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<document id="BB6595614A801DC83786FE26FCDE651E" ID-CLB-Dataset="31852" ID-DOI="10.11646/zootaxa.4335.1.1" ID-GBIF-Dataset="f2ad3cb1-0bad-4be1-9e36-2170a6a08aaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="1010014" ID-ZooBank="3429592D-23A8-4EBF-8412-BD57B3BC4314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507877826645" checkinUser="plazi" docAuthor="Azevedo, Celso O." docDate="2017" docId="3B3687C07C4B9B1CFF67F2A89D86FC63" docLanguage="en" docName="zootaxa.4335.1.1.pdf" docOrigin="Zootaxa 4335 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Dissomphalus vallensis Evans. Eighteen 1979" docType="treatment" docVersion="15" lastPageNumber="57" masterDocId="C70FFFB87C739B24FFF0F23F9F65FFDF" masterDocTitle="Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="57" updateTime="1698455438235" updateUser="plazi">
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<mods:title id="28E2DBE3336026EB88DCD48F2BACA302">Review of Dissomphalus Ashmead (Hymenoptera, Bethylidae) from Panama, with key to the Central American species</mods:title>
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<treatment id="3B3687C07C4B9B1CFF67F2A89D86FC63" ID-DOI="http://doi.org/10.5281/zenodo.5020510" ID-GBIF-Taxon="135435278" ID-Zenodo-Dep="5020510" LSID="urn:lsid:plazi:treatment:3B3687C07C4B9B1CFF67F2A89D86FC63" httpUri="http://treatment.plazi.org/id/3B3687C07C4B9B1CFF67F2A89D86FC63" lastPageNumber="57" pageId="56" pageNumber="57">
<subSubSection id="FB85655D7C4B9B1CFF67F2A89E26FF0C" pageId="56" pageNumber="57" type="nomenclature">
<paragraph id="B32036D67C4B9B1CFF67F2A89D2BFF6D" blockId="56.[151,590,151,211]" box="[151,590,151,178]" pageId="56" pageNumber="57">
<heading id="E86881BA7C4B9B1CFF67F2A89D2BFF6D" bold="true" box="[151,590,151,178]" fontSize="11" level="1" pageId="56" pageNumber="57" reason="1">
<taxonomicName id="749F4D557C4B9B1CFF67F2A89D2BFF6D" authority="Evans, 1979" authorityName="Evans. Eighteen" authorityYear="1979" box="[151,590,151,178]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="vallensis">
<emphasis id="81EBEAC47C4B9B1CFF67F2A89D2BFF6D" bold="true" box="[151,590,151,178]" pageId="56" pageNumber="57">
<emphasis id="81EBEAC47C4B9B1CFF67F2A89ED4FF6E" bold="true" box="[151,433,151,177]" italics="true" pageId="56" pageNumber="57">Dissomphalus vallensis</emphasis>
<bibRefCitation id="D70E4B277C4B9B1CFE49F2A79D2BFF6D" author="Evans" box="[441,590,152,178]" pageId="56" pageNumber="57" refString="Evans, H. E. (1979) The genus Dissomphalus in Northwestern South America (Hymenoptera, Bethylidae). Proceedings of the Entomological Society of Washington, 81, 276 - 284." type="journal article" year="1979">Evans, 1979</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="B32036D67C4B9B1CFF67F2859E26FF0C" blockId="56.[151,590,151,211]" box="[151,323,186,211]" pageId="56" pageNumber="57">
(
<figureCitation id="2BA42A537C4B9B1CFF6FF2859E5EFF0C" box="[159,315,186,211]" captionStart-0="FIGURES 184 197" captionStart-1="FIGURES 198 210" captionStartId-0="50.[151,264,1612,1635]" captionStartId-1="58.[151,264,1578,1601]" captionTargetBox-0="[159,1430,206,1544]" captionTargetBox-1="[185,1414,210,1557]" captionTargetId-0="figure@50.[151,1436,193,1575]" captionTargetId-1="figure@58.[151,1436,195,1557]" captionTargetPageId-0="50" captionTargetPageId-1="58" captionText-0="FIGURES 184 197. Dissomphalus spp., ♂, genitalia, except otherwise indicated. 184 185. D. sartus sp. nov. 184. Dorsal view. 185. Ventral view. 186 187. D. teobus sp. nov. 186. Dorsal view. 187. Ventral view. 188 189. D. terbius sp. nov. 188. Dorsal view. 189. Ventral view. 190 194. D. unitus 190. Dorsal view. 191. Ventral view. 192. Outer lobe of aedeagal dorsal body, dorsal view. 193. Inner lobe of aedeagal dorsal body, dorsal view. 194. Apical lobe of aedeagal dorsal body, lateral view. 195 196. D. dilatatus 195. Dorsal view. 196. Ventral view. 197. D. vallensis. Dorsal view. Scale bars = 250 µm" captionText-1="FIGURES 198 210. Dissomphalus spp., ♂, genitalia, except figs 198, 204 206 outer lobe of apical lobes of aedeagal dorsal body. 198 199. D. vallensis. 198. Lateral view. 199. Ventral view. 200 201. D. rettenmeyeri 200. Dorsal view. 201. Ventral view. 202 206. D. personatus 202. Dorsal view. 203. Ventral view. 204. Dorsal view. 205. Dorsal view. 206. Lateral view. 207 208. D. abarus sp. nov. 207. Dorsal view. 208. Ventral view. 209 210. D. arteobius sp. nov. 209. Dorsal view. 210. Ventral view. Scale bars = 250 µm." httpUri-0="https://zenodo.org/record/1010048/files/figure.png" httpUri-1="https://zenodo.org/record/1010050/files/figure.png" pageId="56" pageNumber="57">Figs 197199</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4B9B1CFF67F33D9E3CFEE8" pageId="56" pageNumber="57" type="reference_group">
<paragraph id="B32036D67C4B9B1CFF67F33D9E3CFEE8" blockId="56.[151,1436,258,311]" pageId="56" pageNumber="57">
<treatmentCitationGroup id="938F11F87C4B9B1CFF67F33D9E3CFEE8" pageId="56" pageNumber="57">
<emphasis id="81EBEAC47C4B9B1CFF67F33D9E43FEC7" box="[151,294,258,280]" italics="true" pageId="56" pageNumber="57">Dissomphlaus</emphasis>
<taxonomicName id="749F4D557C4B9B1CFEDDF33D9D5BFEC7" authority="Evans, 1979: 277" authorityName="Evans. Eighteen" authorityPageNumber="277" authorityYear="1979" box="[301,574,258,280]" class="Insecta" family="Bethylidae" genus="Dissomphalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="vallensis">
<emphasis id="81EBEAC47C4B9B1CFEDDF33D9EE3FEC7" box="[301,390,258,280]" italics="true" pageId="56" pageNumber="57">vallensis</emphasis>
<treatmentCitation id="323E10C77C4B9B1CFE7BF33C9D5BFEC7" author="Evans" box="[395,574,258,280]" page="277" pageId="56" pageNumber="57" year="1979">
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: 277
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, 281 (♂ holotype from USNM);
<treatmentCitation id="323E10C77C4B9B1CFC7BF33D9BC8FEC7" author="Gordh" box="[907,1197,258,280]" page="226" pageId="56" pageNumber="57" year="1990">
<bibRefCitation id="D70E4B277C4B9B1CFC7BF33D9B1FFEC7" author="Gordh" box="[907,1146,258,280]" pageId="56" pageNumber="57" refString="Gordh, G. &amp; Moczar, L. (1990) A catalog of the world Bethylidae (Hymenoptera). Memoirs of the American Entomological Institute, 46, 1 - 364." type="journal article" year="1990">Gordh &amp; Móczár, 1990</bibRefCitation>
: 226
</treatmentCitation>
;
<treatmentCitation id="323E10C77C4B9B1CFB48F33D9AEAFEC7" author="Azevedo" box="[1208,1423,258,280]" page="327" pageId="56" pageNumber="57" year="1999">
<bibRefCitation id="D70E4B277C4B9B1CFB48F33D9A05FEC7" author="Azevedo" box="[1208,1376,258,280]" pageId="56" pageNumber="57" refString="Azevedo, C. O. (1999 a) Revision of the Neotropical Dissomphalus Ashmead, 1893 (Hymenoptera, Bethylidae) with median tergal process. Arquivos de Zoologia, 35 (4), 301 - 394. https: // doi. org / 10.11606 / issn. 2176 - 7793. v 35 i 4 p 301 - 394" type="journal article" year="1999" yearSuffix="a">Azevedo, 1999a</bibRefCitation>
: 327
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328, 162163.
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</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4B9B1CFF67F35B9E1DFD8A" pageId="56" pageNumber="57" type="diagnosis">
<paragraph id="B32036D67C4B9B1CFF67F35B9E1DFD8A" blockId="56.[151,1437,356,957]" pageId="56" pageNumber="57">
<emphasis id="81EBEAC47C4B9B1CFF67F35B9E74FEA2" bold="true" box="[151,273,356,381]" pageId="56" pageNumber="57">Diagnosis.</emphasis>
Mandible bidentate. Clypeus wih median lobe ill-defined. Tergal process median with large, elliptical and deep depression with pair of tubercles with tuft of setae directed each other. Hypopygium with posterior margin concave. Genitalia: paramere with apex truncate and oblique; aedeagal ventral ramus as long as aedeagal dorsal body; surface smooth; inner margin straight; outer margin convex; apical half narrow and divergent; aedeagal dorsal body with apical half arched downward; ventral side with median fold; dorsal margin of apical half arched outward and pair of very small lobes upward in base of aedeagal dorsal body. Apodeme not extending beyond genital ring.
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4B9B1CFF37F0609C3DFCD7" pageId="56" pageNumber="57" type="discussion">
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<emphasis id="81EBEAC47C4B9B1CFF37F0609E5EFDA7" bold="true" box="[199,315,607,632]" pageId="56" pageNumber="57">Remarks.</emphasis>
This species is known from
<collectingCountry id="CB8876467C4B9B1CFD63F0609C64FDA7" box="[659,769,607,632]" name="Colombia" pageId="56" pageNumber="57">Colombia</collectingCountry>
(
<bibRefCitation id="D70E4B277C4B9B1CFCE3F0609CAFFDA7" author="Azevedo" box="[787,970,607,632]" pageId="56" pageNumber="57" refString="Azevedo, C. O. (1999 a) Revision of the Neotropical Dissomphalus Ashmead, 1893 (Hymenoptera, Bethylidae) with median tergal process. Arquivos de Zoologia, 35 (4), 301 - 394. https: // doi. org / 10.11606 / issn. 2176 - 7793. v 35 i 4 p 301 - 394" type="journal article" year="1999" yearSuffix="a">Azevedo 1999a</bibRefCitation>
) and is recorded for the first time from
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<collectingCountry id="CB8876467C4B9B1CFF67F0BA9F96FD43" box="[151,243,645,668]" name="Panama" pageId="56" pageNumber="57">Panama</collectingCountry>
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. Some variations are worth to mention: median clypeal tooth angulate; median clypeal carina conspicuous and complete; vertex crest straight; anterior margin of pronotal disc coarse; aedeagal ventral ramus shorter than aedeagal dorsal body; median region of aedeagal dorsal body wide, apical portion elongated and narrow, outer margin sinuous; genital ring bifurcated distally in ventral view.
</paragraph>
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<collectingMunicipality id="5344ACAC7C4B9B1CFF37F12B9E4AFCF2" box="[199,303,788,813]" pageId="56" pageNumber="57">Material</collectingMunicipality>
examined.
</emphasis>
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,
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,
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<emphasis id="81EBEAC47C4B9B1CFCF2F12A9CCEFCF2" box="[770,939,788,813]" italics="true" pageId="56" pageNumber="57">Peñas Blancas</emphasis>
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:
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,
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, Dept.
<collectingRegion id="715BF8347C4B9B1CFB30F12B9B99FCF2" box="[1216,1276,788,813]" country="Honduras" name="Valle" pageId="56" pageNumber="57">Valle</collectingRegion>
, 2628 1975,
<collectingMethod id="6ADE4EC17C4B9B1CFF67F1089E40FC90" box="[151,293,823,848]" pageId="56" pageNumber="57">Malaise trap</collectingMethod>
,
<collectorName id="1E6A53007C4B9B1CFEC2F1079EADFC8F" box="[306,456,823,848]" pageId="56" pageNumber="57">R. Wilkerson</collectorName>
(
<collectionCode id="D58EAE137C4B9B1CFE27F1079D46FC8F" box="[471,547,824,848]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="56" pageNumber="57">FSCA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4B9B1CFDC7F1079B9AFCAB" ID-GBIF-Occurrence="1671715749" collectingDate="1994-11-19" collectingDateMax="1994-11-20" collectingDateMin="1994-11-19" collectionCode="CNCI" collectorName="El Cope &amp; D. Windson Edwards" country="Panama" county="Coche" elevation="730" latitude="8.619722" location="El Cope" longLatPrecision="21" longitude="-80.58417" municipality="El Cope" pageId="56" pageNumber="57" specimenCount="1" specimenCount-male="1" stateProvince="Cocle" typeStatus="holotype">
<collectingCountry id="CB8876467C4B9B1CFDC7F1079DD4FC8F" box="[567,689,824,848]" name="Panama" pageId="56" pageNumber="57">PANAMA</collectingCountry>
,
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{=
<collectingRegion id="715BF8347C4B9B1CFCD8F1089C09FC8F" box="[808,876,823,848]" country="Panama" name="Cocle" pageId="56" pageNumber="57">Coclé</collectingRegion>
}
</emphasis>
:
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,
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<collectingMunicipality id="5344ACAC7C4B9B1CFC49F1089B70FC8F" box="[953,1045,823,848]" pageId="56" pageNumber="57">El Cope</collectingMunicipality>
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{=
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<collectorName id="1E6A53007C4B9B1CFBC6F1089BFBFC8F" box="[1078,1182,823,848]" pageId="56" pageNumber="57">El Copé</collectorName>
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</elevation>
, [
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,
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],
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</collectingDate>
, F[light] I[nterception] T[rap],
<collectorName id="1E6A53007C4B9B1CFCAAF1629B36FCAA" box="[858,1107,860,885]" pageId="56" pageNumber="57">D. Windson Edwards</collectorName>
[col.] (
<collectionCode id="D58EAE137C4B9B1CFB5EF1639B92FCAB" box="[1198,1271,860,884]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="56" pageNumber="57">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="03F73C8B7C4B9B1CFAFFF1629AF9FC47" ID-GBIF-Occurrence="1671715765" collectingDate="1995-06-04" collectingDateMax="1995-06-07" collectingDateMin="1995-06-04" collectionCode="UFES" collectorName="de Pirre &amp; J. Ashe &amp; R. Brooks" country="Panama" county="Cana" elevation="1450" location="Serrania" municipality="Serraria" pageId="56" pageNumber="57" specimenCount="5" specimenCount-male="5" stateProvince="Darien" typeStatus="holotype">
<collectingRegion id="715BF8347C4B9B1CFAFFF1629A3BFCAB" box="[1295,1374,861,884]" country="Panama" name="Darien" pageId="56" pageNumber="57">
<emphasis id="81EBEAC47C4B9B1CFAFFF1629A3BFCAB" box="[1295,1374,861,884]" italics="true" pageId="56" pageNumber="57">Darién</emphasis>
</collectingRegion>
:
<specimenCount id="A599FD5F7C4B9B1CFA82F1639AF2FCAA" box="[1394,1431,860,885]" pageId="56" pageNumber="57" type="male">5♂</specimenCount>
,
<collectingCounty id="5A414E5A7C4B9B1CFF67F1BF9FB6FC47" box="[151,211,896,920]" pageId="56" pageNumber="57">Cana</collectingCounty>
,
<collectingMunicipality id="5344ACAC7C4B9B1CFF2EF1409E52FC47" box="[222,311,895,920]" pageId="56" pageNumber="57">Serraria</collectingMunicipality>
{=
<location id="B640600D7C4B9B1CFEA5F1409ED8FC47" LSID="urn:lsid:plazi:treatment:3B3687C07C4B9B1CFF67F2A89D86FC63:B640600D7C4B9B1CFEA5F1409ED8FC47" box="[341,445,895,920]" country="Panama" county="Cana" municipality="Serraria" name="Serrania" pageId="56" pageNumber="57" stateProvince="Darien">Serrania</location>
}
<collectorName id="1E6A53007C4B9B1CFE3BF1409D47FC47" box="[459,546,895,920]" pageId="56" pageNumber="57">de Pirre</collectorName>
<elevation id="38B2D1E57C4B9B1CFDD9F1409D1CFC47" box="[553,633,895,920]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="56" pageNumber="57" unit="m" value="1450.0">
<quantity id="74679B337C4B9B1CFDD9F1409D1CFC47" box="[553,633,895,920]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="56" pageNumber="57" unit="m" value="1450.0">1450m</quantity>
</elevation>
,
<collectingDate id="D765E9FE7C4B9B1CFD74F1409C77FC47" box="[644,786,895,920]" pageId="56" pageNumber="57" value="1995-06-04" valueMax="1995-06-07" valueMin="1995-06-04">
<date id="C72110167C4B9B1CFD74F1409C77FC47" box="[644,786,895,920]" pageId="56" pageNumber="57" value="1995-06-04" valueMax="1995-06-07" valueMin="1995-06-04">47.vi. 1995</date>
</collectingDate>
,
<collectorName id="1E6A53007C4B9B1CFCEDF1BF9C14FC47" box="[797,881,895,920]" pageId="56" pageNumber="57">J. Ashe</collectorName>
,
<collectorName id="1E6A53007C4B9B1CFC8DF1BF9C8BFC47" box="[893,1006,895,920]" pageId="56" pageNumber="57">R. Brooks</collectorName>
F[light] I[nterception] T[rap] (
<collectionCode id="D58EAE137C4B9B1CFAB6F1BF9AF5FC47" box="[1350,1424,896,920]" country="Brazil" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="56" pageNumber="57">UFES</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FB85655D7C4B9B1CFF37F19B9D86FC63" box="[199,739,932,957]" pageId="56" pageNumber="57" type="distribution">
<paragraph id="B32036D67C4B9B1CFF37F19B9D86FC63" blockId="56.[151,1437,356,957]" box="[199,739,932,957]" pageId="56" pageNumber="57">
<emphasis id="81EBEAC47C4B9B1CFF37F19B9E3CFC62" bold="true" box="[199,345,932,957]" pageId="56" pageNumber="57">Distribution</emphasis>
(
<figureCitation id="2BA42A537C4B9B1CFE98F19B9EAEFC63" box="[360,459,932,957]" captionStart="FIGURES 244 249" captionStartId="68.[151,264,1212,1235]" captionTargetBox="[156,1430,201,1176]" captionTargetId="figure@68.[148,1433,193,1184]" captionTargetPageId="68" captionText="FIGURES 244 249. Known distribution of Dissomphalus in Panama. 244. D. rufipalpis, D. teraus sp. nov. and D. vallensis. 245. D. sartus sp. nov. and D. trisus sp. nov.. 246. D. teobus sp. nov. and D. urazius sp. nov.. 247. D. terbius sp. nov. and D. unitus. 248. D. ubracus sp. nov.. 249. D. rettenmeyeri." httpUri="https://zenodo.org/record/1010062/files/figure.png" pageId="56" pageNumber="57">Fig. 244</figureCitation>
).
<collectingCountry id="CB8876467C4B9B1CFE2EF19B9D29FC62" box="[478,588,932,957]" name="Colombia" pageId="56" pageNumber="57">Colombia</collectingCountry>
and
<collectingRegion id="715BF8347C4B9B1CFD73F19A9DBAFC63" box="[643,735,933,956]" country="Panama" name="Panama" pageId="56" pageNumber="57">
<collectingCountry id="CB8876467C4B9B1CFD73F19A9DBAFC63" box="[643,735,933,956]" name="Panama" pageId="56" pageNumber="57">Panama</collectingCountry>
</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>