageNumber="26">Description.</emphasis>
Male. Body length
<quantity id="74679B337C6A9B3DFDE6F7D89DE4FA21" box="[534,641,1511,1535]" metricMagnitude="-3" metricUnit="m" metricValue="2.03" pageId="25" pageNumber="26" unit="mm" value="2.03">2.03 mm</quantity>
. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (
<figureCitation id="2BA42A537C6A9B3DFF14F4359E4FF9FC" box="[228,298,1546,1571]" captionStart="FIGURES 1 – 12" captionStartId="10.[151,264,1512,1535]" captionTargetBox="[151,1436,195,1484]" captionTargetId="figure@10.[151,1436,195,1489]" captionTargetPageId="10" captionText="FIGURES 1 – 12. Dissomphalus spp., ♂, head, dorsal view, except 2 lateral view. 1 – 2. D. isteus sp. nov. 3. D. azagus sp. nov. 4. D. asfartus sp. nov. 5. D. iozus sp. nov. 6. D. asitius sp. nov. 7. D. ibocius sp. nov. 8. D. tisinus sp. nov. 9. D. ardonius sp. nov. 10. D. arteus sp. nov. 11. D. ettus sp. nov. 12. D. ibrastus sp. nov.. Scale bars = 20 µm, except fig. 10 = 50 µm and fig. 4 = 100 µm." httpUri="https://zenodo.org/record/1010016/files/figure.png" pageId="25" pageNumber="26">Fig. 6</figureCitation>
). Mandible with two distal teeth. Median clypeal lobe ill-defined, with one angulate tooth; median clypeal carina low in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest straight. Pronotal disc coriaceous; anterior margin coarse. Metasoma (
<figureCitation id="2BA42A537C6A9B3DFB1EF46D9A24F9B4" box="[1262,1345,1618,1643]" captionStart="FIGURES 33 – 44" captionStartId="13.[151,264,1907,1930]" captionTargetBox="[151,1436,553,1886]" captionTargetId="figure@13.[151,1436,553,1886]" captionTargetPageId="13" captionText="FIGURES 33 – 44. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 33. D. isteus sp. nov. 34. D. azagus sp. nov. 35. D. asfartus sp. nov. 36. D. iozus sp. nov. 37. D. asitius sp. nov. 38. D. ibocius sp. nov. 39. D. tisinus sp. nov. 40. D. ardonius sp. nov. 41. D. arteus sp. nov. 42. D. ettus sp. nov. 43. D. ibrastus sp. nov. 44. D. racarus sp. nov.. Scale bars = 20 µm, except fig. 35 = 200 µm." httpUri="https://zenodo.org/record/1010022/files/figure.png" pageId="25" pageNumber="26">Fig. 37</figureCitation>
). Tergal process with median, shallow, subcircular and large depression, with small, low and dorsad tubercle placed on center of depression, pit very small, with short and dense tuft of setae. Posterior hypopygeal margin strongly concave (
<figureCitation id="2BA42A537C6A9B3DFEF4F4819E3FF908" box="[260,346,1726,1751]" captionStart="FIGURES 64 – 75" captionStartId="16.[151,264,1938,1961]" captionTargetBox="[175,1408,606,1889]" captionTargetId="figure@16.[151,1436,569,1917]" captionTargetPageId="16" captionText="FIGURES 64 – 75. Dissomphalus spp., ♂, hypopygium, internal view. 64. D. isteus sp. nov. 65. D. azagus sp. nov. 66. D. asfartus sp. nov. 67. D. iozus sp. nov. 68. D. asitius sp. nov. 69. D. ibocius sp. nov. 70. D. tisinus sp. nov. 71. D. ardonius sp. nov. 72. D. arteus sp. nov. 73. D. ettus sp. nov. 74. D. ibrastus sp. nov. 75. D. racarus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010028/files/figure.png" pageId="25" pageNumber="26">Fig. 68</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C6A9B3DFE18F4819DEFF908" box="[488,650,1726,1751]" captionStart="FIGURES 123 – 135" captionStartId="26.[151,264,1820,1843]" captionTargetBox="[170,1430,415,1784]" captionTargetId="figure@26.[151,1436,411,1799]" captionTargetPageId="26" captionText="FIGURES 123 – 135. Dissomphalus spp., ♂, genitália. 123 – 124. D. angulatus 123. Ventral view. 124. Lateral view. 125 – 126. D. asitius sp. nov. 125. Dorsal view. 126. Ventral view. 127 – 128. D. ibocius sp. nov. 127. Dorsal view. 128. Ventral view. 129 – 130. D. tisinus sp. nov. 129. Dorsal view. 130. Ventral view. 131 – 132. D. cervoides 131. Dorsal view. 132. Ventral view. 133 – 135. D. ardonius sp. nov. 133. Dorsal view. 134. Ventral view. 135. Lateral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010038/files/figure.png" pageId="25" pageNumber="26">Figs 125–126</figureCitation>
): paramere with dorsal margin entirely wide; apical margin truncate. Aedeagal ventral ramus shorter than dorsal body, wide, abruptly narrowing apicad; cross section laminar; surface subvertical; inner margin sinuous; outer margin sinuous; apex long, toothed, angled outward; additional inner ramus absent; basal stub absent. Aedeagal dorsal body two pairs of apical lobes; outer lobe long, horizontal and wide, with apical margin acute and dorsad, inner pair stout, membranous and setose. Apodeme not extending beyond genital ring.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C6A9B3EFF37F5A99ED1FF0A" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="discussion">
<paragraph id="B32036D67C6A9B3EFF37F5A99ED1FF0A" blockId="25.[151,1437,1510,2003]" lastBlockId="26.[151,1437,151,357]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="81EBEAC47C6A9B3DFF37F5A99E5EF870" bold="true" box="[199,315,1942,1967]" pageId="25" pageNumber="26">Remarks.</emphasis>
This species is easily recognized by having the dorsal margin of the paramere almost straight, the apex of the outer lobe of the aedeagal dorsal body abruptly acute, the aedeagal ventral ramus with a large fold and the genital ring bifurcated in the distal region in ventral view. This combination of characters is unusual for
<taxonomicName id="749F4D557C699B3EFF67F2839E6FFF0A" box="[151,266,188,213]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bifoveatus">
<emphasis id="81EBEAC47C699B3EFF67F2839E6FFF0A" box="[151,266,188,213]" italics="true" pageId="26" pageNumber="27">bifoveatus</emphasis>
</taxonomicName>
species-group.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C699B3EFF37F2E09D27FE9F" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="B32036D67C699B3EFF37F2E09D27FE9F" blockId="26.[151,1437,151,357]" pageId="26" pageNumber="27">
<materialsCitation id="03F73C8B7C699B3EFF37F2E09AFCFEC3" ID-GBIF-Occurrence="1671715955" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Teribe, P &amp; Tr &amp; Malaise, A" country="Panama" latitude="9.406944" location="Material" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="26" pageNumber="27" specimenCount="1" specimenCount-male="1" stateProvince="Bocas del Toro" typeStatus="holotype">
<emphasis id="81EBEAC47C699B3EFF37F2E09ED6FF27" bold="true" box="[199,435,223,248]" pageId="26" pageNumber="27">
<location id="B640600D7C699B3EFF37F2E09E4AFF27" LSID="urn:lsid:plazi:treatment:3B3687C07C6A9B3EFF67F7439C64FEBB:B640600D7C699B3EFF37F2E09E4AFF27" box="[199,303,223,248]" country="Panama" latitude="9.406944" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" name="Material" pageId="26" pageNumber="27" stateProvince="Bocas del Toro">Material</location>
examined.
</emphasis>
<typeStatus id="6C2488747C699B3EFE4CF2E09D42FF27" box="[444,551,223,248]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C699B3EFDC5F2DF9D29FF28" box="[565,588,224,247]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
,
<collectingCountry id="CB8876467C699B3EFDAAF2DF9DB1FF27" box="[602,724,224,248]" name="Panama" pageId="26" pageNumber="27">PANAMA</collectingCountry>
,
<collectingRegion id="715BF8347C699B3EFD11F2DE9CEBFF27" box="[737,910,223,248]" country="Panama" name="Bocas del Toro" pageId="26" pageNumber="27">
<emphasis id="81EBEAC47C699B3EFD11F2DE9CEBFF27" box="[737,910,223,248]" italics="true" pageId="26" pageNumber="27">Bocas del Toro</emphasis>
</collectingRegion>
:
<collectingMunicipality id="5344ACAC7C699B3EFC6FF2E09C8BFF27" box="[927,1006,223,248]" pageId="26" pageNumber="27">Wekso</collectingMunicipality>
,
<collectorName id="1E6A53007C699B3EFC0BF2E09B07FF28" box="[1019,1122,223,248]" pageId="26" pageNumber="27">Teribe, P</collectorName>
[arque] I[nternacional] L[a] A[mistad], 50 msnm, [
<geoCoordinate id="D6AB50117C699B3EFE6FF33B9D41FEC2" box="[415,548,260,285]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="9.406944">09º24'25&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C699B3EFDC4F33B9DA4FEC2" box="[564,705,260,285]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-82.93889">82º56'20&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C699B3EFD27F33B9CF2FEC3" box="[727,919,260,284]" pageId="26" pageNumber="27" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">
<date id="C72110167C699B3EFD27F33B9CF2FEC3" box="[727,919,260,284]" pageId="26" pageNumber="27" value="1999-10-17" valueMax="1999-10-24" valueMin="1999-10-17">17–24 oct. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C699B3EFC57F33A9CA6FEC3" box="[935,963,261,284]" pageId="26" pageNumber="27">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C699B3EFBEAF33B9BF9FEC3" box="[1050,1180,260,285]" pageId="26" pageNumber="27">Malaise, A</collectorName>
. Santos[col.] (
<collectionCode id="D58EAE137C699B3EFAB4F33B9AF5FEC3" box="[1348,1424,260,284]" pageId="26" pageNumber="27">MIUP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="03F73C8B7C699B3EFF67F3179D5BFE9F" ID-GBIF-Occurrence="1671715971" box="[151,574,295,320]" collectingDate="1999-10-17" collectingDateMax="1999-10-24" collectingDateMin="1999-10-17" collectionCode="MIUP" collectorName="Teribe, P &amp; Tr &amp; Malaise, A" country="Panama" latitude="9.406944" location="Material" longLatPrecision="21" longitude="-82.93889" municipality="Wekso" pageId="26" pageNumber="27" specimenCount="5" specimenCount-male="5" stateProvince="Bocas del Toro" typeStatus="paratype">
<typeStatus id="6C2488747C699B3EFF67F3179E6CFE9F" box="[151,265,296,320]" pageId="26" pageNumber="27" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A599FD5F7C699B3EFEE3F3189E52FE9F" box="[275,311,295,320]" pageId="26" pageNumber="27" type="male">5♂</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C699B3EFF37F3739C64FEBB" box="[199,769,332,357]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="B32036D67C699B3EFF37F3739C64FEBB" blockId="26.[151,1437,151,357]" box="[199,769,332,357]" pageId="26" pageNumber="27">
<emphasis id="81EBEAC47C699B3EFF37F3739E3CFEBA" bold="true" box="[199,345,332,357]" pageId="26" pageNumber="27">Distribution</emphasis>
(
<figureCitation id="2BA42A537C699B3EFE98F3739EAEFEBB" box="[360,459,332,357]" captionStart="FIGURES 224 – 233" captionStartId="66.[151,264,1814,1837]" captionTargetBox="[182,1404,199,1768]" captionTargetId="figure@66.[169,1413,193,1773]" captionTargetPageId="66" captionText="FIGURES 224 – 233. Known distribution of Dissomphalus in Panama. 224. Provinces and Counties of Panama. 1. Bocas del Toro. 2. Chiriquí. 3. Ngöbe-Buglé County. 4. Veraguas. 5. Herrera. 6. Los Santos. 7. Colón. 8. Coclé. 9. Panama. 10. Guna Yala County. 11. Darién. 12. Emberá Woonan County. 225. D. abarus sp. nov., D. afestus sp. nov. and D. altivolans. 226. D. angulatus sp. nov., D. ardonius sp. nov. and D. aretisus sp. nov.. 227. D. apertus, D. areius sp. nov. and D. arteobius sp. nov.. 228. D. arbeius sp. nov., D. asitius sp. nov. and D. brasiliensis. 229. D. arteus sp. nov., D. cervoides and D. culteratus. 230. D. asfartus sp. nov.. 231. D. azagus sp. nov., D. bilobatus and D. coronatus. 232. D. borus sp. nov., D. citus sp. nov. and D. curvifoveatus. 233. D. declinatus, D. dilatatus and D. ettus sp. nov.." httpUri="https://zenodo.org/record/1010056/files/figure.png" pageId="26" pageNumber="27">Fig. 228</figureCitation>
).
<collectingRegion id="715BF8347C699B3EFE2EF3729D52FEBB" box="[478,567,333,356]" country="Panama" name="Panama" pageId="26" pageNumber="27">
<collectingCountry id="CB8876467C699B3EFE2EF3729D52FEBB" box="[478,567,333,356]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C699B3EFDB6F3739D93FEBB" box="[582,758,332,357]" country="Panama" name="Bocas del Toro" pageId="26" pageNumber="27">Bocas del Toro</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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