treatments-xml/data/03/B9/30/03B9306BF076ED06A573373B4459E577.xml
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<document id="474CB94D2FE71E8A4F43597A60B54807" ID-DOI="10.5281/zenodo.1038228" ID-GBIF-Dataset="6116db9f-6823-4432-98f6-eabc9faa978b" ID-Zenodo-Dep="1038228" IM.taxonomicNames_approvedBy="jeremy" approvalRequired="7" approvalRequired_for_document="1" approvalRequired_for_textStreams="6" checkinTime="1509025528795" checkinUser="jeremy" docAuthor="Olshevsky, G." docDate="1995" docId="03B9306BF076ED06A573373B4459E577" docLanguage="en" docName="Olshevsky_Ford_1995_Dinotyrannus_Stygivenator(almost done).pdf.imf" docOrigin="Dino-Frontline 9" docStyle="DocumentStyle{}" docTitle="Stygivenator cristatus Olshevsky, 1995, n. comb." docType="treatment" docVersion="8" lastPageNumber="119" masterDocId="FF804813F077ED07A33435294264E000" masterDocTitle="The origin and evolution of the Tyrannosaurids, Part 1" masterLastPageNumber="119" masterPageNumber="92" pageNumber="92" updateTime="1695395669354" updateUser="jeremy">
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<mods:titleInfo id="A7292DC5C2F024D389021163F7631ED7">
<mods:title id="CDAEFEE586F87A19AB02EEC4DA0D1F8D">The origin and evolution of the Tyrannosaurids, Part 1</mods:title>
</mods:titleInfo>
<mods:name id="ECB240D7CCBE7178B113806E9846ACE7" type="personal">
<mods:role id="72A3DC711391721546417848532D1685">
<mods:roleTerm id="31EED8E2009E463DABF4A71237CEF5F4">Author</mods:roleTerm>
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<mods:namePart id="0CC66EE97F1B60CD09015FF6EF785953">Olshevsky, G.</mods:namePart>
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<mods:typeOfResource id="0AD2E5ADCE866154854E16ED68F306E6">text</mods:typeOfResource>
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<mods:title id="B43A570062C320176800F40142C0EFF9">Dino-Frontline</mods:title>
</mods:titleInfo>
<mods:part id="A9D6DA2BD0A1D011BA10436A5E7E8FB0">
<mods:date id="6CF5262D88E0EDA84614F3D27536B10D">1995</mods:date>
<mods:detail id="A3AB0E574DCBBFE019B8FCE6E9596AD1" type="volume">
<mods:number id="8F0EC8A9645723BFF60A2EB6E964BA8A">9</mods:number>
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<mods:classification id="F62754820BF68DA1A4F4B0A5E5786370">journal article</mods:classification>
<mods:identifier id="756C2A8FE9D8BE9C513AB1DDA7ED2124" type="DOI">10.5281/zenodo.1038228</mods:identifier>
<mods:identifier id="67D9D1A914489C097F04209B5AB76300" type="GBIF-Dataset">6116db9f-6823-4432-98f6-eabc9faa978b</mods:identifier>
<mods:identifier id="64D9B08DB79E9BE8173FE7E06AB18AE9" type="Zenodo-Dep">1038228</mods:identifier>
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<treatment id="03B9306BF076ED06A573373B4459E577" ID-DOI="http://doi.org/10.5281/zenodo.3483053" ID-GBIF-Taxon="193436296" ID-Zenodo-Dep="3483053" LSID="urn:lsid:plazi:treatment:03B9306BF076ED06A573373B4459E577" httpUri="http://treatment.plazi.org/id/03B9306BF076ED06A573373B4459E577" pageId="1">
<subSubSection id="C30AD2F6F076ED06A573373B403FE267" pageId="1" type="nomenclature">
<paragraph id="8BAF817DF076ED06A573373B403FE267" blockId="1.[341,2282,435,615]" pageId="1">
<emphasis id="B9645D6FF076ED06A573373B4432E238" box="[1607,1622,530,568]" italics="true" pageId="1">?</emphasis>
<taxonomicName id="4C10FAFEF076ED06A56D373B4AA1E238" baseAuthorityName="Marsh" baseAuthorityYear="1892" box="[1625,2245,530,568]" class="Reptilia" family="Tyrannosauridae" genus="Stygivenator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="species" species="cristatus" status="n. comb.">
<emphasis id="B9645D6FF076ED06A56D373B45AEE238" box="[1625,1994,530,568]" italics="true" pageId="1">Stygivenator cristatus</emphasis>
(Marsh, 1892)
</taxonomicName>
<taxonomicNameLabel id="A257E014F076ED06ABE4373B4031E267" pageId="1">n. comb.</taxonomicNameLabel>
,
</paragraph>
</subSubSection>
<subSubSection id="C30AD2F6F076ED06A15D37684A40E267" box="[617,2084,577,615]" pageId="1" type="reference_group">
<paragraph id="8BAF817DF076ED06A15D37684A40E267" blockId="1.[341,2282,435,615]" box="[617,2084,577,615]" pageId="1">
formerly
<treatmentCitation id="0AB1A76CF076ED06A03437684724E267" authorityName="Marsh" authorityYear="1892" box="[768,1344,577,615]" httpUri="http://treatment.plazi.org/id/03B58786-FFCE-FFC6-48A6-F7C6FB32D794" pageId="1">
<taxonomicName id="4C10FAFEF076ED06A03437684724E267" authorityName="Marsh" authorityYear="1892" box="[768,1344,577,615]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="species" species="cristatus">
<emphasis id="B9645D6FF076ED06A03437684600E267" box="[768,1124,577,615]" italics="true" pageId="1">Aublysodon cristatus</emphasis>
Marsh, 1892
</taxonomicName>
</treatmentCitation>
(=
<emphasis id="B9645D6FF076ED06A648376844B0E267" box="[1404,1748,577,615]" italics="true" pageId="1">
<taxonomicName id="4C10FAFEF076ED06A648376844A5E267" baseAuthorityName="Marsh" baseAuthorityYear="1892" box="[1404,1729,577,615]" class="Reptilia" family="Tyrannosauridae" genus="Deinodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="species" species="cristatus">Deinodon crlstatus</taxonomicName>
),
</emphasis>
See text for details.
</paragraph>
</subSubSection>
<subSubSection id="C30AD2F6F076ED06A26037FD4459E577" pageId="1" type="discussion">
<paragraph id="8BAF817DF076ED06A26037FD46BBE486" blockId="1.[340,2292,724,1399]" pageId="1">
Discussion: Since
<taxonomicName id="4C10FAFEF076ED06A1BB37FD4101E2FA" authorityName="Olshevsky" authorityYear="1995" box="[655,869,724,762]" class="Reptilia" family="Tyrannosauridae" genus="Stygivenator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="genus">
<emphasis id="B9645D6FF076ED06A1BB37FD4101E2FA" box="[655,869,724,762]" italics="true" pageId="1">Stygivenator</emphasis>
</taxonomicName>
is so strikingly different from other tyrannosaurids, it is first necessary to justify its inclusion in
<taxonomicName id="4C10FAFEF076ED06A18E362C41BDE32B" authorityName="Osborn" authorityYear="1906" box="[698,985,773,811]" class="Reptilia" family="Tyrannosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="family">Tyrannosauridae</taxonomicName>
. Naturally,
<taxonomicName id="4C10FAFEF076ED06A7A3362C4709E32B" authorityName="Olshevsky" authorityYear="1995" box="[1175,1389,773,811]" class="Reptilia" family="Tyrannosauridae" genus="Stygivenator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="genus">
<emphasis id="B9645D6FF076ED06A7A3362C4709E32B" box="[1175,1389,773,811]" italics="true" pageId="1">Stygivenator</emphasis>
</taxonomicName>
differs strongly enough from the “orthodox” tyrannosaurids that at least subfamilial separation
<emphasis id="B9645D6FF076ED06A61B361F472EE35C" box="[1327,1354,822,860]" italics="true" pageId="1">is</emphasis>
warranted. But the smooth, constricted nasals; low, pointed maxillae; long frontals; unsenrated D-sectioned premaxillary tooth; relatively slender dentaxy; and lateral buttress above the surangular foramen are
<emphasis id="B9645D6FF076ED06A61736B64751E3C5" bold="true" box="[1315,1333,927,965]" pageId="1">a</emphasis>
combination of characters present in
<taxonomicName id="4C10FAFEF076ED06A49F36B64AE7E3C5" authorityName="Olshevsky" authorityYear="1995" box="[1963,2179,927,965]" class="Reptilia" family="Tyrannosauridae" genus="Stygivenator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="genus">
<emphasis id="B9645D6FF076ED06A49F36B64AE7E3C5" box="[1963,2179,927,965]" italics="true" pageId="1">Stygivenator</emphasis>
</taxonomicName>
that are diagnostic of the tyrannosaurid parasubfamily
<taxonomicName id="4C10FAFEF076ED06A60636E64414E3F5" authorityName="Olshevsky" authorityYear="1995" box="[1330,1648,975,1013]" class="Reptilia" family="Tyrannosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="subFamily" subFamily="Shanshanosaurinae">Shanshanosaurinae</taxonomicName>
(= Aublysodontidae [
<emphasis id="B9645D6FF076ED06A4EA36E64ABEE3F5" box="[2014,2266,975,1013]" italics="true" pageId="1">nomen dubium</emphasis>
] of authors). The surangular buttress and morphology of the articulation point of the dentary in particular are quite similar to that of
<emphasis id="B9645D6FF076ED06A05C311944C0E456" box="[872,1700,1072,1110]" italics="true" pageId="1">
<taxonomicName id="4C10FAFEF076ED06A05C311947D9E456" box="[872,1469,1072,1110]" class="Reptilia" family="Tyrannosauridae" genus="Shanshanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="species" species="huoyanshanensls">Shanshanosaurus huoyanshanensls</taxonomicName>
,
<taxonomicName id="4C10FAFEF076ED06A6F9311944C0E456" authorityName="Olshevsky" authorityYear="1995" box="[1485,1700,1072,1110]" class="Reptilia" family="Tyrannosauridae" genus="Stygivenator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="genus">Stygivenator</taxonomicName>
</emphasis>
is thus best viewed as a highly derived, late-occurring member of that taxon.
</paragraph>
<paragraph id="8BAF817DF076ED06A2DE31E84459E577" blockId="1.[340,2292,724,1399]" pageId="1">
Most of the diagnostic characters may be found in the description of this genus (as the “Jordan theropod” and as
<emphasis id="B9645D6FF076ED06A16931D84150E517" box="[605,820,1265,1303]" italics="true" pageId="1">
<taxonomicName id="4C10FAFEF076ED06A16931D8414CE517" authorityName="Leidy" authorityYear="1868" box="[605,808,1265,1303]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" phylum="Chordata" rank="genus">Aublysodon</taxonomicName>
)
</emphasis>
by Molnar (1978) and Molnar &amp; Carpenter (1989). The specimen was personally examined by the author on a visit to the University of California at Berkeley Museum of Paleontology in the early 1980s, when the specimen was on loan to that institution.
</paragraph>
</subSubSection>
</treatment>
</document>