treatments-xml/data/FC/50/54/FC50544FA5EE5F8692A716D33BD72276.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="FC50544FA5EE5F8692A716D33BD72276" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Incasarcus viracocha Kury &amp; Maury 1998" docType="treatment" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
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<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:date>2022</mods:date>
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<mods:number>2022-08-16</mods:number>
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3.35. 
<taxonomicName LSID="FC50544F-A5EE-5F86-92A7-16D33BD72276" authority="Kury &amp; Maury, 1998" authorityName="Kury &amp; Maury" authorityYear="1998" class="Arachnida" family="Metasarcidae" genus="Incasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Incasarcus viracocha" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="viracocha">Incasarcus viracocha Kury &amp; Maury, 1998</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">Figs 4F</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male trochanter-patella / tibia IV of Incasarcus. A - B Trochanter-tibia IV of I. argenteus Kury &amp; Maury, 1998, dorsal and ventral view, respectively; C - D Trochanter-tibia IV of I. dianae Kury &amp; Maury, 1998, dorsal and ventral view, respectively; E-F Trochanter-tibia IV of I. pictus Kury &amp; Maury, 1998, dorsal and ventral view, respectively; G - H Trochanter-tibia IV of I. viracocha Kury &amp; Maury, 1998, dorsal and ventral view, respectively; I Trochanter-patella of I. viracocha, lateral view; J Trochanter-patella of I. ochoai Kury &amp; Maury, 1998, large male, dorsal view; K - L Trochanter-tibia of I. ochoai, small male, dorsal and ventral view, respectively; M Trochanter-patella of I. ochoai, large male, lateral view; N Trochanter-patella of I. ochoai, small male, lateral view; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure9" httpUri="https://binary.pensoft.net/fig/729883" pageId="0" pageNumber="309">, 9G-I</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Penis of Huancabamba gen. nov. and Incasarcus, dorsal, lateral and ventral views, respectively. A - C H. kubricki gen. et sp. nov.; D - F I. ochoai Kury &amp; Maury, 1998; G - I I. viracocha Kury &amp; Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure18" httpUri="https://binary.pensoft.net/fig/729892" pageId="0" pageNumber="309">, 18G-I</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Distribution of Incasarcus in Cusco, Peru." figureDoi="10.3897/asp.80.e73829.figure30" httpUri="https://binary.pensoft.net/fig/729904" pageId="0" pageNumber="309">, 30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="reference_group">
<paragraph pageId="0" pageNumber="309">
<taxonomicName authorityName="Kury &amp; Maury" authorityYear="1998" class="Arachnida" family="Metasarcidae" genus="Incasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Incasarcus viracocha" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="viracocha">Incasarcus viracocha</taxonomicName>
Kury &amp; Maury, 1998: 157 (desc.), 160 (key), figs. 38 (male dorsal habitus, chelicerae, pedipalpus, trochanter-patella IV), 39 (penis dorsal view), 40 (penis lateral view), 41 (male lateral habitus), 42 (female dorsal habitus), 43 (male femur IV); 
<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
: 145 (cat.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="redescription">
<paragraph pageId="0" pageNumber="309">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
MALE: 
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) DSW: 4.9-5.0 (4.9); DSL: 5.9-6.6 (5.9); CL: 2.4-3.0 (2.4). FIVL: 11.7-11.8 (11.8). ChL: 3.5-4.2 (4.2). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
(in ethanol): Brownish yellow body with reticular spots covering practically the entire carapace and chelicerae. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">4F</figureCitation>
) Anterior margin of carapace with granular median elevation. Ocularium with very marked median depression; with sparse granules or smooth. Carapace with sparse granules. Areas I-IV with granules throughout their surface. Areas I-II and IV unarmed. Area III with a pair of median spines, facing posteriorly. Lateral and posterior margins of DS with granules throughout their length. Free tergites I-III with a row of granules. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">4F</figureCitation>
) Swollen in large males (as in the holotype). Segment I with densely distributed granules. Segment II with small granules or smooth; finger with five teeth. Segment III with three teeth. 
<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: Covered with sparse granules across the dorsal surface of the femur-tibia. Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of 6-7 small setiferous tubercles and a retro-subapical spine. Patella with a small retroapical setiferous tubercle. Tibia: prolateral iIiIIi, retrolateral IIii. Tarsus: prolateral iiiIiIIi, retrolateral iIiiIi. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with a median row of 5-6 setiferous tubercles. Coxae II-IV with setiferous granules. Rows of four tubercles between the coxae II-III and seven between the coxae III-IV. Genital area with few granules. Free sternites with transverse rows of small granules. Anal few granular. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">4F</figureCitation>
, 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male trochanter-patella / tibia IV of Incasarcus. A - B Trochanter-tibia IV of I. argenteus Kury &amp; Maury, 1998, dorsal and ventral view, respectively; C - D Trochanter-tibia IV of I. dianae Kury &amp; Maury, 1998, dorsal and ventral view, respectively; E-F Trochanter-tibia IV of I. pictus Kury &amp; Maury, 1998, dorsal and ventral view, respectively; G - H Trochanter-tibia IV of I. viracocha Kury &amp; Maury, 1998, dorsal and ventral view, respectively; I Trochanter-patella of I. viracocha, lateral view; J Trochanter-patella of I. ochoai Kury &amp; Maury, 1998, large male, dorsal view; K - L Trochanter-tibia of I. ochoai, small male, dorsal and ventral view, respectively; M Trochanter-patella of I. ochoai, large male, lateral view; N Trochanter-patella of I. ochoai, small male, lateral view; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure9" httpUri="https://binary.pensoft.net/fig/729883" pageId="0" pageNumber="309">9G-I</figureCitation>
) Coxa I with a retrolateral and a prolateral apophysis. Coxa II with a prolateral apophysis. Coxa III unarmed. Coxa IV with sparse setiferous granules. Trochanters I-IV unarmed and granular. Femora I-III unarmed and with sparse granules. Femur IV with small granules; a retroventral row of 30-35 acuminate tubercles, distributed along the entire length of the article, except at the basal and apical ends (the most basal and apical tubercles are smaller than the median ones). Patellae I-III unarmed. Patella IV with a retroapical spiniform apophysis; a ventromedian row of four acuminate tubercles. Tibiae I-IV unarmed and granular. Tarsal formula: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) 11, 21-22 (22), 11, 14-15 (15). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig. 
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Penis of Huancabamba gen. nov. and Incasarcus, dorsal, lateral and ventral views, respectively. A - C H. kubricki gen. et sp. nov.; D - F I. ochoai Kury &amp; Maury, 1998; G - I I. viracocha Kury &amp; Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure18" httpUri="https://binary.pensoft.net/fig/729892" pageId="0" pageNumber="309">18G-I</figureCitation>
) VP rectangular, slightly tapering at the apex, with distal margin straight; straight in lateral view. MS C1-C3(C4) subapical long and apically curved; MS A1 median long and straight (⅓ of the length of the MS C); MS B1-B2 sub basal; MS B1 long and straight, MS B2 long and spatulated; MS D1 short, placed beneath MS C. Lateral sacs robust, with a blunt apex; with long T3-like microsetae. Stylus with slightly broad apex; with ventral projections. Promontory convex and tapered. - 
<emphasis bold="true" pageId="0" pageNumber="309">
FEMALES: 
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) DSW: 4.5-4.9; DSL: 5.5-5.8; CL: 2.0. FIVL: 12.4-12.5. ChL: 2.4-2.5. Chelicerae not swollen. Femur IV unarmed. Pedipalpal femur with five tubercles. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) 9-10, 15-20, 10-12, 12-13.
</paragraph>
<caption doi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309" start="Figure 23" startId="F23">
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">Figure 23.</emphasis>
Live specimens of 
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="0" pageNumber="309">A</emphasis>
<taxonomicName authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho spiniger" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho spiniger</emphasis>
</taxonomicName>
(Roewer, 1957) 
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
, male; 
<emphasis bold="true" pageId="0" pageNumber="309">B</emphasis>
female; 
<emphasis bold="true" pageId="0" pageNumber="309">C</emphasis>
<taxonomicName genus="A." lsidName="A. tapacocha" pageId="0" pageNumber="309" rank="species" species="tapacocha">
<emphasis italics="true" pageId="0" pageNumber="309">A. tapacocha</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">nom. nov.</emphasis>
, male; 
<emphasis bold="true" pageId="0" pageNumber="309">D</emphasis>
female; 
<emphasis bold="true" pageId="0" pageNumber="309">E</emphasis>
<taxonomicName genus="A." lsidName="A. titschacki" pageId="0" pageNumber="309" rank="species" species="titschacki">
<emphasis italics="true" pageId="0" pageNumber="309">A. titschacki</emphasis>
</taxonomicName>
Roewer, 1949, male; 
<emphasis bold="true" pageId="0" pageNumber="309">F</emphasis>
<taxonomicName genus="A." lsidName="A. vargasllosai" pageId="0" pageNumber="309" rank="species" species="vargasllosai">
<emphasis italics="true" pageId="0" pageNumber="309">A. vargasllosai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male; 
<emphasis bold="true" pageId="0" pageNumber="309">G</emphasis>
<taxonomicName genus="A." lsidName="A. uniseriatus" pageId="0" pageNumber="309" rank="species" species="uniseriatus">
<emphasis italics="true" pageId="0" pageNumber="309">A. uniseriatus</emphasis>
</taxonomicName>
(Roewer, 1959) 
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
, male; 
<emphasis bold="true" pageId="0" pageNumber="309">H</emphasis>
: female.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="diagnosis">
<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="309">
It differs from other species of the genus by the set of following characters: DS without white spots on DS; DS granular; ocularium unarmed and with few granules or smooth; areas I, II and IV unarmed; area III with a pair of spines; free tergites I-III with a row of tubercles, unarmed (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury &amp; Maury, 1998; C I. dianae Kury &amp; Maury, 1998; D I. ochoai Kury &amp; Maury, 1998; E I. pictus Kury &amp; Maury, 1998; F I. viracocha Kury &amp; Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">4F</figureCitation>
); male femur IV with a retroventral row of 30-35 acuminate tubercles (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male trochanter-patella / tibia IV of Incasarcus. A - B Trochanter-tibia IV of I. argenteus Kury &amp; Maury, 1998, dorsal and ventral view, respectively; C - D Trochanter-tibia IV of I. dianae Kury &amp; Maury, 1998, dorsal and ventral view, respectively; E-F Trochanter-tibia IV of I. pictus Kury &amp; Maury, 1998, dorsal and ventral view, respectively; G - H Trochanter-tibia IV of I. viracocha Kury &amp; Maury, 1998, dorsal and ventral view, respectively; I Trochanter-patella of I. viracocha, lateral view; J Trochanter-patella of I. ochoai Kury &amp; Maury, 1998, large male, dorsal view; K - L Trochanter-tibia of I. ochoai, small male, dorsal and ventral view, respectively; M Trochanter-patella of I. ochoai, large male, lateral view; N Trochanter-patella of I. ochoai, small male, lateral view; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure9" httpUri="https://binary.pensoft.net/fig/729883" pageId="0" pageNumber="309">9G-I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="distribution">
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="309">
(Fig. 
<figureCitation captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Distribution of Incasarcus in Cusco, Peru." figureDoi="10.3897/asp.80.e73829.figure30" httpUri="https://binary.pensoft.net/fig/729904" pageId="0" pageNumber="309">30</figureCitation>
) PERU. Cusco. Urubamba province.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
<emphasis italics="true" pageId="0" pageNumber="309">Type material</emphasis>
: 
<materialsCitation collectingDate="1994-02-25" collectorName="J. Ochoa, J. Achicahuala" country="PERU" elevation="2350" latitude="-13.116667" location="Machu Picchu" longLatPrecision="1291" longitude="-72.566666" municipality="Urubamba province" specimenCount="♂" stateProvince="Cusco" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
<specimenCount>♂</specimenCount>
, ' 
<collectingCountry name="Peru">PERU</collectingCountry>
, 
<collectingRegion country="Peru" name="Cusco">Cusco</collectingRegion>
, 
<collectingMunicipality>Urubamba province</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:FC50544FA5EE5F8692A716D33BD72276:5F1A1010FC45FD0788E2F2029D2DB7E1" country="PERU" latitude="-13.116667" longLatPrecision="1291" longitude="-72.566666" municipality="Urubamba province" name="Machu Picchu" stateProvince="Cusco">Machu Picchu</location>
, 
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" unit="m" value="2350.0" valueMax="2500.0" valueMin="2200.0">2,200-2,500m</quantity>
a.s.l.
</elevation>
, 
<geoCoordinate degrees="13" direction="south" minutes="07" orientation="latitude" precision="925" value="-13.116667">13°07′S</geoCoordinate>
<geoCoordinate degrees="72" direction="west" minutes="34" orientation="longitude" precision="925" value="-72.566666">72°34′W</geoCoordinate>
, 
<collectingDate value="1994-02-25">25/II/1994</collectingDate>
, 
<collectorName>J. Ochoa</collectorName>
&amp; 
<collectorName>J. Achicahuala</collectorName>
leg. (MACN 9547)
</materialsCitation>
</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount>♀</specimenCount>
, 
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
(MACN 9548); 
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>ParatypeS</typeStatus>
</emphasis>
<specimenCount type="female">2 ♀</specimenCount>
, 
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
(MNRJ 5400). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Additional material</emphasis>
: 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="PERU" location="Cusco'" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<specimenCount type="male">2 ♂</specimenCount>
, 
<specimenCount type="female">3 ♀</specimenCount>
' 
<collectingCountry name="Peru">PERU</collectingCountry>
, 
<normalizedToken originalValue="Cusco’">
<location LSID="urn:lsid:plazi:treatment:FC50544FA5EE5F8692A716D33BD72276:DCAAF5C6DE14CE9A50FC4E0253B383C3" country="PERU" name="Cusco'">Cusco'</location>
</normalizedToken>
(MUBI)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>