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<document id="2824660391E47979BA7A50A688B16D40" ID-CLB-Dataset="182452" ID-DOI="10.11646/zootaxa.5169.4.8" ID-GBIF-Dataset="bd9d01e4-d98e-4318-a101-ce2e7bb46666" ID-ISSN="1175-5326" ID-Zenodo-Dep="6952610" ID-ZooBank="77313217-7775-4EF1-A39D-435EB8405EDB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659440142825" checkinUser="plazi" docAuthor="Li, Hua-Zhao &amp; Liu, Zhe" docDate="2022" docId="8122CC127A6CA2141EE0FE51FB91FEE5" docLanguage="en" docName="zootaxa.5169.4.8.pdf" docOrigin="Zootaxa 5169 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stibochiona nicea subsp. wangyukunae Li &amp; Liu 2022, ssp. nov." docType="treatment" docUuid="6A8D23C2-FD10-4FDF-90D6-CA762A46BCD5" docUuidSource="ZooBank" docVersion="7" lastPageNumber="393" masterDocId="7D1BB46A7A6DA2101E77FF83FFFDFF80" masterDocTitle="Notes on Stibochiona nicea (Gray, 1846) from China with description of a new subspecies from Hainan Island (Lepidoptera, Nymphalidae)" masterLastPageNumber="398" masterPageNumber="392" pageNumber="393" updateTime="1699363299037" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C91F256E95C47080021BBB30458FD007">Notes on Stibochiona nicea (Gray, 1846) from China with description of a new subspecies from Hainan Island (Lepidoptera, Nymphalidae)</mods:title>
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<mods:namePart id="6263759F1B71BA28CC863291CC31CA9B">Li, Hua-Zhao</mods:namePart>
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<mods:affiliation id="981FB7AA87BF2639EB211D91C62AEC3F">College of Animal Science and Technology, Huazhong Agricultural University, Wuhan, Hubei, P. R. China. &amp; lhz _ euphony @ foxmail. com; https: // orcid. org / 0000 - 0002 - 3656 - 193 X</mods:affiliation>
<mods:nameIdentifier id="418780BEADDEE0D47A28040876B86BCC" type="email">lhz_euphony@foxmail.com</mods:nameIdentifier>
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<mods:namePart id="159F108A793582CE9D484AAAB3CFAF86">Liu, Zhe</mods:namePart>
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<mods:affiliation id="07460735A5E420BB7C45F5BB9E9D8C2A">Nanfanghangkongcheng, Sanya, Hainan, P. R. China. &amp; 149058809 @ qq. com; https: // orcid. org / 0000 - 0001 - 7036 - 7164</mods:affiliation>
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<mods:date id="9A1AB3361610B3E035DFA5BF050D98B7">2022</mods:date>
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<taxonomicName id="CE8B06877A6CA2111EE0FE51FDDFFE6D" authority="Li &amp; Liu, 2022" authorityName="Li &amp; Liu" authorityYear="2022" box="[151,546,466,493]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" status="subsp. nov." subSpecies="wangyukunae">
<emphasis id="3BFFA1167A6CA2111EE0FE51FDDFFE6D" bold="true" box="[151,546,466,493]" italics="true" pageId="1" pageNumber="393">Stibochiona nicea wangyukunae</emphasis>
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<taxonomicNameLabel id="20CC1C6D7A6CA2111C5EFE50FD6FFE6D" box="[553,658,467,493]" pageId="1" pageNumber="393" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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(
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,
<figureCitation id="91B061817A6CA2111F60FE74FE9DFD91" box="[279,352,503,529]" captionStart="FIGURE 1936" captionStartId="3.[151,250,1876,1900]" captionTargetBox="[151,1436,181,1852]" captionTargetId="figure-20@3.[151,1436,181,1852]" captionTargetPageId="3" captionText="FIGURE 1936. Ventral view of Stibochiona nicea sspp. 1926, wangyukunae ssp. nov. from Hainan; 2734, nicea from S. China; 3536, subucula from W. Yunnan." figureDoi="http://doi.org/10.5281/zenodo.6952617" httpUri="https://zenodo.org/record/6952617/files/figure.png" pageId="1" pageNumber="393">1926</figureCitation>
,
<figureCitation id="91B061817A6CA2111F1CFE74FE7AFD91" box="[363,391,503,529]" captionStart="FIGURE 3739" captionStartId="4.[151,250,929,953]" captionTargetBox="[157,1428,388,897]" captionTargetId="figure-118@4.[151,1436,381,905]" captionTargetPageId="4" captionText="FIGURE 3739. Male genitalia of Stibochiona nicea sspp with left valva removed. 37, wangyukunae ssp. nov., holotype, HN1; 38, nicea, GX1; 39, subucula, YN2; a = lateral view of vinculum; b = lateral view of valvae; c = lateral view of phallus; d = posterior view of juxta." figureDoi="http://doi.org/10.5281/zenodo.6952619" httpUri="https://zenodo.org/record/6952619/files/figure.png" pageId="1" pageNumber="393">37</figureCitation>
)
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LSID
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<treatmentCitation id="882A5B157A6CA2111EE0FDDCFD55FDF7" author="Joicey, J. J. &amp; Talbot, G." box="[151,680,607,631]" page="10" pageId="1" pageNumber="393" year="1928">
<taxonomicName id="CE8B06877A6CA2111EE0FDDCFD55FDF7" ID-CoL="52HRW" authority=": Joicey &amp; Talbot, 1928: 10" authorityName=": Joicey &amp; Talbot" authorityPageNumber="10" authorityYear="1928" box="[151,680,607,631]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="nicea">
<emphasis id="3BFFA1167A6CA2111EE0FDDCFE70FDF7" box="[151,397,607,631]" italics="true" pageId="1" pageNumber="393">Stibochiona nicea nicea</emphasis>
:
<bibRefCitation id="6D1A00F57A6CA2111FECFDDCFD55FDF7" author="Joicey, J. J. &amp; Talbot, G." box="[411,680,607,631]" pageId="1" pageNumber="393" pagination="3 - 18" refId="ref3405" refString="Joicey, J. J. &amp; Talbot, G. (1928) A catalogue of the Lepidoptera of Hainan. Bulletin of the Hill Museum, 2, 3 - 18 + 183 - 191." type="journal article" year="1928">Joicey &amp; Talbot, 1928: 10</bibRefCitation>
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, Five Finger Mt. [Mt. Wuzhi], June,
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,
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, May,
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;
<treatmentCitation id="882A5B157A6CA2111B68FDDCFED2FD16" author="Gu, M. B. &amp; Chen, P. Z." page="177" pageId="1" pageNumber="393" year="1997">
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, fig. 174.
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:
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male,
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<collectingDate id="6D71A22C7A6CA2111F2DFD42FE25FD5B" box="[346,472,705,731]" pageId="1" pageNumber="393" value="2022-04-19">19.IV.2022</collectingDate>
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,
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,
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,
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,
<collectorName id="A47E18D27A6CA2111AD4FD42FB00FD5B" box="[1187,1277,705,731]" pageId="1" pageNumber="393">Zhe Liu</collectorName>
leg. (
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;
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)
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.
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<typeStatus id="D630C3A67A6CA2111E90FD67FE9EFD7E" box="[231,355,740,766]" pageId="1" pageNumber="393" type="paratype">Paratypes</typeStatus>
:
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male,
<materialsCitation id="B9E377597A6CA2111FC0FD67FF2DFCA1" ID-GBIF-Occurrence="3962316301" collectingDate="2020-03-07" collectionCode="CZL" collectorName="Zhe Liu" country="China" county="Ledong Li Autonomous County" location="Collection of Zhe Liu" pageId="1" pageNumber="393" specimenCode="HN2" specimenCount="1" stateProvince="Hainan" typeStatus="paratype">
<date id="7D355BC47A6CA2111FC0FD67FDD4FD7E" box="[439,553,740,766]" pageId="1" pageNumber="393" value="2020-03-07">
<collectingDate id="6D71A22C7A6CA2111FC0FD67FDD4FD7E" box="[439,553,740,766]" pageId="1" pageNumber="393" value="2020-03-07">7.III.2020</collectingDate>
</date>
, same locality as holotype,
<collectorName id="A47E18D27A6CA2111D10FD67FC3EFD7E" box="[871,963,740,766]" pageId="1" pageNumber="393">Zhe Liu</collectorName>
leg. (
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,
<collectionCode id="6F9AE5C17A6CA2111B78FD67FABAFD7E" box="[1295,1351,740,766]" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="1" pageNumber="393">CZL</collectionCode>
;
<specimenCode id="592DD57F7A6CA2111B22FD67FA6DFD7E" box="[1365,1424,740,766]" collectionCode="CZL" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="1" pageNumber="393">HN2</specimenCode>
); male
</materialsCitation>
,
<materialsCitation id="B9E377597A6CA2111E97FC84FBD9FCA1" ID-GBIF-Occurrence="3962316303" box="[224,1060,775,801]" collectingDate="2020-03-27" collectionCode="CZL" collectorName="Zhe Liu" country="China" county="Ledong Li Autonomous County" location="Jianfengling" pageId="1" pageNumber="393" specimenCode="HN3" specimenCount="1" stateProvince="Hainan" typeStatus="paratype">
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<collectingDate id="6D71A22C7A6CA2111E97FC84FE9DFCA1" box="[224,352,775,801]" pageId="1" pageNumber="393" value="2020-03-27">27.III.2020</collectingDate>
</date>
, same locality as holotype,
<collectorName id="A47E18D27A6CA2111CDDFC84FCF5FCA1" box="[682,776,775,801]" pageId="1" pageNumber="393">Zhe Liu</collectorName>
leg. (
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;
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); male
</materialsCitation>
,
<materialsCitation id="B9E377597A6CA2111A44FC84FD88FCC3" ID-GBIF-Occurrence="3962316310" collectingDate="2020-04-14" collectionCode="CHZL" collectorName="Zhe Liu" country="China" location="Sanya" municipality="Mt. Liuluo" pageId="1" pageNumber="393" specimenCode="HN4" specimenCount="1" stateProvince="Hainan" typeStatus="paratype">
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<collectingDate id="6D71A22C7A6CA2111A44FC84FB4FFCA1" box="[1075,1202,775,801]" pageId="1" pageNumber="393" value="2020-04-14">14.IV.2020</collectingDate>
</date>
,
<collectingMunicipality id="E950E77E7A6CA2111AB6FC84FABCFCA1" box="[1217,1345,775,801]" pageId="1" pageNumber="393">Mt. Liuluo</collectingMunicipality>
,
<location id="0C542BDF7A6CA2111B26FC84FA65FCA1" LSID="urn:lsid:plazi:treatment:8122CC127A6CA2141EE0FE51FB91FEE5:0C542BDF7A6CA2111B26FC84FA65FCA1" box="[1361,1432,775,801]" country="China" municipality="Mt. Liuluo" name="Sanya" pageId="1" pageNumber="393" stateProvince="Hainan">Sanya</location>
,
<collectingRegion id="CB4FB3E67A6CA2111EE0FCAAFF16FCC3" box="[151,235,809,835]" country="China" name="Hainan" pageId="1" pageNumber="393">Hainan</collectingRegion>
,
<collectorName id="A47E18D27A6CA2111E80FCAAFEACFCC3" box="[247,337,809,835]" pageId="1" pageNumber="393">Zhe Liu</collectorName>
leg. (
<collectionCode id="6F9AE5C17A6CA2111FE5FCAAFE1DFCC3" box="[402,480,809,835]" pageId="1" pageNumber="393">CHZL</collectionCode>
;
<specimenCode id="592DD57F7A6CA2111F9BFCAAFDD5FCC3" box="[492,552,809,835]" collectionCode="CHZL" pageId="1" pageNumber="393">HN4</specimenCode>
); male
</materialsCitation>
,
<materialsCitation id="B9E377597A6CA2111CF6FCAAFA65FCC3" ID-GBIF-Occurrence="3962316308" box="[641,1432,809,836]" collectingDate="2020-11-02" collectionCode="CZL" collectorName="Zhe Liu" country="China" county="Ledong Li Autonomous County" location="Jianfengling" pageId="1" pageNumber="393" specimenCode="HN5" specimenCount="1" stateProvince="Hainan" typeStatus="paratype">
<date id="7D355BC47A6CA2111CF6FCAAFD08FCC3" box="[641,757,809,835]" pageId="1" pageNumber="393" value="2020-11-02">
<collectingDate id="6D71A22C7A6CA2111CF6FCAAFD08FCC3" box="[641,757,809,835]" pageId="1" pageNumber="393" value="2020-11-02">2.XI.2020</collectingDate>
</date>
, same locality as holotype,
<collectorName id="A47E18D27A6CA2111A59FCAAFB74FCC3" box="[1070,1161,809,835]" pageId="1" pageNumber="393">Zhe Liu</collectorName>
leg. (
<collectionCode id="6F9AE5C17A6CA2111ABDFCAAFAF9FCC3" box="[1226,1284,809,835]" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="1" pageNumber="393">CZL</collectionCode>
;
<specimenCode id="592DD57F7A6CA2111B78FCAAFABAFCC4" box="[1295,1351,809,836]" collectionCode="CZL" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="1" pageNumber="393">HN5</specimenCode>
); male
</materialsCitation>
,
<date id="7D355BC47A6CA2111EE0FCCFFEF6FCE6" box="[151,267,844,870]" pageId="1" pageNumber="393" value="2022-06-03">
3.
<collectionCode id="6F9AE5C17A6CA2111ED0FCCFFF35FCE6" box="[167,200,844,870]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="393">VI</collectionCode>
<materialsCitation id="B9E377597A6CA2111EBFFCCFFEF6FCE6" ID-GBIF-Occurrence="3962316302" box="[200,267,844,870]" collectionCode="CZL" collectorName="Zhe Liu" country="China" location="Wuzhishan" municipality="Mt. Wuzhi" pageId="1" pageNumber="393" specimenCode="HN6" specimenCount="1" stateProvince="Hainan" typeStatus="paratype">.2022</materialsCitation>
</date>
,
<collectingMunicipality id="E950E77E7A6CA2111F6FFCCEFE69FCE6" box="[280,404,844,871]" pageId="1" pageNumber="393">Mt. Wuzhi</collectingMunicipality>
,
<location id="0C542BDF7A6CA2111FD6FCCFFDDCFCE6" LSID="urn:lsid:plazi:treatment:8122CC127A6CA2141EE0FE51FB91FEE5:0C542BDF7A6CA2111FD6FCCFFDDCFCE6" box="[417,545,844,870]" country="China" municipality="Mt. Wuzhi" name="Wuzhishan" pageId="1" pageNumber="393" stateProvince="Hainan">Wuzhishan</location>
,
<collectingRegion id="CB4FB3E67A6CA2111C59FCCFFD7FFCE6" box="[558,642,844,870]" country="China" name="Hainan" pageId="1" pageNumber="393">Hainan</collectingRegion>
,
<collectorName id="A47E18D27A6CA2111CF9FCCFFD14FCE6" box="[654,745,844,870]" pageId="1" pageNumber="393">Zhe Liu</collectorName>
leg. (
<collectionCode id="6F9AE5C17A6CA2111D5BFCCFFC9BFCE6" box="[812,870,844,870]" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="1" pageNumber="393">CZL</collectionCode>
;
<specimenCode id="592DD57F7A6CA2111D05FCCFFC53FCE6" box="[882,942,844,870]" collectionCode="CZL" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="1" pageNumber="393">HN6</specimenCode>
); female,
<date id="7D355BC47A6CA2111A68FCCFFB6FFCE6" box="[1055,1170,844,870]" pageId="1" pageNumber="393" value="2022-06-01">
1.
<collectionCode id="6F9AE5C17A6CA2111A5BFCCFFBB3FCE6" box="[1068,1102,844,870]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="393">VI</collectionCode>
<materialsCitation id="B9E377597A6CA2111A3AFCCFFB6FFCE6" ID-GBIF-Occurrence="3962316305" box="[1101,1170,844,870]" collectionCode="CHZL" collectorName="Bo-Tao Huang" country="China" location="Wuzhishan" municipality="Mt. Wuzhi" pageId="1" pageNumber="393" specimenCode="HN7" specimenCount="1" stateProvince="Hainan" typeStatus="paratype">.2022</materialsCitation>
</date>
, same locality as HN6,
<collectorName id="A47E18D27A6CA2111EE0FCECFEC0FC09" box="[151,317,879,905]" pageId="1" pageNumber="393">Bo-Tao Huang</collectorName>
leg. (
<collectionCode id="6F9AE5C17A6CA2111F0CFCECFE34FC09" box="[379,457,879,905]" pageId="1" pageNumber="393">CHZL</collectionCode>
;
<specimenCode id="592DD57F7A6CA2111FA2FCECFDEDFC09" box="[469,528,879,905]" collectionCode="CHZL" pageId="1" pageNumber="393">HN7</specimenCode>
); female,
<date id="7D355BC47A6CA2111C09FCECFD0CFC09" box="[638,753,879,905]" pageId="1" pageNumber="393" value="2022-06-04">
4.
<collectionCode id="6F9AE5C17A6CA2111CE7FCECFD4DFC09" box="[656,688,879,905]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="393">VI</collectionCode>
<materialsCitation id="B9E377597A6CA2111CC7FCECFD0CFC09" ID-GBIF-Occurrence="3962316306" box="[688,753,879,905]" collectionCode="CZL" collectorName="Zhe Liu" country="China" location="Wuzhishan" municipality="Mt. Wuzhi" pageId="1" pageNumber="393" specimenCode="HN8" specimenCount="1" stateProvince="Hainan" typeStatus="paratype">.2022</materialsCitation>
</date>
, same locality as HN6,
<collectorName id="A47E18D27A6CA2111D8CFCECFBA8FC09" box="[1019,1109,879,905]" pageId="1" pageNumber="393">Zhe Liu</collectorName>
leg. (
<collectionCode id="6F9AE5C17A6CA2111AE3FCECFB33FC09" box="[1172,1230,879,905]" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="1" pageNumber="393">CZL</collectionCode>
;
<specimenCode id="592DD57F7A6CA2111AAEFCECFAE9FC09" box="[1241,1300,879,905]" collectionCode="CZL" country="Portugal" httpUri="http://grbio.org/cool/03yf-yar9" name="Centro de Zoologia" pageId="1" pageNumber="393">HN8</specimenCode>
).
</paragraph>
</subSubSection>
<subSubSection id="41912E8F7A6CA2111EB0FC12FDADFAEE" pageId="1" pageNumber="393" type="description">
<paragraph id="09347D047A6CA2111EB0FC12FE88FB63" blockId="1.[151,1437,705,1459]" pageId="1" pageNumber="393">
<emphasis id="3BFFA1167A6CA2111EB0FC12FE5BFC2B" bold="true" box="[199,422,913,939]" pageId="1" pageNumber="393">Description. Male.</emphasis>
Length of forewing
<quantity id="CE73D0E17A6CA2111CEFFC12FCEEFC2C" box="[664,787,913,940]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.3" metricValueMin="2.9" pageId="1" pageNumber="393" unit="mm" value="31.0" valueMax="33.0" valueMin="29.0">2933 mm</quantity>
(n=
<quantity id="CE73D0E17A6CA2111D32FC12FC49FC2C" box="[837,948,913,940]" metricMagnitude="-3" metricUnit="m" metricValue="6.33" pageId="1" pageNumber="393" unit="mm" value="6.33">6, 33 mm</quantity>
in
<typeStatus id="D630C3A67A6CA2111DABFC12FBBEFC2B" box="[988,1091,913,939]" pageId="1" pageNumber="393" type="holotype">holotype</typeStatus>
). Forewing upper side ground color black with three rows of concrete white spots in discal, postdiscal, and submarginal areas. A diffused zigzag blue line present between submarginal and marginal series of white spots. Discal cell scattered with several blue spots. Forewing under side generally similar to upper side but with all spots more prominent and without zigzag blue line in submarginal area. Hindwing upper side ground color black. A serrate blue line running from cell 1b to cell
<quantity id="CE73D0E17A6CA2111EBFFBBDFF09FBD9" box="[200,244,1086,1113]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="393" unit="in" value="5.0">5 in</quantity>
postdiscal area. A series of characteristic marginal patches presented from cells 1b
<quantity id="CE73D0E17A6CA2111AD8FBBCFB27FBD9" box="[1199,1242,1087,1113]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="1" pageNumber="393" unit="in" value="6.0">6 in</quantity>
submarginal and marginal areas. Blue section of patch usually double broader than white section. Hindwing under side with series of blue spots present from base to postmedial area, marginal area with a series of white patches with inner margin concave.
<emphasis id="3BFFA1167A6CA2111F77FB24FEA2FB41" bold="true" box="[256,351,1191,1217]" pageId="1" pageNumber="393">Female.</emphasis>
Length of forewing
<quantity id="CE73D0E17A6CA2111C33FB24FD40FB41" box="[580,701,1191,1217]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.7" metricValueMin="3.6" pageId="1" pageNumber="393" unit="mm" value="36.5" valueMax="37.0" valueMin="36.0">3637 mm</quantity>
(n=2). Similar to male but larger in size. Forewing more rounded, ground color duller.
</paragraph>
<paragraph id="09347D047A6CA2111EB0FB6FFDADFAEE" blockId="1.[151,1437,705,1459]" pageId="1" pageNumber="393">
<emphasis id="3BFFA1167A6CA2111EB0FB6FFE84FA86" bold="true" box="[199,377,1260,1286]" pageId="1" pageNumber="393">Male genitalia.</emphasis>
Uncus lobe-like with distal end forming a small bifurcated hook. Tegumen moderately broad. Gnathos ring-like. Vinculum slender. Juxta Y-shaped. Saccus shorter than half length of vinculum. Valva trapezoid. Costa simple. Sacculus with a rectangular process basally with dense dentation presented at its dorsal ridge. Phallus slender, broader at base. Coecum short.
</paragraph>
</subSubSection>
<subSubSection id="41912E8F7A6CA2111EB0FAF5FC74F949" pageId="1" pageNumber="393" type="diagnosis">
<paragraph id="09347D047A6CA2111EB0FAF5FD51FA33" blockId="1.[151,1437,705,1459]" pageId="1" pageNumber="393">
<emphasis id="3BFFA1167A6CA2111EB0FAF5FEBFFA10" bold="true" box="[199,322,1398,1424]" pageId="1" pageNumber="393">Diagnosis.</emphasis>
The new subspecies can be distinguished from the other two subspecies,
<taxonomicName id="CE8B06877A6CA2111A15FAF4FB30FA11" authorityName=": Joicey &amp; Talbot" authorityYear="1928" box="[1122,1229,1399,1425]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="nicea">ssp. nicea</taxonomicName>
and
<taxonomicName id="CE8B06877A6CA2111B77FAF4FA68FA11" authorityName="Fruhstorfer" authorityYear="1898" box="[1280,1429,1399,1425]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="subucula">
ssp.
<emphasis id="3BFFA1167A6CA2111B47FAF4FA68FA11" box="[1328,1429,1399,1425]" italics="true" pageId="1" pageNumber="393">subucula</emphasis>
</taxonomicName>
, by the combination of the following characters:
</paragraph>
<paragraph id="09347D047A6CA2111EE0FA5CFCBCF99B" blockId="1.[151,1437,1503,1737]" pageId="1" pageNumber="393">
1. The blue discal spots on the underside of the forewing in cells 1b, 2 and 3 are situated inline, whereas in
<taxonomicName id="CE8B06877A6CA2111B06FA5CFEF9F99B" authorityName=": Joicey &amp; Talbot" authorityYear="1928" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="nicea">
ssp.
<emphasis id="3BFFA1167A6CA2111EB0F981FEF9F99B" box="[199,260,1538,1563]" italics="true" pageId="1" pageNumber="393">nicea</emphasis>
</taxonomicName>
and
<taxonomicName id="CE8B06877A6CA2111F4DF981FE2CF99B" authorityName="Fruhstorfer" authorityYear="1898" box="[314,465,1537,1564]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="subucula">
ssp.
<emphasis id="3BFFA1167A6CA2111F1BF982FE2CF99B" box="[364,465,1537,1563]" italics="true" pageId="1" pageNumber="393">subucula</emphasis>
</taxonomicName>
the spot in cell 2 shifted basally.
</paragraph>
<paragraph id="09347D047A6CA2111EE0F9A7FCCBF9E1" blockId="1.[151,1437,1503,1737]" pageId="1" pageNumber="393">
2. On the hindwing upper side, the blue-white marginal patches are compact and rectangular as in
<taxonomicName id="CE8B06877A6CA2111A89F9A6FA68F9BE" authorityName="Fruhstorfer" authorityYear="1898" box="[1278,1429,1572,1599]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="subucula">
ssp.
<emphasis id="3BFFA1167A6CA2111B47F9A7FA68F9BE" box="[1328,1429,1572,1598]" italics="true" pageId="1" pageNumber="393">subucula</emphasis>
</taxonomicName>
, whereas in
<taxonomicName id="CE8B06877A6CA2111F30F9C4FE48F9E0" authorityName=": Joicey &amp; Talbot" authorityYear="1928" box="[327,437,1607,1633]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="nicea">
ssp.
<emphasis id="3BFFA1167A6CA2111F0FF9C4FE48F9E0" box="[376,437,1607,1632]" italics="true" pageId="1" pageNumber="393">nicea</emphasis>
</taxonomicName>
, the patches are variable in shape.
</paragraph>
<paragraph id="09347D047A6CA2111EE0F9EAFC74F949" blockId="1.[151,1437,1503,1737]" pageId="1" pageNumber="393">
3. On the hindwing upper side, the blue-white marginal patches have blue section usually broader than the twice the width of the white section, whereas in
<taxonomicName id="CE8B06877A6CA2111CD5F90EFCC4F926" authorityName="Fruhstorfer" authorityYear="1898" box="[674,825,1676,1703]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="subucula">
ssp.
<emphasis id="3BFFA1167A6CA2111CA3F90FFCC4F926" box="[724,825,1676,1702]" italics="true" pageId="1" pageNumber="393">subucula</emphasis>
</taxonomicName>
the white section broader than the blue section, and in
<taxonomicName id="CE8B06877A6CA2111EB0F92CFECBF948" authorityName=": Joicey &amp; Talbot" authorityYear="1928" box="[199,310,1711,1737]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="nicea">
ssp.
<emphasis id="3BFFA1167A6CA2111E8EF92CFECBF948" box="[249,310,1711,1736]" italics="true" pageId="1" pageNumber="393">nicea</emphasis>
</taxonomicName>
, the ratio between white and blue section is variable.
</paragraph>
</subSubSection>
<subSubSection id="41912E8F7A6CA2111EB0F977FC4BF88E" box="[199,950,1780,1806]" pageId="1" pageNumber="393" type="distribution">
<paragraph id="09347D047A6CA2111EB0F977FC4BF88E" blockId="1.[151,1437,1780,2017]" box="[199,950,1780,1806]" pageId="1" pageNumber="393">
<emphasis id="3BFFA1167A6CA2111EB0F977FE9CF88E" bold="true" box="[199,353,1780,1806]" pageId="1" pageNumber="393">Distribution.</emphasis>
Currently only known from
<collectingRegion id="CB4FB3E67A6CA2111CD4F977FD09F88E" box="[675,756,1780,1806]" country="China" name="Hainan" pageId="1" pageNumber="393">Hainan</collectingRegion>
Island, S.
<collectingCountry id="719C3D947A6CA2111D1CF977FC4FF88E" box="[875,946,1780,1806]" name="China" pageId="1" pageNumber="393">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="41912E8F7A6CA2111EB0F894FE44F8D3" pageId="1" pageNumber="393" type="etymology">
<paragraph id="09347D047A6CA2111EB0F894FE44F8D3" blockId="1.[151,1437,1780,2017]" pageId="1" pageNumber="393">
<emphasis id="3BFFA1167A6CA2111EB0F894FEB3F8B1" bold="true" box="[199,334,1815,1841]" pageId="1" pageNumber="393">Etymology.</emphasis>
This subspecific name is dedicated to Dr. Yu-Kun Wang, who financially supports the senior author for his taxonomic studies.
</paragraph>
</subSubSection>
<subSubSection id="41912E8F7A6CA2141EE0F8DFFB91FEE5" lastPageId="4" lastPageNumber="396" pageId="1" pageNumber="393" type="discussion">
<paragraph id="09347D047A6CA2141EE0F8DFFB91FEE5" blockId="1.[151,1437,1780,2017]" lastBlockId="4.[151,1437,151,357]" lastPageId="4" lastPageNumber="396" pageId="1" pageNumber="393">
<emphasis id="3BFFA1167A6CA2111EE0F8DFFEF6F8F6" bold="true" box="[151,267,1884,1910]" pageId="1" pageNumber="393">Remarks.</emphasis>
No individual from Hainan is found resembling specimens from
<collectingCountry id="719C3D947A6CA2111A62F8DFFB89F8F6" box="[1045,1140,1884,1910]" name="Vietnam" pageId="1" pageNumber="393">Vietnam</collectingCountry>
(
<bibRefCitation id="6D1A00F57A6CA2111AF0F8DFFA69F8F6" author="Inayoshi, Y. &amp; Saito, K." box="[1159,1428,1884,1910]" pageId="1" pageNumber="393" refId="ref3360" refString="Inayoshi, Y. &amp; Saito, K. (2021) A Check List of Butterflies in Indo-China, Chiefly from Thailand, Laos &amp; Vietnam. Available from: http: // yutaka. it-n. jp (accessed 7 April 2022)" type="url" year="2021">Inayoshi &amp; Saito 2021</bibRefCitation>
) nor specimens used in the present study or in literatures (e.g.
<bibRefCitation id="6D1A00F57A6CA2111D15F8FCFC1CF819" author="Chen, J. L." box="[866,993,1919,1945]" pageId="1" pageNumber="393" refId="ref2993" refString="Chen, J. L. (2017) Genus Stibochiona. In: Wu, C. S. &amp; Hsu, Y. F. (Eds.), Butterflies of China. The straits publishing house, Fuzhou, pp. 0881. [in Chinese]" type="book" year="2017">Chen 2017</bibRefCitation>
) from S.
<collectingCountry id="719C3D947A6CA2111A25F8FCFB65F819" box="[1106,1176,1919,1945]" name="China" pageId="1" pageNumber="393">China</collectingCountry>
, which are adjacent to
<collectingRegion id="CB4FB3E67A6CA2111EE0F820FF11F83D" box="[151,236,1955,1981]" country="China" name="Hainan" pageId="1" pageNumber="393">Hainan</collectingRegion>
. Hence, we hereby treat population from
<collectingRegion id="CB4FB3E67A6CA2111CCFF820FCF4F83D" box="[696,777,1955,1981]" country="China" name="Hainan" pageId="1" pageNumber="393">Hainan</collectingRegion>
as a new subspecies via superficial features. Regarding the publications related to butterflies of
<collectingRegion id="CB4FB3E67A6CA2111C5DF844FD83F861" box="[554,638,1991,2017]" country="China" name="Hainan" pageId="1" pageNumber="393">Hainan</collectingRegion>
, 1)
<bibRefCitation id="6D1A00F57A6CA2111CD1F844FC8BF861" author="Gu, M. B. &amp; Chen, P. Z." box="[678,886,1991,2017]" pageId="1" pageNumber="393" refId="ref3207" refString="Gu, M. B. &amp; Chen, P. Z. (1997) Butterflies in Hainan Island. China Forestry Publishing House, Beijing, 355 pp. [in Chinese]" type="book" year="1997">Gu &amp; Chen (1997)</bibRefCitation>
assigned the insular samples to the nominate ssp. Moreover, the figured female should actually be male. 2) A living adult with characters close to the nominotypical subspecies was photographed from Yinggeling,
<collectingRegion id="CB4FB3E67A69A2141CC1FF38FCFAFF55" box="[694,775,187,213]" country="China" name="Hainan" pageId="4" pageNumber="396">Hainan</collectingRegion>
and figured in
<bibRefCitation id="6D1A00F57A69A2141DC0FF38FB72FF55" author="Wang, M. &amp; Liu, J. &amp; Fu, C. R." box="[951,1167,187,213]" pageId="4" pageNumber="396" refId="ref3601" refString="Wang, M., Liu, J. &amp; Fu, C. R. (2020) A Photographic Guide to the Butterflies of Yinggeling, Hainan. Nan Hai Publishing Company, Haikou, 369 pp. [in Chinese]" type="book" year="2020">
Wang
<emphasis id="3BFFA1167A69A2141A77FF38FBC4FF55" box="[1024,1081,187,213]" italics="true" pageId="4" pageNumber="396">et al.</emphasis>
(2020)
</bibRefCitation>
. However, according to the photographer Hai-Feng Jia (pers. comm.), this individual was actually photographed in
<collectingRegion id="CB4FB3E67A69A2141AD0FF5CFB06FF79" box="[1191,1275,223,249]" country="China" name="Jiangxi" pageId="4" pageNumber="396">Jiangxi</collectingRegion>
, E.
<collectingCountry id="719C3D947A69A2141B5FFF5CFA96FF79" box="[1320,1387,223,249]" name="China" pageId="4" pageNumber="396">China</collectingCountry>
and thus belongs to
<taxonomicName id="CE8B06877A69A2141F3BFE80FE41FE9C" authorityName=": Joicey &amp; Talbot" authorityYear="1928" box="[332,444,259,285]" class="Insecta" family="Nymphalidae" genus="Stibochiona" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="396" phylum="Arthropoda" rank="subSpecies" species="nicea" subSpecies="nicea">
ssp.
<emphasis id="3BFFA1167A69A2141F08FE80FE41FE9C" box="[383,444,259,284]" italics="true" pageId="4" pageNumber="396">nicea</emphasis>
</taxonomicName>
without doubt. 3)
<bibRefCitation id="6D1A00F57A69A2141CE7FE80FC64FE9D" author="Zhang, S. K. &amp; Zhang, H. Q." box="[656,921,259,285]" pageId="4" pageNumber="396" refId="ref3645" refString="Zhang, S. K. &amp; Zhang, H. Q. (2018) An Ecological Illustrated Handbook of Butterflies in Nanjing. Nanjing Normal University Press, Nanjing, 365 pp. [in Chinese]" type="book" year="2018">Zhang &amp; Zhang (2018)</bibRefCitation>
figured an individual from
<collectingRegion id="CB4FB3E67A69A2141AA1FE80FAD0FE9D" box="[1238,1325,259,285]" country="China" name="Jiangsu" pageId="4" pageNumber="396">Jiangsu</collectingRegion>
, E.
<collectingCountry id="719C3D947A69A2141B2EFE80FA61FE9D" box="[1369,1436,259,285]" name="China" pageId="4" pageNumber="396">China</collectingCountry>
which is identical to the new subspecies. However, the source of this specimen cannot be determined by Song-Kui Zhang (pers. comm.). It probably is based on a wrongly labeled specimen from
<collectingRegion id="CB4FB3E67A69A2141A63FEC8FB94FEE5" box="[1044,1129,331,357]" country="China" name="Hainan" pageId="4" pageNumber="396">Hainan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>