243 lines
30 KiB
XML
243 lines
30 KiB
XML
<document id="DC83802EE864DA84D7F53DE2F0A2B195" ID-CLB-Dataset="29087" ID-DOI="10.11646/zootaxa.4455.3.4" ID-GBIF-Dataset="c9bf5059-abe4-414e-a7c2-f3f9e75f2979" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457467" ID-ZooBank="69209FF7-80B0-409D-A257-0B88C65E903F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539262799757" checkinUser="plazi" docAuthor="Kamiński, Marcin J. & Schoeman, Colin S." docDate="2018" docId="038BC66EFFD69650F7ABFD42FB11FE02" docLanguage="en" docName="zootaxa.4455.3.4.pdf" docOrigin="Zootaxa 4455 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anaxius bloubergensis Kamiński & Schoeman 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="475" masterDocId="FFB2BE16FFD49654F73CFFF7FF95FFF2" masterDocTitle="Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina)" masterLastPageNumber="485" masterPageNumber="471" pageNumber="473" updateTime="1698576062251" updateUser="ExternalLinkService">
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<mods:mods id="5A8EDCBB65BEFD6467A4CC1536B8D783" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3B6D210FDFC2D0E47F48252E92F1FA5D">
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<mods:title id="3D6E8CAE011F98B71ABD94AADF520BEC">Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina)</mods:title>
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<mods:name id="CC40F7D4669DB96FB15B6028098B74D0" type="personal">
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<mods:namePart id="C4FAC9171DE85C553B09FCB1CD69A9F2">Kamiński, Marcin J.</mods:namePart>
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<mods:roleTerm id="5B5BB0D7A6F5EEAB1197C2C1F25FBA2B">Author</mods:roleTerm>
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<mods:namePart id="92101EF69B91A4B0F0A98C006B04127C">Schoeman, Colin S.</mods:namePart>
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<mods:title id="60ACA7B47B065E30F22FD9686D89CDC7">Zootaxa</mods:title>
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<mods:part id="0787F63A2C0A8F4E4DB3E7890B3AE053">
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<mods:date id="22E72BA14124BC7D0C24DF0547D5F88A">2018</mods:date>
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<mods:number id="1FF3AD34DA549788C5C407A7D65AA85C">2018-08-03</mods:number>
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<mods:number id="7E003AC1354FEFADB04F863FA8B917F7">4455</mods:number>
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<treatment id="038BC66EFFD69650F7ABFD42FB11FE02" ID-DOI="http://doi.org/10.5281/zenodo.5975953" ID-GBIF-Taxon="148691418" ID-Zenodo-Dep="5975953" LSID="urn:lsid:plazi:treatment:038BC66EFFD69650F7ABFD42FB11FE02" httpUri="http://treatment.plazi.org/id/038BC66EFFD69650F7ABFD42FB11FE02" lastPageId="4" lastPageNumber="475" pageId="2" pageNumber="473">
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<subSubSection id="C33824F3FFD69656F7ABFD42FB1CFD02" pageId="2" pageNumber="473" type="nomenclature">
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<paragraph id="8B9D7778FFD69656F7ABFD42FD85FD3D" blockId="2.[151,1161,693,789]" box="[151,528,693,719]" pageId="2" pageNumber="473">
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<heading id="D0D5C014FFD69656F7ABFD42FD85FD3D" bold="true" box="[151,528,693,719]" fontSize="11" level="1" pageId="2" pageNumber="473" reason="1">
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<emphasis id="B956AB6AFFD69656F7ABFD42FD85FD3D" bold="true" box="[151,528,693,719]" pageId="2" pageNumber="473">
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<taxonomicName id="4C220CFBFFD69656F7ABFD42FE3EFD3D" authority="Kamiński & Schoeman, 2018" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[151,427,693,719]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="473" phylum="Arthropoda" rank="species" species="bloubergensis" status="sp. nov.">
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<emphasis id="B956AB6AFFD69656F7ABFD42FE3EFD3D" bold="true" box="[151,427,693,719]" italics="true" pageId="2" pageNumber="473">Anaxius bloubergensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2651611FFD69656F68EFD42FD85FD3D" box="[434,528,693,719]" pageId="2" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8B9D7778FFD69656F7ABFD20FB1CFD02" blockId="2.[151,1161,693,789]" box="[151,1161,727,752]" pageId="2" pageNumber="473">http://zoobank.org/urn:lsid:zoobank.org:act:81D10A96-9238-4501-AF69-048DFF06453F</paragraph>
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</subSubSection>
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<subSubSection id="C33824F3FFD69656F7ABFD0BFE0BFCE7" box="[151,414,764,789]" pageId="2" pageNumber="473" type="description">
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<paragraph id="8B9D7778FFD69656F7ABFD0BFE0BFCE7" blockId="2.[151,1161,693,789]" box="[151,414,764,789]" pageId="2" pageNumber="473">
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(
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||
<figureCitation id="13196BFDFFD69656F7A3FD0BFE96FCE6" box="[159,259,764,789]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="2" pageNumber="473">Figs. 1A</figureCitation>
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,
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<figureCitation id="13196BFDFFD69656F632FD0BFED9FCE6" box="[270,332,764,788]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[151,1435,681,1847]" captionTargetId="figure@5.[151,1435,681,1847]" captionTargetPageId="5" captionText="FIGURE 2. Diagnostic characters proposed for Anaxius: mentum (A), proleg (B, G), mesoleg (C, F), metaleg (D), metaventrite (H, I). Anaxius bloubergensis (E, F), A. campbellae (A, H), A. meletsensis (G, I), and A. prozeskyi (B, C, D). Abbreviations: d—denticles, ld—surface between labial palpi, lk—longitudinal keel, lw—lateral wing, p—palpifer, t—apical denticle, tub—tubercles." httpUri="https://zenodo.org/record/1457471/files/figure.png" pageId="2" pageNumber="473">2E, F</figureCitation>
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,
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<figureCitation id="13196BFDFFD69656F66BFD0BFEEEFCE6" box="[343,379,764,788]" captionStart="FIGURE 3" captionStartId="7.[151,250,1440,1462]" captionTargetBox="[200,1386,725,1434]" captionTargetId="figure@7.[200,1386,725,1434]" captionTargetPageId="7" captionText="FIGURE 3. Aedeagal tegmen morphology. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457473/files/figure.png" pageId="2" pageNumber="473">3A</figureCitation>
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,
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<figureCitation id="13196BFDFFD69656F6BBFD0BFE00FCE7" box="[391,405,764,789]" captionStart="FIGURE 5" captionStartId="11.[151,250,1731,1753]" captionTargetBox="[161,1437,844,1704]" captionTargetId="figure@11.[151,1437,844,1710]" captionTargetPageId="11" captionText="FIGURE 5. Distribution of the species representing Anaxius." httpUri="https://zenodo.org/record/1457477/files/figure.png" pageId="2" pageNumber="473">5</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33824F3FFD69656F7ABFCB3FB9DFC1E" pageId="2" pageNumber="473" type="materials_examined">
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<paragraph id="8B9D7778FFD69656F7ABFCB3FB9DFC1E" blockId="2.[151,1437,836,2013]" pageId="2" pageNumber="473">
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<materialsCitation id="3B4A7D25FFD69656F7ABFCB3FC34FC72" ID-GBIF-Occurrence="1917075943" collectingDate="2016-11-27" collectionCode="TMSA" collectorName="S. Afr. & Ruth Muller" latitude="-22.59" location="Limpopo" longLatPrecision="755" longitude="29.08" pageId="2" pageNumber="473" specimenCount="2" stateProvince="Limpopo" typeStatus="holotype">
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||
<emphasis id="B956AB6AFFD69656F7ABFCB3FE0DFCAF" bold="true" box="[151,408,836,861]" pageId="2" pageNumber="473">
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<typeStatus id="5499C9DAFFD69656F7ABFCB3FF47FCAF" box="[151,210,836,861]" pageId="2" pageNumber="473">Type</typeStatus>
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data.
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||
<typeStatus id="5499C9DAFFD69656F617FCB3FE0DFCAF" box="[299,408,836,861]" pageId="2" pageNumber="473" type="holotype">Holotype</typeStatus>
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</emphasis>
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, male (
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||
<collectionCode id="ED33EFBDFFD69656F6C7FCB3FDC5FCAE" box="[507,592,836,860]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="2" pageNumber="473">TMSA</collectionCode>
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): “
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||
<collectorName id="26D712AEFFD69656F54BFCB3FD59FCAF" box="[631,716,836,861]" pageId="2" pageNumber="473">S. Afr.</collectorName>
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||
;
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<collectingRegion id="49E6B99AFFD69656F5E2FCB3FCD0FCAF" box="[734,837,836,861]" country="South Africa" name="Limpopo" pageId="2" pageNumber="473">Limpopo</collectingRegion>
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Prov. / Blouberg Nat. Res. /
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||
<geoCoordinate id="EE1611BFFFD69656F382FCB3FA8CFCAE" box="[1214,1305,836,861]" direction="south" orientation="latitude" pageId="2" pageNumber="473" precision="555" value="-22.59">22.59 S</geoCoordinate>
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||
–
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||
<geoCoordinate id="EE1611BFFFD69656F215FCB3FA19FCAE" box="[1321,1420,836,860]" direction="east" orientation="longitude" pageId="2" pageNumber="473" precision="555" value="29.08">29.08 E</geoCoordinate>
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||
”, “25–
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||
<collectingDate id="EFD8A850FFD69656F7F6FC90FED9FC72" box="[202,332,871,896]" pageId="2" pageNumber="473" value="2016-11-27">
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||
<date id="FF9C51B8FFD69656F7F6FC90FED9FC72" box="[202,332,871,896]" pageId="2" pageNumber="473" value="2016-11-27">27.11.2016</date>
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||
</collectingDate>
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||
E-Y:3987 / day, sandy bushveld / leg.
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<collectorName id="26D712AEFFD69656F439FC90FC02FC72" box="[773,919,871,896]" pageId="2" pageNumber="473">Ruth Müller</collectorName>
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||
”
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B4A7D25FFD69656F491FC90FF68FC57" ID-GBIF-Occurrence="1917075902" collectingDate="2016-11-27" collectionCode="TMSA" collectorName="S. Afr. & Ruth Muller" latitude="-22.59" location="Limpopo" longLatPrecision="755" longitude="29.08" pageId="2" pageNumber="473" specimenCount="3" specimenCount-male="3" stateProvince="Limpopo" typeStatus="paratype">
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||
<typeStatus id="5499C9DAFFD69656F491FC90FBB1FC72" box="[941,1060,871,896]" pageId="2" pageNumber="473" type="paratype">
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<emphasis id="B956AB6AFFD69656F491FC90FBB1FC72" bold="true" box="[941,1060,871,896]" pageId="2" pageNumber="473">Paratypes</emphasis>
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</typeStatus>
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||
,
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<specimenCount id="9D24BCF1FFD69656F30AFC9FFB1BFC72" box="[1078,1166,871,896]" pageId="2" pageNumber="473" type="male">3 males</specimenCount>
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||
(
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<collectionCode id="ED33EFBDFFD69656F39CFC9FFB61FC72" box="[1184,1268,872,896]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="2" pageNumber="473">TMSA</collectionCode>
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||
): same data as holotype
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</materialsCitation>
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||
;
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<materialsCitation id="3B4A7D25FFD69656F633FC7BFB9DFC1E" ID-GBIF-Occurrence="1917075936" collectingDate="2012-06-12" collectionCode="TMSA, MIZ, PAN" collectorName="S. Afr. & NR North & Roodeberg Bushveld & Colin Schoeman & Blouberg NR & Rodeberg Bushveld" elevation="866" location="Limpopo" longitude="29.12" pageId="2" pageNumber="473" specimenCount="1" stateProvince="Limpopo" typeStatus="paratype">
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male (
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<collectionCode id="ED33EFBDFFD69656F660FC7BFE38FC56" box="[348,429,908,932]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="2" pageNumber="473">TMSA</collectionCode>
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||
) and male (
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<collectionCode id="ED33EFBDFFD69656F57AFC7BFDEEFC56" box="[582,635,908,932]" pageId="2" pageNumber="473">MIZ</collectionCode>
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||
<collectionCode id="ED33EFBDFFD69656F5B6FC7BFD53FC56" box="[650,710,908,932]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13841" name="Panjab University" pageId="2" pageNumber="473">PAN</collectionCode>
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||
): “
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||
<collectorName id="26D712AEFFD69656F5D0FC7BFCD4FC57" box="[748,833,908,933]" pageId="2" pageNumber="473">S. Afr.</collectorName>
|
||
;
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||
<collectingRegion id="49E6B99AFFD69656F46FFC7BFC2FFC57" box="[851,954,908,933]" country="South Africa" name="Limpopo" pageId="2" pageNumber="473">Limpopo</collectingRegion>
|
||
Prov. Blouberg
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<collectorName id="26D712AEFFD69656F3B4FC7AFB6BFC57" box="[1160,1278,908,933]" pageId="2" pageNumber="473">NR North</collectorName>
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||
/ dd22 98S, dd
|
||
<geoCoordinate id="EE1611BFFFD69656F788FC58FE9DFC3A" box="[180,264,943,968]" direction="east" orientation="longitude" pageId="2" pageNumber="473" precision="555" value="29.12">29.12E</geoCoordinate>
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||
”, “
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||
<collectingDate id="EFD8A850FFD69656F614FC58FE0BFC3A" box="[296,414,943,968]" pageId="2" pageNumber="473" value="2012-06-12">
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||
<date id="FF9C51B8FFD69656F614FC58FE0BFC3A" box="[296,414,943,968]" pageId="2" pageNumber="473" value="2012-06-12">6.12.2012</date>
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||
</collectingDate>
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||
;
|
||
<elevation id="000F904BFFD69656F695FC58FE7EFC3A" box="[425,491,943,968]" metricMagnitude="2" metricUnit="m" metricValue="8.66" pageId="2" pageNumber="473" unit="m" value="866.0">
|
||
<quantity id="4CDADA9DFFD69656F695FC58FE7EFC3A" box="[425,491,943,968]" metricMagnitude="2" metricUnit="m" metricValue="8.66" pageId="2" pageNumber="473" unit="m" value="866.0">866m</quantity>
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</elevation>
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||
, BLN1 /
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<collectorName id="26D712AEFFD69656F56EFC58FCABFC3A" box="[594,830,943,968]" pageId="2" pageNumber="473">Roodeberg Bushveld</collectorName>
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||
/ leg.
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<collectorName id="26D712AEFFD69656F4B8FC58FBD1FC3A" box="[900,1092,943,968]" pageId="2" pageNumber="473">Colin Schoeman</collectorName>
|
||
”; male (
|
||
<collectionCode id="ED33EFBDFFD69656F3A2FC47FB46FC3A" box="[1182,1235,944,968]" pageId="2" pageNumber="473">MIZ</collectionCode>
|
||
<collectionCode id="ED33EFBDFFD69656F3E7FC47FA83FC3A" box="[1243,1302,944,968]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13841" name="Panjab University" pageId="2" pageNumber="473">PAN</collectionCode>
|
||
): “
|
||
<collectorName id="26D712AEFFD69656F20EFC58FF2BFC1E" pageId="2" pageNumber="473">Blouberg NR</collectorName>
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||
/
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||
<collectorName id="26D712AEFFD69656F7E8FC23FE24FC1F" box="[212,433,980,1005]" pageId="2" pageNumber="473">Rodeberg Bushveld</collectorName>
|
||
/ -22.98; 29.12”, “leg.
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<collectorName id="26D712AEFFD69656F588FC23FCFBFC1F" box="[692,878,980,1005]" pageId="2" pageNumber="473">Colin Schoeman</collectorName>
|
||
/ BLN7c / I”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33824F3FFD69656F7FBFC00FAA5FBE2" box="[199,1328,1015,1040]" pageId="2" pageNumber="473" type="etymology">
|
||
<paragraph id="8B9D7778FFD69656F7FBFC00FAA5FBE2" blockId="2.[151,1437,836,2013]" box="[199,1328,1015,1040]" pageId="2" pageNumber="473">
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||
<emphasis id="B956AB6AFFD69656F7FBFC00FEDBFBE2" bold="true" box="[199,334,1015,1040]" pageId="2" pageNumber="473">Etymology.</emphasis>
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||
The name refers to the
|
||
<typeStatus id="5499C9DAFFD69656F56BFC0FFD12FBE2" box="[599,647,1016,1040]" pageId="2" pageNumber="473">type</typeStatus>
|
||
locality, Blouberg Nature Reserve (
|
||
<collectingRegion id="49E6B99AFFD69656F325FC00FB11FBE2" box="[1049,1156,1015,1040]" country="South Africa" name="Limpopo" pageId="2" pageNumber="473">Limpopo</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33537E8FFD69656F3B3FC00FAB0FBE2" box="[1167,1317,1015,1040]" name="South Africa" pageId="2" pageNumber="473">
|
||
<collectingRegion id="49E6B99AFFD69656F3B3FC00FB45FBE2" box="[1167,1232,1015,1040]" country="Tanzania" name="Kusini Unguja" pageId="2" pageNumber="473">South</collectingRegion>
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||
Africa
|
||
</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33824F3FFD69656F7FBFBEBFF6BFB52" pageId="2" pageNumber="473" type="diagnosis">
|
||
<paragraph id="8B9D7778FFD69656F7FBFBEBFF6BFB52" blockId="2.[151,1437,836,2013]" pageId="2" pageNumber="473">
|
||
<emphasis id="B956AB6AFFD69656F7FBFBEBFED4FBC7" bold="true" box="[199,321,1052,1077]" pageId="2" pageNumber="473">Diagnosis.</emphasis>
|
||
This species is distinguished from all other congeners by a unique structure of pro- (first tarsomere extremely widened, following tarsomeres narrow;
|
||
<figureCitation id="13196BFDFFD69656F5C7FBC8FCC9FBAA" box="[763,860,1087,1112]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="2" pageNumber="473">Fig. 1A</figureCitation>
|
||
), mesotarsi (first tarsomere widened;
|
||
<figureCitation id="13196BFDFFD69656F20EFBC8FA1AFBAA" box="[1330,1423,1087,1112]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[151,1435,681,1847]" captionTargetId="figure@5.[151,1435,681,1847]" captionTargetPageId="5" captionText="FIGURE 2. Diagnostic characters proposed for Anaxius: mentum (A), proleg (B, G), mesoleg (C, F), metaleg (D), metaventrite (H, I). Anaxius bloubergensis (E, F), A. campbellae (A, H), A. meletsensis (G, I), and A. prozeskyi (B, C, D). Abbreviations: d—denticles, ld—surface between labial palpi, lk—longitudinal keel, lw—lateral wing, p—palpifer, t—apical denticle, tub—tubercles." httpUri="https://zenodo.org/record/1457471/files/figure.png" pageId="2" pageNumber="473">Fig. 2F</figureCitation>
|
||
), mesotibiae (flattened laterally; protruding outwards apically;
|
||
<figureCitation id="13196BFDFFD69656F46BFB93FC3BFB8E" box="[855,942,1124,1149]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[151,1435,681,1847]" captionTargetId="figure@5.[151,1435,681,1847]" captionTargetPageId="5" captionText="FIGURE 2. Diagnostic characters proposed for Anaxius: mentum (A), proleg (B, G), mesoleg (C, F), metaleg (D), metaventrite (H, I). Anaxius bloubergensis (E, F), A. campbellae (A, H), A. meletsensis (G, I), and A. prozeskyi (B, C, D). Abbreviations: d—denticles, ld—surface between labial palpi, lk—longitudinal keel, lw—lateral wing, p—palpifer, t—apical denticle, tub—tubercles." httpUri="https://zenodo.org/record/1457471/files/figure.png" pageId="2" pageNumber="473">Fig. 2F</figureCitation>
|
||
), and aedeagal tegmen (without basal gap;
|
||
<figureCitation id="13196BFDFFD69656F7ABFB70FF65FB52" box="[151,240,1159,1184]" captionStart="FIGURE 3" captionStartId="7.[151,250,1440,1462]" captionTargetBox="[200,1386,725,1434]" captionTargetId="figure@7.[200,1386,725,1434]" captionTargetPageId="7" captionText="FIGURE 3. Aedeagal tegmen morphology. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457473/files/figure.png" pageId="2" pageNumber="473">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33824F3FFD69650F7FBFB5BFA93FE5B" lastPageId="4" lastPageNumber="475" pageId="2" pageNumber="473" type="description">
|
||
<paragraph id="8B9D7778FFD69656F7FBFB5BFC34FB36" blockId="2.[151,1437,836,2013]" box="[199,929,1196,1221]" pageId="2" pageNumber="473">
|
||
<emphasis id="B956AB6AFFD69656F7FBFB5BFECCFB37" bold="true" box="[199,345,1196,1221]" pageId="2" pageNumber="473">Description.</emphasis>
|
||
Body—length = 8.0-9.0 mm. Habitus as in
|
||
<figureCitation id="13196BFDFFD69656F479FB5BFC0BFB36" box="[837,926,1196,1221]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1992]" captionTargetBox="[170,1407,373,1936]" captionTargetId="figure@3.[151,1436,373,1948]" captionTargetPageId="3" captionText="FIGURE 1. Habitus images of the studied species. Anaxius bloubergensis (A), A. campbellae (B), A. limpopoensis (C), A. meletsensis (D), A. obesus (E), A. prozeskyi (F), and A. pseudoloensus (G)." httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="2" pageNumber="473">Fig. 1A</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9D7778FFD69656F7FBFB38FE44F9A2" blockId="2.[151,1437,836,2013]" pageId="2" pageNumber="473">Dorsal side of head covered with fine confluent punctures (0.5–1.0 diameters apart). Frontoclypeal suture rough. Canthus rounded at base; narrower than head at level of eyes. Clypeal emargination shallow (clypeal emargination width: depth ratio ca. 8.0). Labrum narrow (width: length ratio ca. 1.1); strongly emarginate medially; with basal membrane exposed. Ligula exposed, not covered by mentum. Mentum widest in apical part; with well-developed medial keel; with exposed lateral lobes; not covering cardo or basistipes; apically with median indentation. Submentum pentagonal; short. Apical segment of maxillary palpus trapezoidal; with sensory field occupying whole proximal edge. Second segment of maxillary palpus elongate (length: width ratio ca. 2.4). Palpifer located laterally on basistipes. Anterior tentorial pit circular. Eye not dorsally shielded by any keel. Margin between maxillary fossa and antenna insertion well marked. Antenna relatively long (antenna: pronotum length ratio ca. 1.4); third antennomere short (3rd: 2nd antennomere length ratio ca. 3.0); fifth antennomere elongated (length: width ratio ca. 2.0).</paragraph>
|
||
<paragraph id="8B9D7778FFD69656F7FBF9AAFDD0F882" blockId="2.[151,1437,836,2013]" pageId="2" pageNumber="473">Pronotum narrower than elytra (elytra: pronotum width ratio ca. 1.3); relatively narrow (length: width ratio of pronotal disc ca. 0.7); disc dull, covered with fine punctures (3–4 diameters apart). Lateral sides of pronotal disc straight. Disc widest in basal half; without apophyseal depressions. Basal and lateral emargination of pronotal disc present; anterior interrupted in middle. Base of pronotum straight. Posterior angles not protruding beyond base. Pronotal hypomeron dull; without punctures; covered with very fine longitudinal rugosities; without submarginal indentation. Intercoxal process of prosternum depressed apically (lateral view); apically with oval depression on ventral surface; slightly protruding towards first abdominal segment. Procoxae narrowly separated (procoxa: intercoxal process width ratio ca. 2.0).</paragraph>
|
||
<paragraph id="8B9D7778FFD69657F7FBF88BFBB5FF27" blockId="2.[151,1437,836,2013]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="474" pageId="2" pageNumber="473">Elytral striae and intervals not clearly visible. Elytron covered by shiny tubercles (0.1 diameters apart) displayed on dull surface. 1st, 3rd, 5th and 7th intervals protruding on disc; transforming into sparsely (1–2 diameters apart) distributed, large, tubercles on elytral apex. Elytral base straight; not emarginate. Elytral humerus rounded. Epipleuron slightly narrowing apically, then of constant width; basally covering all elytral intervals (ventral view). Scutellum relatively large; triangular. Metathoracic wings absent.</paragraph>
|
||
<paragraph id="8B9D7778FFD79657F7FBFF28FAF5FEB2" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="474">Metaventrite extremely short (metacoxal cavity: metaventrite (between insertions of meso- and metacoxae) length ratio ca. 10.0). Process of first abdominal ventrite wide (distances between mesocoxae: metacoxae ratio ca. 0.9). Fifth abdominal ventrite without submarginal sulcus; covered with fine punctures (4–5 diameters apart).</paragraph>
|
||
<caption id="DF5D27F0FFD79657F7ABF845FC19F815" httpUri="https://zenodo.org/record/1457469/files/figure.png" pageId="3" pageNumber="474" startId="3.[151,250,1970,1992]" targetBox="[170,1407,373,1936]" targetPageId="3">
|
||
<paragraph id="8B9D7778FFD79657F7ABF845FC19F815" blockId="3.[151,1437,1970,2023]" pageId="3" pageNumber="474">
|
||
<emphasis id="B956AB6AFFD79657F7ABF845FE9AF835" bold="true" box="[151,271,1970,1992]" pageId="3" pageNumber="474">FIGURE 1</emphasis>
|
||
. Habitus images of the studied species.
|
||
<taxonomicName id="4C220CFBFFD79657F58BF844FC0AF83A" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[695,927,1970,1992]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="bloubergensis">
|
||
<emphasis id="B956AB6AFFD79657F58BF844FC0AF83A" box="[695,927,1970,1992]" italics="true" pageId="3" pageNumber="474">Anaxius bloubergensis</emphasis>
|
||
</taxonomicName>
|
||
(A),
|
||
<taxonomicName id="4C220CFBFFD79657F4E6F844FBF8F83A" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[986,1133,1970,1992]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="campbellae">
|
||
<emphasis id="B956AB6AFFD79657F4E6F844FBF8F83A" box="[986,1133,1970,1992]" italics="true" pageId="3" pageNumber="474">A. campbellae</emphasis>
|
||
</taxonomicName>
|
||
(B),
|
||
<taxonomicName id="4C220CFBFFD79657F394F844FAD9F83A" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[1192,1356,1970,1992]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="limpopoensis">
|
||
<emphasis id="B956AB6AFFD79657F394F844FAD9F83A" box="[1192,1356,1970,1992]" italics="true" pageId="3" pageNumber="474">A. limpopoensis</emphasis>
|
||
</taxonomicName>
|
||
(C),
|
||
<taxonomicName id="4C220CFBFFD79657F2BAF844FE9EF815" authorityName="Kamiński & Schoeman" authorityYear="2018" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="meletsensis">
|
||
<emphasis id="B956AB6AFFD79657F2BAF844FE9EF815" italics="true" pageId="3" pageNumber="474">A. meletsensis</emphasis>
|
||
</taxonomicName>
|
||
(D),
|
||
<taxonomicName id="4C220CFBFFD79657F67CF826FE34F815" authorityName="Fahraeus" authorityYear="1870" box="[320,417,2001,2023]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="B956AB6AFFD79657F67CF826FE34F815" box="[320,417,2001,2023]" italics="true" pageId="3" pageNumber="474">A. obesus</emphasis>
|
||
</taxonomicName>
|
||
(E),
|
||
<taxonomicName id="4C220CFBFFD79657F6E8F826FDDAF815" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[468,591,2001,2023]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="prozeskyi">
|
||
<emphasis id="B956AB6AFFD79657F6E8F826FDDAF815" box="[468,591,2001,2023]" italics="true" pageId="3" pageNumber="474">A. prozeskyi</emphasis>
|
||
</taxonomicName>
|
||
(F), and
|
||
<taxonomicName id="4C220CFBFFD79657F590F826FCC9F815" authorityName="Kamiński & Schoeman" authorityYear="2018" box="[684,860,2001,2023]" class="Insecta" family="Tenebrionidae" genus="Anaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="474" phylum="Arthropoda" rank="species" species="pseudoloensus">
|
||
<emphasis id="B956AB6AFFD79657F590F826FCC9F815" box="[684,860,2001,2023]" italics="true" pageId="3" pageNumber="474">A. pseudoloensus</emphasis>
|
||
</taxonomicName>
|
||
(G).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9D7778FFD09650F7FBFF60FF71FECC" blockId="4.[151,1436,151,496]" pageId="4" pageNumber="475">Male pro- and mesotarsi with basal tarsomere widened. Protibia with deep longitudinal cavity. Profemora simple. Mesotibiae flattened laterally; protruding outwards apically. Mesofemora with internal extension in apical part. Metatibia and metafemora with fringe of setae distributed on whole length. Metatibia curved inwards. First tarsomere of metatarsi elongated (1.5× longer than 4th one). Length of metatarsus equal to half of metatibial length.</paragraph>
|
||
<paragraph id="8B9D7778FFD09650F7FBFEBEFA93FE5B" blockId="4.[151,1436,151,496]" pageId="4" pageNumber="475">Aedeagal tegmen widest in middle; unipartite; without basal gap. Apical part divided (up to 0.15 of whole tegmen length). Clava straight. Penis of even width along most of its length; with apical part not covered by tegmen, exposed dorsally. Basal apophyses of penis relatively short (ca. 0.4 of whole tegmen length).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33824F3FFD09650F7FBFE44FB11FE02" pageId="4" pageNumber="475" type="distribution">
|
||
<paragraph id="8B9D7778FFD09650F7FBFE44FB11FE02" blockId="4.[151,1436,151,496]" pageId="4" pageNumber="475">
|
||
<emphasis id="B956AB6AFFD09650F7FBFE44FEF5FE3E" bold="true" box="[199,352,435,460]" pageId="4" pageNumber="475">Distribution.</emphasis>
|
||
This species has been collected in the following ecoregions of
|
||
<collectingCountry id="F33537E8FFD09650F3B1FE44FAB3FE3E" box="[1165,1318,435,460]" name="South Africa" pageId="4" pageNumber="475">
|
||
<collectingRegion id="49E6B99AFFD09650F3B1FE44FB5BFE3E" box="[1165,1230,435,460]" country="Tanzania" name="Kusini Unguja" pageId="4" pageNumber="475">South</collectingRegion>
|
||
Africa
|
||
</collectingCountry>
|
||
(
|
||
<figureCitation id="13196BFDFFD09650F203FE44FA18FE3E" box="[1343,1421,435,460]" captionStart="FIGURE 5" captionStartId="11.[151,250,1731,1753]" captionTargetBox="[161,1437,844,1704]" captionTargetId="figure@11.[151,1437,844,1710]" captionTargetPageId="11" captionText="FIGURE 5. Distribution of the species representing Anaxius." httpUri="https://zenodo.org/record/1457477/files/figure.png" pageId="4" pageNumber="475">Fig. 5</figureCitation>
|
||
): Drakensberg montane grasslands, woodlands and forests; and Southern African bushveld.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |