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<document id="B7652DCE9ACF7386CB3033776AC1C8DD" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C4F9B18FF67F2A89B6DFAF0" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus borus Azevedo 2017, sp. nov." docType="treatment" docVersion="15" lastPageNumber="61" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="61" updateTime="1698455438235" updateUser="plazi">
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<mods:title id="429EBA60E5F3BDDCE5BAA0B68CCA1416">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
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<mods:namePart id="48544E451408BB89381E38503FCC7BE7">Azevedo, Celso O.</mods:namePart>
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<mods:title id="5E453E914CFAFD26B3A4B9F206DB052D">Zootaxa</mods:title>
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<mods:date id="F0367634D706D9685E49148585322E89">2017</mods:date>
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<mods:number id="CFDF446013B3529D8C85E6D78F900998">2017-10-13</mods:number>
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<mods:number id="3F0B1DB0706F2424B1F109D63B951ED8">4335</mods:number>
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<mods:identifier id="98D292AB30E93EE83C316541C5007065" type="DOI">10.11646/zootaxa.4335.1.1</mods:identifier>
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<treatment id="3B3687C07C4F9B18FF67F2A89B6DFAF0" ID-DOI="http://doi.org/10.5281/zenodo.5020522" ID-GBIF-Taxon="135435281" ID-Zenodo-Dep="5020522" LSID="urn:lsid:plazi:treatment:3B3687C07C4F9B18FF67F2A89B6DFAF0" httpUri="http://treatment.plazi.org/id/3B3687C07C4F9B18FF67F2A89B6DFAF0" lastPageNumber="61" pageId="60" pageNumber="61">
<subSubSection id="FB85655D7C4F9B18FF67F2A89EA5FF0C" pageId="60" pageNumber="61" type="nomenclature">
<paragraph id="B32036D67C4F9B18FF67F2A89DB6FF6D" blockId="60.[151,723,151,211]" box="[151,723,151,178]" pageId="60" pageNumber="61">
<heading id="E86881BA7C4F9B18FF67F2A89DB6FF6D" bold="true" box="[151,723,151,178]" fontSize="11" level="1" pageId="60" pageNumber="61" reason="1">
<emphasis id="81EBEAC47C4F9B18FF67F2A89DB6FF6D" bold="true" box="[151,723,151,178]" pageId="60" pageNumber="61">
<taxonomicName id="749F4D557C4F9B18FF67F2A89D0CFF6D" authority="Brito &amp; Azevedo" authorityName="Azevedo" authorityYear="2017" box="[151,617,151,178]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="borus" status="sp. nov.">
<emphasis id="81EBEAC47C4F9B18FF67F2A89EE8FF6E" bold="true" box="[151,397,151,177]" italics="true" pageId="60" pageNumber="61">Dissomphalus borus</emphasis>
Brito &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="9AD857BF7C4F9B18FD85F2A79DB6FF6D" box="[629,723,152,178]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B32036D67C4F9B18FF67F2859EA5FF0C" blockId="60.[151,723,151,211]" box="[151,448,186,211]" pageId="60" pageNumber="61">
(
<figureCitation id="2BA42A537C4F9B18FF6FF2859F90FF0C" box="[159,245,186,211]" captionStart="FIGURES 25 32" captionStartId="12.[151,264,1546,1569]" captionTargetBox="[151,1436,555,1525]" captionTargetId="figure@12.[151,1436,555,1525]" captionTargetPageId="12" captionText="FIGURES 25 32. Dissomphalus spp., ♂, head in dorsal view. 25. D. abarus sp. nov. 26. D. arteobius sp. nov. 27. D. borus sp. nov. 28. D. geortus sp. nov. 29. D. ilarius sp. nov. 30. D. isortus sp. nov. 31. D. trisus sp. nov. 32. D. ubracus sp. nov.. Scale bars = 20 µm, except fig. 31 = 50 µm and fig. 32 = 100 µm." httpUri="https://zenodo.org/record/1010020/files/figure.png" pageId="60" pageNumber="61">Figs 27</figureCitation>
,
<figureCitation id="2BA42A537C4F9B18FEF1F2859E45FF0C" box="[257,288,186,211]" captionStart="FIGURES 57 63" captionStartId="15.[151,264,1212,1235]" captionTargetBox="[151,1436,197,1188]" captionTargetId="figure@15.[151,1436,193,1191]" captionTargetPageId="15" captionText="FIGURES 57 63. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 57. D. arteobius sp. nov. 58. D. borus sp. nov. 59. D. geortus sp. nov. 60. D. ilarius sp. nov. 61. D. isortus sp. nov. 62. D. trisus sp. nov. 63. D. ubracus sp. nov. Scale bars = 20 µm except, fig. 62 = 50 µm and fig. 63 = 200 µm." httpUri="https://zenodo.org/record/1010026/files/figure.png" pageId="60" pageNumber="61">58</figureCitation>
,
<figureCitation id="2BA42A537C4F9B18FEDBF2849E2FFF0C" box="[299,330,187,211]" captionStart="FIGURES 88 94" captionStartId="18.[151,264,1101,1124]" captionTargetBox="[190,1378,187,1060]" captionTargetId="figure@18.[179,1393,179,1082]" captionTargetPageId="18" captionText="FIGURES 88 94. Dissomphalus spp., ♂, hypopygium, internal view. 88. D. arteobius sp. nov. 89. D. borus sp. nov. 90. D. geortus sp. nov. 91. D. ilarius sp. nov. 92. D. isortus sp. nov. 93. D. trisus sp. nov. 94. D. ubracus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010032/files/figure.png" pageId="60" pageNumber="61">89</figureCitation>
,
<figureCitation id="2BA42A537C4F9B18FEA5F2849EDDFF0C" box="[341,440,187,211]" captionStart="FIGURES 211 222" captionStartId="61.[151,264,1820,1843]" captionTargetBox="[153,1424,424,1779]" captionTargetId="figure@61.[151,1436,411,1799]" captionTargetPageId="61" captionText="FIGURES 211 222. Dissomphalus spp., ♂, genitalia. 211 212. D. borus sp. nov. 211. Dorsal view. 212. Ventral view. 213 214. D. geortus sp. nov. 213. Dorsal view. 214. Ventral view. 215 216. D. ilarius sp. nov. 215. Dorsal view. 216. Ventral view. 217 218. D. isortus sp. nov. 217. Dorsal view. 218. Ventral view. 219 220. D. trisus sp. nov. 219. Dorsal view. 220. Ventral view. 221 222. D. ubracus sp. nov. 221. Dorsal view. 222. Ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010052/files/figure.png" pageId="60" pageNumber="61">211212</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4F9B18FF67F33D9C22FD15" pageId="60" pageNumber="61" type="description">
<paragraph id="B32036D67C4F9B18FF67F33D9FAEFD78" blockId="60.[151,1437,258,1327]" pageId="60" pageNumber="61">
<emphasis id="81EBEAC47C4F9B18FF67F33D9E4CFEC4" bold="true" box="[151,297,258,283]" pageId="60" pageNumber="61">Description.</emphasis>
Male. Body length
<quantity id="74679B337C4F9B18FDE6F33C9DE4FEC5" box="[534,641,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" pageId="60" pageNumber="61" unit="mm" value="1.73">1.73 mm</quantity>
. Color: head and mesosoma dark castaneous; metasoma castaneous. Head (
<figureCitation id="2BA42A537C4F9B18FF13F3199E5DFEE0" box="[227,312,294,319]" captionStart="FIGURES 25 32" captionStartId="12.[151,264,1546,1569]" captionTargetBox="[151,1436,555,1525]" captionTargetId="figure@12.[151,1436,555,1525]" captionTargetPageId="12" captionText="FIGURES 25 32. Dissomphalus spp., ♂, head in dorsal view. 25. D. abarus sp. nov. 26. D. arteobius sp. nov. 27. D. borus sp. nov. 28. D. geortus sp. nov. 29. D. ilarius sp. nov. 30. D. isortus sp. nov. 31. D. trisus sp. nov. 32. D. ubracus sp. nov.. Scale bars = 20 µm, except fig. 31 = 50 µm and fig. 32 = 100 µm." httpUri="https://zenodo.org/record/1010020/files/figure.png" pageId="60" pageNumber="61">Fig. 27</figureCitation>
). Mandible with four distal teeth. Median clypeal lobe ill-defined, with one rounded tooth; median clypeal carina high in profile, complete or nearly so, straight in profile or nearly so. Frons weakly coriaceous and punctures small. Vertex crest weakly concave. Pronotal disc weakly coriaceous; anterior margin ecarinate. Metasoma (
<figureCitation id="2BA42A537C4F9B18FEECF3AD9E15FE74" box="[284,368,402,427]" captionStart="FIGURES 57 63" captionStartId="15.[151,264,1212,1235]" captionTargetBox="[151,1436,197,1188]" captionTargetId="figure@15.[151,1436,193,1191]" captionTargetPageId="15" captionText="FIGURES 57 63. Dissomphalus spp., ♂, metasomal tergal process, dorsal view. 57. D. arteobius sp. nov. 58. D. borus sp. nov. 59. D. geortus sp. nov. 60. D. ilarius sp. nov. 61. D. isortus sp. nov. 62. D. trisus sp. nov. 63. D. ubracus sp. nov. Scale bars = 20 µm except, fig. 62 = 50 µm and fig. 63 = 200 µm." httpUri="https://zenodo.org/record/1010026/files/figure.png" pageId="60" pageNumber="61">Fig. 58</figureCitation>
). Tergal process lateral, without depression, with very small, low and dorsad tubercle, pit very small, with short tuft of setae. Posterior hypopygeal margin sinuous (
<figureCitation id="2BA42A537C4F9B18FC52F3899C92FE10" box="[930,1015,438,463]" captionStart="FIGURES 88 94" captionStartId="18.[151,264,1101,1124]" captionTargetBox="[190,1378,187,1060]" captionTargetId="figure@18.[179,1393,179,1082]" captionTargetPageId="18" captionText="FIGURES 88 94. Dissomphalus spp., ♂, hypopygium, internal view. 88. D. arteobius sp. nov. 89. D. borus sp. nov. 90. D. geortus sp. nov. 91. D. ilarius sp. nov. 92. D. isortus sp. nov. 93. D. trisus sp. nov. 94. D. ubracus sp. nov.. Scale bars = 125 µm." httpUri="https://zenodo.org/record/1010032/files/figure.png" pageId="60" pageNumber="61">Fig. 89</figureCitation>
). Genitalia (
<figureCitation id="2BA42A537C4F9B18FB71F3899A44FE10" box="[1153,1313,438,463]" captionStart="FIGURES 211 222" captionStartId="61.[151,264,1820,1843]" captionTargetBox="[153,1424,424,1779]" captionTargetId="figure@61.[151,1436,411,1799]" captionTargetPageId="61" captionText="FIGURES 211 222. Dissomphalus spp., ♂, genitalia. 211 212. D. borus sp. nov. 211. Dorsal view. 212. Ventral view. 213 214. D. geortus sp. nov. 213. Dorsal view. 214. Ventral view. 215 216. D. ilarius sp. nov. 215. Dorsal view. 216. Ventral view. 217 218. D. isortus sp. nov. 217. Dorsal view. 218. Ventral view. 219 220. D. trisus sp. nov. 219. Dorsal view. 220. Ventral view. 221 222. D. ubracus sp. nov. 221. Dorsal view. 222. Ventral view. Scale bars = 250 µm." httpUri="https://zenodo.org/record/1010052/files/figure.png" pageId="60" pageNumber="61">Figs 211212</figureCitation>
): paramere with dorsal margin wide basally; apical margin truncate. Aedeagal ventral ramus higher than dorsal body, wide, progressively narrowing apicad; cross section laminar; surface horizontal; inner margin straight; outer margin sinuous; apex long, simple, parallel; additional inner ramus absent; basal stub very short and narrow. Aedeagal dorsal body with two pairs of apical lobes; outer lobe very short, horizontal and narrow, with apical margin abruptly acute and slightly mesad; inner pair stout, membranous and setose. Apodeme extending beyond genital ring.
</paragraph>
<paragraph id="B32036D67C4F9B18FF37F08D9C22FD15" blockId="60.[151,1437,258,1327]" box="[199,839,690,715]" pageId="60" pageNumber="61">
<emphasis id="81EBEAC47C4F9B18FF37F08D9E59FD14" bold="true" box="[199,316,690,715]" pageId="60" pageNumber="61">Variation.</emphasis>
Pit of tubercle of tergal process without setae.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4F9B18FF37F0E99F82FC18" pageId="60" pageNumber="61" type="discussion">
<paragraph id="B32036D67C4F9B18FF37F0E99F82FC18" blockId="60.[151,1437,258,1327]" pageId="60" pageNumber="61">
<emphasis id="81EBEAC47C4F9B18FF37F0E99E5EFD30" bold="true" box="[199,315,726,751]" pageId="60" pageNumber="61">Remarks.</emphasis>
This species is easily recognized by having the dorsal margin of the paramere straight and the apex very excavated in the apical region and the apical lobes of the aedeagal dorsal body very short. This species is similar to
<taxonomicName id="749F4D557C4F9B18FEFCF1209E1BFCE9" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[268,382,799,822]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="azagus" status="sp. nov.">
<emphasis id="81EBEAC47C4F9B18FEFCF1209E1BFCE9" box="[268,382,799,822]" italics="true" pageId="60" pageNumber="61">D. azagus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C4F9B18FE77F1219E86FCE9" box="[391,483,798,823]" pageId="60" pageNumber="61" rank="species">
<emphasis id="81EBEAC47C4F9B18FE77F1219EB9FCE8" bold="true" box="[391,476,798,823]" pageId="60" pageNumber="61">sp. nov</emphasis>
.
</taxonomicNameLabel>
by having the dorsal margin of the paramere straight and the apex very excavated. However, this species has the tergal process present, the apical lobes of the aedeagal dorsal body very short and the aedeagal ventral ramus progressively narrowing apicad, whereas
<taxonomicName id="749F4D557C4F9B18FC8BF1589C89FCA1" authority="Azevedo, 2017" authorityName="Azevedo" authorityYear="2017" box="[891,1004,871,894]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="azagus" status="sp. nov.">
<emphasis id="81EBEAC47C4F9B18FC8BF1589C89FCA1" box="[891,1004,871,894]" italics="true" pageId="60" pageNumber="61">D. azagus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9AD857BF7C4F9B18FC04F1599B2FFCA0" box="[1012,1098,870,895]" pageId="60" pageNumber="61" rank="species">
<emphasis id="81EBEAC47C4F9B18FC04F1599B2FFCA0" bold="true" box="[1012,1098,870,895]" pageId="60" pageNumber="61">sp. nov</emphasis>
</taxonomicNameLabel>
has the tergal process absent, the apical lobes of the aedeagal dorsal body long and the apex of the aedeagal ventral ramus narrowing abruptly apicad.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4F9B18FF37F1ED9B68FAD4" pageId="60" pageNumber="61" type="materials_examined">
<paragraph id="B32036D67C4F9B18FF37F1ED9B68FAD4" blockId="60.[151,1437,258,1327]" pageId="60" pageNumber="61">
<materialsCitation id="03F73C8B7C4F9B18FF37F1ED9A71FC34" ID-GBIF-Occurrence="1671715805" box="[199,1300,978,1003]" collectorName="Veraguas Pr &amp; Coiba" country="Panama" location="PANAMA" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="PANAMA" typeStatus="holotype">
<emphasis id="81EBEAC47C4F9B18FF37F1ED9ED7FC34" bold="true" box="[199,434,978,1003]" pageId="60" pageNumber="61">Material examined.</emphasis>
<typeStatus id="6C2488747C4F9B18FE4BF1ED9D43FC34" box="[443,550,978,1003]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
,
<specimenCount id="A599FD5F7C4F9B18FDC3F1EC9D2FFC35" box="[563,586,979,1002]" pageId="60" pageNumber="61" type="male"></specimenCount>
,
<collectingRegion id="715BF8347C4F9B18FDA8F1EC9DB7FC34" box="[600,722,979,1003]" country="Panama" name="Panama" pageId="60" pageNumber="61">
<collectingCountry id="CB8876467C4F9B18FDA8F1EC9DB7FC34" box="[600,722,979,1003]" name="Panama" pageId="60" pageNumber="61">PANAMA</collectingCountry>
</collectingRegion>
,
<emphasis id="81EBEAC47C4F9B18FD2EF1EB9CB1FC35" box="[734,980,979,1003]" italics="true" pageId="60" pageNumber="61">
<collectorName id="1E6A53007C4F9B18FD2EF1EB9C0EFC35" box="[734,875,979,1003]" pageId="60" pageNumber="61">
<collectingRegion id="715BF8347C4F9B18FD2EF1EB9C23FC34" box="[734,838,980,1003]" country="Panama" name="Veraguas" pageId="60" pageNumber="61">Veraguas</collectingRegion>
<collectingRegion id="715BF8347C4F9B18FCBFF1EC9C0EFC35" box="[847,875,979,1002]" country="Brazil" name="Para" pageId="60" pageNumber="61">Pr</collectingRegion>
</collectorName>
[ovincia]
</emphasis>
: P[arque] N[acional]
<collectorName id="1E6A53007C4F9B18FB3EF1ED9A71FC34" box="[1230,1300,978,1003]" pageId="60" pageNumber="61">Coiba</collectorName>
</materialsCitation>
,
<materialsCitation id="03F73C8B7C4F9B18FAD1F1ED9F96FBEC" ID-GBIF-Occurrence="1671715809" collectingDate="1929-07-04" collectionCode="MIUP" collectorName="Malaise, R &amp; Cambra, S &amp; Bermudez" country="Iceland" elevation="120" latitude="7.4691668" location="Cerro la Equis" longLatPrecision="21" longitude="-81.75667" pageId="60" pageNumber="61" specimenCount="1" typeStatus="holotype">
<collectingCountry id="CB8876467C4F9B18FAD1F1ED9A2FFC34" box="[1313,1354,978,1003]" name="Iceland" pageId="60" pageNumber="61">Isla</collectingCountry>
Coiba,
<location id="B640600D7C4F9B18FF67F1C89E27FBD0" LSID="urn:lsid:plazi:treatment:3B3687C07C4F9B18FF67F2A89B6DFAF0:B640600D7C4F9B18FF67F1C89E27FBD0" box="[151,322,1014,1039]" country="Iceland" latitude="7.4691668" longLatPrecision="21" longitude="-81.75667" name="Cerro la Equis" pageId="60" pageNumber="61">Cerro la Equis</location>
,
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<quantity id="74679B337C4F9B18FEA2F1C89EFAFBD1" box="[338,415,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="60" pageNumber="61" unit="m" value="120.0">120 m</quantity>
</elevation>
, [
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,
<geoCoordinate id="D6AB50117C4F9B18FDBBF1C99DB2FBD0" box="[587,727,1014,1039]" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="15" value="-81.75667">81º45'24&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C4F9B18FD1DF1C89C2BFBD0" box="[749,846,1014,1039]" pageId="60" pageNumber="61" value="1929-07-04">
<date id="C72110167C4F9B18FD1DF1C89C2BFBD0" box="[749,846,1014,1039]" pageId="60" pageNumber="61" value="1929-07-04">29 jul4</date>
</collectingDate>
ag 1998,
<collectorName id="1E6A53007C4F9B18FC37F1C99B2DFBD1" box="[967,1096,1014,1039]" pageId="60" pageNumber="61">Malaise, R</collectorName>
.
<collectorName id="1E6A53007C4F9B18FBA7F1C99BB6FBD0" box="[1111,1235,1014,1039]" pageId="60" pageNumber="61">Cambra, S</collectorName>
.
<collectorName id="1E6A53007C4F9B18FB12F1C99A30FBD0" box="[1250,1365,1014,1039]" pageId="60" pageNumber="61">Bermudez</collectorName>
[col.] (
<collectionCode id="D58EAE137C4F9B18FF6FF6249F8EFBEC" box="[159,235,1051,1075]" pageId="60" pageNumber="61">MIUP</collectionCode>
)
</materialsCitation>
. Paratypes: 1♂ same as holotype except
<date id="C72110167C4F9B18FD38F6259CFAFBEC" box="[712,927,1050,1075]" pageId="60" pageNumber="61" value="1998-11-30" valueMax="1998-12-04" valueMin="1998-11-30">30nov.4dic.1998</date>
(MIUP); 1♂ same as holotype except
<quantity id="74679B337C4F9B18FAACF6249AF9FBEC" box="[1372,1436,1051,1075]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="60" pageNumber="61" unit="m" value="200.0">200m</quantity>
(MIUP);
<materialsCitation id="03F73C8B7C4F9B18FF0EF6009E7DFBA4" ID-GBIF-Occurrence="1671715822" collectingDate="1999-03-22" collectingDateMax="1999-03-26" collectingDateMin="1999-03-22" collectionCode="MIUP" collectorName="Col. A. &amp; Santos" country="Iceland" elevation="770" latitude="8.518194" location="Altos de Piedra" longLatPrecision="1" longitude="-81.108444" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Santa Fe" typeStatus="holotype">
<specimenCount id="A599FD5F7C4F9B18FF0EF6009E78FB88" box="[254,285,1087,1111]" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
,
<collectingRegion id="715BF8347C4F9B18FED8F6009EE9FB88" box="[296,396,1087,1111]" country="Argentina" name="Santa Fe" pageId="60" pageNumber="61">Santa Fé</collectingRegion>
,
<location id="B640600D7C4F9B18FE66F6019D23FB88" LSID="urn:lsid:plazi:treatment:3B3687C07C4F9B18FF67F2A89B6DFAF0:B640600D7C4F9B18FE66F6019D23FB88" box="[406,582,1086,1111]" country="Iceland" latitude="8.518194" longLatPrecision="1" longitude="-81.108444" name="Altos de Piedra" pageId="60" pageNumber="61" stateProvince="Santa Fe">Altos de Piedra</location>
,
<elevation id="38B2D1E57C4F9B18FDA0F6009DF3FB89" box="[592,662,1087,1111]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="60" pageNumber="61" unit="m" value="770.0">
<quantity id="74679B337C4F9B18FDA0F6009DF3FB89" box="[592,662,1087,1111]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="60" pageNumber="61" unit="m" value="770.0">770 m</quantity>
</elevation>
[
<geoCoordinate id="D6AB50117C4F9B18FD54F6019C56FB88" box="[676,819,1086,1111]" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="1" value="8.518194">08°31'5.5&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C4F9B18FCCFF6019C86FB88" box="[831,995,1086,1111]" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="1" value="-81.108444">81°06'30.4&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C4F9B18FC05F6019BADFB88" box="[1013,1224,1086,1111]" pageId="60" pageNumber="61" value="1999-03-22" valueMax="1999-03-26" valueMin="1999-03-22">
<date id="C72110167C4F9B18FC05F6019BADFB88" box="[1013,1224,1086,1111]" pageId="60" pageNumber="61" value="1999-03-22" valueMax="1999-03-26" valueMin="1999-03-22">2226 marzo 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C4F9B18FB24F6019A1CFB88" box="[1236,1401,1086,1111]" pageId="60" pageNumber="61">Col. A. Santos</collectorName>
, L. D. (
<collectionCode id="D58EAE137C4F9B18FF34F65C9E75FBA4" box="[196,272,1123,1147]" pageId="60" pageNumber="61">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4F9B18FED6F65C9EA8FBA4" ID-GBIF-Occurrence="1671715808" box="[294,461,1122,1147]" country="Iceland" location="Chiriqui" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="holotype">
[
<collectingRegion id="715BF8347C4F9B18FEDFF65D9EEBFBA4" box="[303,398,1122,1147]" country="Panama" name="Chiriqui" pageId="60" pageNumber="61">
<emphasis id="81EBEAC47C4F9B18FEDFF65D9EEBFBA4" box="[303,398,1122,1147]" italics="true" pageId="60" pageNumber="61">Chiriqui</emphasis>
</collectingRegion>
]:
<specimenCount id="A599FD5F7C4F9B18FE59F65C9EA8FBA4" box="[425,461,1123,1147]" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
</materialsCitation>
,
<materialsCitation id="03F73C8B7C4F9B18FE2CF65C9C45FB40" ID-GBIF-Occurrence="1671715811" collectingDate="1994-05-28" collectingDateMax="1994-06-02" collectingDateMin="1994-05-28" collectionCode="UCDC" collectorName="F. Andrews &amp; A. Gilbert" country="Taiwan" county="Finca La Suiza" latitude="8.666667" location="Hornitos" longLatPrecision="1300" longitude="-82.23333" municipality="Km" pageId="60" pageNumber="61" specimenCount="1" stateProvince="Formosa" typeStatus="holotype">
<collectingRegion id="715BF8347C4F9B18FE2CF65C9D24FBA5" box="[476,577,1123,1146]" country="Argentina" name="Formosa" pageId="60" pageNumber="61">
<collectingCountry id="CB8876467C4F9B18FE2CF65C9D24FBA5" box="[476,577,1123,1146]" name="Taiwan" pageId="60" pageNumber="61">Formosa</collectingCountry>
</collectingRegion>
,
<collectingCounty id="5A414E5A7C4F9B18FDBFF65D9D99FBA4" box="[591,764,1122,1147]" pageId="60" pageNumber="61">Finca La Suiza</collectingCounty>
1
<collectingMunicipality id="5344ACAC7C4F9B18FCEDF65C9C22FBA5" box="[797,839,1123,1146]" pageId="60" pageNumber="61">Km</collectingMunicipality>
se.
<location id="B640600D7C4F9B18FC89F65D9CBBFBA4" LSID="urn:lsid:plazi:treatment:3B3687C07C4F9B18FF67F2A89B6DFAF0:B640600D7C4F9B18FC89F65D9CBBFBA4" box="[889,990,1122,1147]" country="Taiwan" county="Finca La Suiza" latitude="8.666667" longLatPrecision="1300" longitude="-82.23333" municipality="Km" name="Hornitos" pageId="60" pageNumber="61" stateProvince="Formosa">Hornitos</location>
, [
<geoCoordinate id="D6AB50117C4F9B18FC02F65C9B31FBA4" box="[1010,1108,1123,1147]" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="925" value="8.666667">08°40'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C4F9B18FB92F65C9BACFBA4" box="[1122,1225,1123,1147]" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="925" value="-82.23333">82°14'W</geoCoordinate>
],
<collectingDate id="D765E9FE7C4F9B18FB2EF65C9AFDFBA4" box="[1246,1432,1123,1147]" pageId="60" pageNumber="61" value="1994-05-28" valueMax="1994-06-02" valueMin="1994-05-28">
<date id="C72110167C4F9B18FB2EF65C9AFDFBA4" box="[1246,1432,1123,1147]" pageId="60" pageNumber="61" value="1994-05-28" valueMax="1994-06-02" valueMin="1994-05-28">28.V2.VI.1994</date>
</collectingDate>
,
<collectingMethod id="6ADE4EC17C4F9B18FF67F6B99E57FB41" box="[151,306,1158,1183]" pageId="60" pageNumber="61">Malaise Trap</collectingMethod>
,
<collectorName id="1E6A53007C4F9B18FEB1F6B89EA3FB40" box="[321,454,1158,1183]" pageId="60" pageNumber="61">F. Andrews</collectorName>
&amp;
<collectorName id="1E6A53007C4F9B18FE03F6B89D0CFB40" box="[499,617,1158,1183]" pageId="60" pageNumber="61">A. Gilbert</collectorName>
[col.] (
<collectionCode id="D58EAE137C4F9B18FD34F6B89C7DFB40" box="[708,792,1159,1183]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="60" pageNumber="61">UCDC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4F9B18FCC0F6B99C7AFB38" ID-GBIF-Occurrence="1671715813" collectingDate="1999-09-26" collectingDateMax="1999-09-30" collectingDateMin="1999-09-26" collectionCode="MIUP" collectorName="de Division &amp; Omar Torrijos H &amp; Tr &amp; Malaise, A &amp; Santos, P" country="Taiwan" county="Par" elevation="620" latitude="8.698889" location="General de Division" longLatPrecision="21" longitude="-80.654724" municipality="Nac" pageId="60" pageNumber="61" specimenCount="4" specimenCount-male="4" stateProvince="Cocle Pr" typeStatus="holotype">
<emphasis id="81EBEAC47C4F9B18FCC0F6B99B64FB41" box="[816,1025,1158,1183]" italics="true" pageId="60" pageNumber="61">
<collectingRegion id="715BF8347C4F9B18FCC0F6B99CFCFB41" box="[816,921,1158,1183]" isEnumeration="true" pageId="60" pageNumber="61">
<collectingRegion id="715BF8347C4F9B18FCC0F6B99C14FB40" box="[816,881,1158,1183]" country="Panama" name="Cocle" pageId="60" pageNumber="61">Coclé</collectingRegion>
<collectingRegion id="715BF8347C4F9B18FC8DF6B89CFCFB41" box="[893,921,1159,1182]" country="Brazil" name="Para" pageId="60" pageNumber="61">Pr</collectingRegion>
</collectingRegion>
[ovincia]
</emphasis>
:
<specimenCount id="A599FD5F7C4F9B18FBE4F6B89B5CFB40" box="[1044,1081,1159,1183]" pageId="60" pageNumber="61" type="male">4♂</specimenCount>
, El
<collectingRegion id="715BF8347C4F9B18FB9EF6B99BD7FB40" box="[1134,1202,1158,1183]" country="Panama" name="Cocle" pageId="60" pageNumber="61">Coclé</collectingRegion>
,
<collectingCounty id="5A414E5A7C4F9B18FB32F6B89B8EFB41" box="[1218,1259,1159,1182]" pageId="60" pageNumber="61">Par</collectingCounty>
[que]
<collectingMunicipality id="5344ACAC7C4F9B18FADDF6B99A39FB40" box="[1325,1372,1158,1183]" pageId="60" pageNumber="61">Nac</collectingMunicipality>
[ional
<location id="B640600D7C4F9B18FF67F6959EE0FB1C" LSID="urn:lsid:plazi:treatment:3B3687C07C4F9B18FF67F2A89B6DFAF0:B640600D7C4F9B18FF67F6959EE0FB1C" box="[151,389,1194,1219]" country="Taiwan" county="Par" latitude="8.698889" longLatPrecision="21" longitude="-80.654724" municipality="Nac" name="General de Division" pageId="60" pageNumber="61" stateProvince="Cocle Pr">
General
<collectorName id="1E6A53007C4F9B18FF0CF6959EE0FB1C" box="[252,389,1194,1219]" pageId="60" pageNumber="61">de División</collectorName>
</location>
]
<collectorName id="1E6A53007C4F9B18FE68F6949D38FB1D" box="[408,605,1194,1219]" pageId="60" pageNumber="61">Omar Torrijos H</collectorName>
[errera],
<elevation id="38B2D1E57C4F9B18FD33F6959C2EFB1C" box="[707,843,1194,1219]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="60" pageNumber="61" unit="m" value="620.0" valueMax="670.0" valueMin="570.0">
<quantity id="74679B337C4F9B18FD33F6959C2EFB1C" box="[707,843,1194,1219]" metricMagnitude="2" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="60" pageNumber="61" unit="m" value="620.0" valueMax="670.0" valueMin="570.0">570670 m</quantity>
</elevation>
.s.n.m.[
<geoCoordinate id="D6AB50117C4F9B18FC64F6959B7BFB1C" box="[916,1054,1194,1219]" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="15" value="8.698889">08º41'56&quot;N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C4F9B18FBDFF6949BD9FB1C" box="[1071,1212,1195,1219]" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="15" value="-80.654724">80º39'17&quot;W</geoCoordinate>
],
<collectingDate id="D765E9FE7C4F9B18FB23F6959AFDFB1C" box="[1235,1432,1194,1219]" pageId="60" pageNumber="61" value="1999-09-26" valueMax="1999-09-30" valueMin="1999-09-26">
<date id="C72110167C4F9B18FB23F6959AFDFB1C" box="[1235,1432,1194,1219]" pageId="60" pageNumber="61" value="1999-09-26" valueMax="1999-09-30" valueMin="1999-09-26">2630 sep. 1999</date>
</collectingDate>
,
<collectorName id="1E6A53007C4F9B18FF67F6F09FD6FB39" box="[151,179,1231,1254]" pageId="60" pageNumber="61">Tr</collectorName>
[ampa]
<collectorName id="1E6A53007C4F9B18FEF8F6F19EEDFB38" box="[264,392,1230,1255]" pageId="60" pageNumber="61">Malaise, A</collectorName>
.
<collectorName id="1E6A53007C4F9B18FE65F6F09E9AFB39" box="[405,511,1231,1255]" pageId="60" pageNumber="61">Santos, P</collectorName>
. González [col.] (
<collectionCode id="D58EAE137C4F9B18FD3BF6F09C72FB38" box="[715,791,1231,1255]" pageId="60" pageNumber="61">MIUP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4F9B18FCDCF6F09B6CFAD4" ID-GBIF-Occurrence="1671715799" collectionCode="UCDC" collectorName="R. B. &amp; L. S. Kimsey" country="Panama" latitude="9.15" location="Zone Barro Colorado" longLatPrecision="1300" longitude="-79.85" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Panama Province" typeStatus="holotype">
[
<collectingRegion id="715BF8347C4F9B18FCC5F6F09C9AFB39" box="[821,1023,1231,1254]" country="Panama" name="Panama" pageId="60" pageNumber="61">
<emphasis id="81EBEAC47C4F9B18FCC5F6F09C9AFB39" box="[821,1023,1231,1254]" italics="true" pageId="60" pageNumber="61">
<collectingCountry id="CB8876467C4F9B18FCC5F6F09CF7FB39" box="[821,914,1231,1254]" name="Panama" pageId="60" pageNumber="61">Panama</collectingCountry>
Province
</emphasis>
</collectingRegion>
]:
<specimenCount id="A599FD5F7C4F9B18FBE9F6F09B5BFB38" box="[1049,1086,1231,1255]" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
, C[anal]
<location id="B640600D7C4F9B18FB59F6F09AF9FB38" LSID="urn:lsid:plazi:treatment:3B3687C07C4F9B18FF67F2A89B6DFAF0:B640600D7C4F9B18FB59F6F09AF9FB38" box="[1193,1436,1230,1255]" country="Panama" latitude="9.15" longLatPrecision="1300" longitude="-79.85" name="Zone Barro Colorado" pageId="60" pageNumber="61" stateProvince="Panama Province">Zone Barro Colorado</location>
I[sland], [
<geoCoordinate id="D6AB50117C4F9B18FEF3F6CC9E01FAD4" box="[259,356,1267,1291]" direction="north" orientation="latitude" pageId="60" pageNumber="61" precision="925" value="9.15">09°09'N</geoCoordinate>
,
<geoCoordinate id="D6AB50117C4F9B18FE80F6CD9EB2FAD4" box="[368,471,1266,1291]" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="925" value="-79.85">79°51'W</geoCoordinate>
], V.11.1981,
<collectorName id="1E6A53007C4F9B18FD99F6CC9C3EFAD4" box="[617,859,1266,1291]" pageId="60" pageNumber="61">R. B. &amp; L. S. Kimsey</collectorName>
[col.] (
<collectionCode id="D58EAE137C4F9B18FC5DF6CC9B64FAD4" box="[941,1025,1267,1291]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="60" pageNumber="61">UCDC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4F9B18FF37F7299B6DFAF0" box="[199,1032,1302,1327]" pageId="60" pageNumber="61" type="distribution">
<paragraph id="B32036D67C4F9B18FF37F7299B6DFAF0" blockId="60.[151,1437,258,1327]" box="[199,1032,1302,1327]" pageId="60" pageNumber="61">
<emphasis id="81EBEAC47C4F9B18FF37F7299E3CFAF0" bold="true" box="[199,345,1302,1327]" pageId="60" pageNumber="61">Distribution</emphasis>
(
<figureCitation id="2BA42A537C4F9B18FE98F7299EAEFAF0" box="[360,459,1302,1327]" captionStart="FIGURES 224 233" captionStartId="66.[151,264,1814,1837]" captionTargetBox="[182,1404,199,1768]" captionTargetId="figure@66.[169,1413,193,1773]" captionTargetPageId="66" captionText="FIGURES 224 233. Known distribution of Dissomphalus in Panama. 224. Provinces and Counties of Panama. 1. Bocas del Toro. 2. Chiriquí. 3. Ngöbe-Buglé County. 4. Veraguas. 5. Herrera. 6. Los Santos. 7. Colón. 8. Coclé. 9. Panama. 10. Guna Yala County. 11. Darién. 12. Emberá Woonan County. 225. D. abarus sp. nov., D. afestus sp. nov. and D. altivolans. 226. D. angulatus sp. nov., D. ardonius sp. nov. and D. aretisus sp. nov.. 227. D. apertus, D. areius sp. nov. and D. arteobius sp. nov.. 228. D. arbeius sp. nov., D. asitius sp. nov. and D. brasiliensis. 229. D. arteus sp. nov., D. cervoides and D. culteratus. 230. D. asfartus sp. nov.. 231. D. azagus sp. nov., D. bilobatus and D. coronatus. 232. D. borus sp. nov., D. citus sp. nov. and D. curvifoveatus. 233. D. declinatus, D. dilatatus and D. ettus sp. nov.." httpUri="https://zenodo.org/record/1010056/files/figure.png" pageId="60" pageNumber="61">Fig. 232</figureCitation>
).
<collectingRegion id="715BF8347C4F9B18FE2EF7289D52FAF1" box="[478,567,1303,1326]" country="Panama" name="Panama" pageId="60" pageNumber="61">
<collectingCountry id="CB8876467C4F9B18FE2EF7289D52FAF1" box="[478,567,1303,1326]" name="Panama" pageId="60" pageNumber="61">Panama</collectingCountry>
</collectingRegion>
(
<collectingRegion id="715BF8347C4F9B18FDB5F7299DC2FAF0" box="[581,679,1302,1327]" country="Panama" name="Chiriqui" pageId="60" pageNumber="61">Chiriqui</collectingRegion>
,
<collectingRegion id="715BF8347C4F9B18FD42F7289C79FAF0" box="[690,796,1303,1327]" country="Panama" name="Veraguas" pageId="60" pageNumber="61">Veraguas</collectingRegion>
,
<collectingRegion id="715BF8347C4F9B18FCD7F7299C0DFAF0" box="[807,872,1302,1327]" country="Panama" name="Cocle" pageId="60" pageNumber="61">Coclé</collectingRegion>
and
<collectingRegion id="715BF8347C4F9B18FC6FF7289C98FAF0" box="[927,1021,1303,1327]" country="Panama" name="Panama" pageId="60" pageNumber="61">
<collectingCountry id="CB8876467C4F9B18FC6FF7289C98FAF0" box="[927,1021,1303,1327]" name="Panama" pageId="60" pageNumber="61">Panama</collectingCountry>
</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>