235 lines
33 KiB
XML
235 lines
33 KiB
XML
<document id="3CE16B786C78E6C41857AFBC1EF001CE" ID-CLB-Dataset="22824" ID-DOI="10.11646/zootaxa.4764.1.1" ID-GBIF-Dataset="55f6b110-0a37-45fe-be1c-34e9a035fe0e" ID-ISSN="1175-5334" ID-Zenodo-Dep="3764923" ID-ZooBank="urn:lsid:zoobank.org:pub:53C5289A-156C-4AF1-B84A-73099A332C05" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587733789776" checkinUser="carolina" docAuthor="Van Ofwegen, Leen P." docDate="2020" docId="03BCB7172348FFB9FF358173FD27F86F" docLanguage="en" docName="zootaxa.4764.1.1.pdf.imf" docOrigin="Zootaxa 4764 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Litophyton horseshoeensis Van 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="106" masterDocId="FF85CF6F2321FFD0FFA2856EFFC1FFB1" masterDocTitle="The genus Litophyton Forskål, 1775 (Octocorallia: Alcyonacea: Nephtheidae) from Australia" masterLastPageNumber="131" masterPageNumber="1" pageNumber="106" updateTime="1698837010817" updateUser="ExternalLinkService">
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<mods:mods id="D6288C8BDDEBEFE2AF25B917095D3D99" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="38626B50563B15D0C14DF3EEEA9C5178">
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<mods:title id="ADCFE4EB37C41EEB39DCC3B287505E94">The genus Litophyton Forskål, 1775 (Octocorallia: Alcyonacea: Nephtheidae) from Australia</mods:title>
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</mods:titleInfo>
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<mods:name id="B5DC2ED25C78AD1F8BC2276B2E742EF2" type="personal">
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<mods:role id="4C46B10A3113BFE888BC53286358438A">
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<mods:roleTerm id="5084D6EAACFDAF2D1762457BA241A71D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7491187C74996C51471B6A317DFE874A">Van Ofwegen, Leen P.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2364EEC7BEEB61279DEE8FB713FE83B3">text</mods:typeOfResource>
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<mods:titleInfo id="7D48BDB54A296737C90AD903E4F18CD2">
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<mods:title id="BA833A2CF117A4041E187486D1CE42F3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A4FC04BA3945BC29C991B482346E7821">
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<mods:date id="3A341C2B086BAA6676B7C7F3C4ABEDDB">2020</mods:date>
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<mods:detail id="6ADFA0FA1ACD25783D8B91918D5636A0" type="pubDate">
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<mods:number id="6A319C077AA152BD7DF8AD6A09C9F564">2020-04-16</mods:number>
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</mods:detail>
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<mods:detail id="84CACBDED71504C4B1AF0DC82B48921B" type="volume">
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<mods:number id="0AAF44FB012021B69FE6F2F884FB9B56">4764</mods:number>
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</mods:detail>
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<mods:detail id="F14BF1FEAF7EA79FCB7A05FE2C834B3F" type="issue">
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<mods:number id="D38408C3AE98A716C927AA59D47D6C6D">1</mods:number>
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<mods:extent id="DF3559CFCC2476CD68DA2F4F457E2953" unit="page">
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<mods:start id="10C1F78721C85502424E6569A1452750">1</mods:start>
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<mods:end id="D67D1317DD07A5DE42714129CB3AAC9A">131</mods:end>
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<mods:classification id="D71E1513464300314C4CA009E8857698">journal article</mods:classification>
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<mods:identifier id="1F79E82293A5442EDD67F95A293A5450" type="CLB-Dataset">22824</mods:identifier>
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<mods:identifier id="6D3C244D082309FCEAE6C095F36D8F0B" type="DOI">10.11646/zootaxa.4764.1.1</mods:identifier>
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<mods:identifier id="A56749BDFA0CD04ED9D18326183DA954" type="GBIF-Dataset">55f6b110-0a37-45fe-be1c-34e9a035fe0e</mods:identifier>
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<mods:identifier id="C6B781F69FB775588A4B57D9F21202EE" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="D1486EF5EC5BAEC9EAAD0A5C2AA57E6F" type="Zenodo-Dep">3764923</mods:identifier>
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<mods:identifier id="2E1267A55A1967AE6CBA898B6E945B88" type="ZooBank">urn:lsid:zoobank.org:pub:53C5289A-156C-4AF1-B84A-73099A332C05</mods:identifier>
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</mods:mods>
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<treatment id="03BCB7172348FFB9FF358173FD27F86F" ID-DOI="http://doi.org/10.5281/zenodo.3803580" ID-GBIF-Taxon="163737701" ID-Zenodo-Dep="3803580" LSID="urn:lsid:plazi:treatment:03BCB7172348FFB9FF358173FD27F86F" httpUri="http://treatment.plazi.org/id/03BCB7172348FFB9FF358173FD27F86F" lastPageNumber="106" pageId="105" pageNumber="106">
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<subSubSection id="C30F558A2348FFB9FF358173FDF5FB86" box="[151,564,1053,1079]" pageId="105" pageNumber="106" type="nomenclature">
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<paragraph id="8BAA06012348FFB9FF358173FDF5FB86" blockId="105.[151,564,1053,1115]" box="[151,564,1053,1079]" pageId="105" pageNumber="106">
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<heading id="D0E2B16D2348FFB9FF358173FDF5FB86" box="[151,564,1053,1079]" pageId="105" pageNumber="106">
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<taxonomicName id="4C157D822348FFB9FF358173FE11FB86" ID-CoL="866MX" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[151,464,1053,1079]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="105" pageNumber="106" phylum="Cnidaria" rank="species" species="horseshoeensis" status="sp. nov.">
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<emphasis id="B961DA132348FFB9FF358173FE11FB86" box="[151,464,1053,1079]" italics="true" pageId="105" pageNumber="106">Litophyton horseshoeensis</emphasis>
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</taxonomicName>
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<emphasis id="B961DA132348FFB9FE758173FDF5FB86" bold="true" box="[471,564,1053,1079]" pageId="105" pageNumber="106">
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<taxonomicNameLabel id="A25267682348FFB9FE758173FDF5FB86" box="[471,564,1053,1079]" pageId="105" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F558A2348FFB9FF35812EFE53FBEA" box="[151,402,1088,1115]" pageId="105" pageNumber="106" type="description">
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<paragraph id="8BAA06012348FFB9FF35812EFE53FBEA" blockId="105.[151,564,1053,1115]" box="[151,402,1088,1115]" pageId="105" pageNumber="106">
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(
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<figureCitation id="132E1A842348FFB9FF3D812EFEC9FBEA" box="[159,264,1088,1115]" captionStart="FIGURE 85" captionStartId="106.[151,248,1418,1443]" captionTargetBox="[158,1420,189,1384]" captionTargetId="figure@106.[151,1436,181,1393]" captionTargetPageId="106" captionText="FIGURE 85. Lateral views polyp armature of Litophyton species: a-b, L. folium n. sp., a, holotype NTM C4128, b, paratype NTM C10456; c, L. fusum n. sp., holotype NTM C4366; d; L. horseshoeensis, n. sp., holotype NTM C2256; e-f, L. sanctuaryensis n. sp., e, holotype NTM C1387; f, paratype NTM C4383; g. L. swainensis n. sp., holotype NTM C1371; h, L. verrucosum n. sp., holotype NTM C4071." figureDoi="http://doi.org/10.5281/zenodo.3765101" httpUri="https://zenodo.org/record/3765101/files/figure.png" pageId="105" pageNumber="106">Figs. 85d</figureCitation>
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,
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<figureCitation id="132E1A842348FFB9FEB1812EFEFFFBEB" box="[275,318,1088,1114]" captionStart="FIGURE 89" captionStartId="110.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@110.[178,1409,181,1919]" captionTargetPageId="110" captionText="FIGURE 89. Litophyton type specimens: a, L. fusum n. sp., holotype NTM C4366; b, L. horseshoeensis n. sp., holotype NTM C2256; c-d, L. mollis (Macfadyen, 1936), c, holotype BMNH 1934.3.28.51, d, NTM C2262. Scale at d 1 cm, all others natural size." figureDoi="http://doi.org/10.5281/zenodo.3765109" httpUri="https://zenodo.org/record/3765109/files/figure.png" pageId="105" pageNumber="106">89b</figureCitation>
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,
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<figureCitation id="132E1A842348FFB9FEEB812FFE4BFBEA" box="[329,394,1089,1115]" captionStart-0="FIGURE 92" captionStart-1="FIGURE 93" captionStartId-0="113.[151,248,1945,1970]" captionStartId-1="114.[151,250,1248,1273]" captionTargetBox-0="[178,1409,181,1919]" captionTargetBox-1="[151,1435,181,1222]" captionTargetId-0="figure@113.[178,1409,181,1919]" captionTargetId-1="figure@114.[151,1435,181,1222]" captionTargetPageId-0="113" captionTargetPageId-1="114" captionText-0="FIGURE 92. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, abaxial point spindles; b, lateral point spindles; c, adaxial and tentacle scales; d, polyp stalk rods; e, supporting bundle spindle, partly; f-g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk. Scale at a also applies to e, g; scale at b also applies to c, f, h." captionText-1="FIGURE 93. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, sclerites of surface layer base of stalk; b-c, spindles of interior base of stalk; d, tuberculation of interior base stalk spindle. Scale at a also applies to b." figureDoi-0="http://doi.org/10.5281/zenodo.3765115" figureDoi-1="http://doi.org/10.5281/zenodo.3765117" httpUri-0="https://zenodo.org/record/3765115/files/figure.png" httpUri-1="https://zenodo.org/record/3765117/files/figure.png" pageId="105" pageNumber="106">92-93</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F558A2348FFB9FF3581E6FE93FAE7" pageId="105" pageNumber="106" type="materials_examined">
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<paragraph id="8BAA06012348FFB9FF3581E6FE93FAE7" blockId="105.[151,1437,1160,2015]" pageId="105" pageNumber="106">
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<emphasis id="B961DA132348FFB9FF3581E6FEB4FB12" bold="true" box="[151,373,1160,1187]" pageId="105" pageNumber="106">Material examined</emphasis>
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.
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<materialsCitation id="3B7D0C5C2348FFB9FE2681E6FEEFFB77" ID-GBIF-Occurrence="2608549241" collectingDate="1980-11-11" collectionCode="NTM" collectorName="P. Alderslade" location="Bommie Top" municipality="Horseshoe" pageId="105" pageNumber="106" specimenCode="NTM C2256" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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<typeStatus id="54AEB8A32348FFB9FE2681E6FE2BFB13" box="[388,490,1160,1186]" pageId="105" pageNumber="106" type="holotype">Holotype</typeStatus>
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<specimenCode id="DBB3AE7A2348FFB9FE5381E7FD44FB12" box="[497,645,1160,1187]" pageId="105" pageNumber="106">NTM C2256</specimenCode>
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,
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<collectingRegion id="49D1C8E32348FFB9FD3081E6FCD8FB13" box="[658,793,1160,1186]" country="Australia" name="Queensland" pageId="105" pageNumber="106">Queensland</collectingRegion>
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,
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<collectingMunicipality id="6BCE9C7B2348FFB9FC8481E6FC5DFB13" box="[806,924,1160,1186]" pageId="105" pageNumber="106">Horseshoe</collectingMunicipality>
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reef,
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<location id="8ECA50DA2348FFB9FC7D81E6FBB4FB12" LSID="urn:lsid:plazi:treatment:03BCB7172348FFB9FF358173FD27F86F:8ECA50DA2348FFB9FC7D81E6FBB4FB12" box="[991,1141,1160,1187]" municipality="Horseshoe" name="Bommie Top" pageId="105" pageNumber="106" stateProvince="Queensland">Bommie Top</location>
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,
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<date id="FFAB20C12348FFB9FB2081E7FA9EFB12" box="[1154,1375,1160,1187]" pageId="105" pageNumber="106" value="1980-11-11">
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<collectingDate id="EFEFD9292348FFB9FB2081E7FA9EFB12" box="[1154,1375,1160,1187]" pageId="105" pageNumber="106" value="1980-11-11">11 November 1980</collectingDate>
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</date>
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, coll.
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<collectorName id="26E063D72348FFB9FF3581C3FEEFFB77" box="[151,302,1196,1223]" pageId="105" pageNumber="106">P. Alderslade</collectorName>
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</materialsCitation>
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;
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<typeStatus id="54AEB8A32348FFB9FE9981C3FE6AFB76" box="[315,427,1197,1223]" pageId="105" pageNumber="106" type="paratype">paratypes</typeStatus>
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: NTM C
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<date id="FFAB20C12348FFB9FDAC81C2FDAAFB77" box="[526,619,1196,1222]" pageId="105" pageNumber="106" value="1366" valueMax="1368">1366-68</date>
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, 1370, 1379,
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<collectingRegion id="49D1C8E32348FFB9FCA081C2FC47FB77" box="[770,902,1196,1222]" country="Australia" name="Queensland" pageId="105" pageNumber="106">Queensland</collectingRegion>
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, Swain reef, Sweet-lip reef,
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<date id="FFAB20C12348FFB9FB6281C2FA59FB76" box="[1216,1432,1196,1223]" pageId="105" pageNumber="106" value="1980-11-15">15 November 1980</date>
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, coll. P. Alderslade; NTM C
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<date id="FFAB20C12348FFB9FE6981BEFDEAFB5A" box="[459,555,1232,1259]" pageId="105" pageNumber="106" value="1392" valueMax="1395">1392-95</date>
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,
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<collectingRegion id="49D1C8E32348FFB9FD9A81BEFD7EFB5B" box="[568,703,1232,1258]" country="Australia" name="Queensland" pageId="105" pageNumber="106">Queensland</collectingRegion>
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, Bills reef,
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<date id="FFAB20C12348FFB9FCE581BFFBD8FB5A" box="[839,1049,1232,1259]" pageId="105" pageNumber="106" value="1980-11-08">8 November 1980</date>
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, coll. P. Alderslade; NTM C1396,
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<collectingRegion id="49D1C8E32348FFB9FF35819AFEDAFABF" box="[151,283,1268,1294]" country="Australia" name="Queensland" pageId="105" pageNumber="106">Queensland</collectingRegion>
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; Central reef, coll. P. Alderslade; NTM C2255;
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<collectingRegion id="49D1C8E32348FFB9FC82819AFC66FABF" box="[800,935,1268,1294]" country="Australia" name="Queensland" pageId="105" pageNumber="106">Queensland</collectingRegion>
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, Horseshoe reef, Bommie Top,
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<date id="FFAB20C12348FFB9FAAA819BFF11FA82" pageId="105" pageNumber="106" value="1980-11-11">11 November 1980</date>
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, coll. P. Alderslade; NTM C3992,
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<collectingRegion id="49D1C8E32348FFB9FDF28076FD16FA83" box="[592,727,1304,1330]" country="Australia" name="Queensland" pageId="105" pageNumber="106">Queensland</collectingRegion>
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, Great Barrier Reef, Davies reef, depth
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<quantity id="4CEDABE42348FFB9FB3B8076FB2DFA82" box="[1177,1260,1304,1331]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="105" pageNumber="106" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">5-10 m</quantity>
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,
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<date id="FFAB20C12348FFB9FB5A8076FA56FA82" box="[1272,1431,1304,1331]" pageId="105" pageNumber="106" value="1981-04-16">16 April 1981</date>
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, coll. Z. Dinesen.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F558A2348FFB9FF65800EFC85F89A" pageId="105" pageNumber="106" type="description">
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<paragraph id="8BAA06012348FFB9FF65800EFCD6FA2F" blockId="105.[151,1437,1160,2015]" pageId="105" pageNumber="106">
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<emphasis id="B961DA132348FFB9FF65800EFE8FFACA" bold="true" box="[199,334,1376,1403]" pageId="105" pageNumber="106">Description</emphasis>
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. The
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<typeStatus id="54AEB8A32348FFB9FE2F800EFE2DFACB" box="[397,492,1376,1402]" pageId="105" pageNumber="106" type="holotype">holotype</typeStatus>
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is
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<quantity id="4CEDABE42348FFB9FDAE800EFD96FACA" box="[524,599,1376,1403]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="105" pageNumber="106" unit="cm" value="7.5">7.5 cm</quantity>
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high and
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<quantity id="4CEDABE42348FFB9FD66800EFCCEFACA" box="[708,783,1376,1403]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="105" pageNumber="106" unit="cm" value="4.5">4.5 cm</quantity>
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wide, the stalk is
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<quantity id="4CEDABE42348FFB9FC76800EFBDEFACA" box="[980,1055,1376,1403]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="105" pageNumber="106" unit="cm" value="1.5">1.5 cm</quantity>
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long (
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<figureCitation id="132E1A842348FFB9FBC7800EFB04FACA" box="[1125,1221,1376,1403]" captionStart="FIGURE 89" captionStartId="110.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@110.[178,1409,181,1919]" captionTargetPageId="110" captionText="FIGURE 89. Litophyton type specimens: a, L. fusum n. sp., holotype NTM C4366; b, L. horseshoeensis n. sp., holotype NTM C2256; c-d, L. mollis (Macfadyen, 1936), c, holotype BMNH 1934.3.28.51, d, NTM C2262. Scale at d 1 cm, all others natural size." figureDoi="http://doi.org/10.5281/zenodo.3765109" httpUri="https://zenodo.org/record/3765109/files/figure.png" pageId="105" pageNumber="106">Fig. 89b</figureCitation>
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). The polyps are up to
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<quantity id="4CEDABE42348FFB9FF1180EAFEC6FA2E" box="[179,263,1412,1439]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="105" pageNumber="106" unit="mm" value="0.6">0.6 mm</quantity>
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wide and 0.5 high, grouped in spherical catkins.
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</paragraph>
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<paragraph id="8BAA06012348FFB9FF6580C7FBEDF9E2" blockId="105.[151,1437,1160,2015]" pageId="105" pageNumber="106">
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<emphasis id="B961DA132348FFB9FF6580C7FED5FA72" bold="true" box="[199,276,1449,1475]" pageId="105" pageNumber="106">Polyps</emphasis>
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. With irregular arrangement of sclerites in eight points (
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<figureCitation id="132E1A842348FFB9FCDF80C6FC1CFA72" box="[893,989,1448,1475]" captionStart="FIGURE 85" captionStartId="106.[151,248,1418,1443]" captionTargetBox="[158,1420,189,1384]" captionTargetId="figure@106.[151,1436,181,1393]" captionTargetPageId="106" captionText="FIGURE 85. Lateral views polyp armature of Litophyton species: a-b, L. folium n. sp., a, holotype NTM C4128, b, paratype NTM C10456; c, L. fusum n. sp., holotype NTM C4366; d; L. horseshoeensis, n. sp., holotype NTM C2256; e-f, L. sanctuaryensis n. sp., e, holotype NTM C1387; f, paratype NTM C4383; g. L. swainensis n. sp., holotype NTM C1371; h, L. verrucosum n. sp., holotype NTM C4071." figureDoi="http://doi.org/10.5281/zenodo.3765101" httpUri="https://zenodo.org/record/3765101/files/figure.png" pageId="105" pageNumber="106">Fig. 85d</figureCitation>
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). Abaxially and lateral point sclerites are spindles with simple tubercles and spines; abaxially up to
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<quantity id="4CEDABE42348FFB9FC8280A2FC43FA56" box="[800,898,1484,1511]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="105" pageNumber="106" unit="mm" value="0.5">0.50 mm</quantity>
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long, laterally they are up to
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<quantity id="4CEDABE42348FFB9FB6580A2FAE8FA56" box="[1223,1321,1484,1511]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="105" pageNumber="106" unit="mm" value="0.25">0.25 mm</quantity>
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long (
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<figureCitation id="132E1A842348FFB9FAD280A2FF75F9BA" captionStart="FIGURE 92" captionStartId="113.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@113.[178,1409,181,1919]" captionTargetPageId="113" captionText="FIGURE 92. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, abaxial point spindles; b, lateral point spindles; c, adaxial and tentacle scales; d, polyp stalk rods; e, supporting bundle spindle, partly; f-g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk. Scale at a also applies to e, g; scale at b also applies to c, f, h." figureDoi="http://doi.org/10.5281/zenodo.3765115" httpUri="https://zenodo.org/record/3765115/files/figure.png" pageId="105" pageNumber="106">Fig. 92</figureCitation>
|
|
a-b). Adaxially there are small, curved scales, up to
|
|
<quantity id="4CEDABE42348FFB9FD52809FFC93F9BA" box="[752,850,1521,1547]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="105" pageNumber="106" unit="mm" value="0.1">0.10 mm</quantity>
|
|
long, similar to those of the tentacles (
|
|
<figureCitation id="132E1A842348FFB9FAA2809EFA9EF9BA" box="[1280,1375,1520,1547]" captionStart="FIGURE 92" captionStartId="113.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@113.[178,1409,181,1919]" captionTargetPageId="113" captionText="FIGURE 92. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, abaxial point spindles; b, lateral point spindles; c, adaxial and tentacle scales; d, polyp stalk rods; e, supporting bundle spindle, partly; f-g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk. Scale at a also applies to e, g; scale at b also applies to c, f, h." figureDoi="http://doi.org/10.5281/zenodo.3765115" httpUri="https://zenodo.org/record/3765115/files/figure.png" pageId="105" pageNumber="106">Fig. 92c</figureCitation>
|
|
). The polyp stalk has straight rods, up to
|
|
<quantity id="4CEDABE42348FFB9FDB2837BFDB1F99E" box="[528,624,1557,1583]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="105" pageNumber="106" unit="mm" value="0.1">0.10 mm</quantity>
|
|
long (
|
|
<figureCitation id="132E1A842348FFB9FD17837AFCD4F99E" box="[693,789,1556,1583]" captionStart="FIGURE 92" captionStartId="113.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@113.[178,1409,181,1919]" captionTargetPageId="113" captionText="FIGURE 92. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, abaxial point spindles; b, lateral point spindles; c, adaxial and tentacle scales; d, polyp stalk rods; e, supporting bundle spindle, partly; f-g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk. Scale at a also applies to e, g; scale at b also applies to c, f, h." figureDoi="http://doi.org/10.5281/zenodo.3765115" httpUri="https://zenodo.org/record/3765115/files/figure.png" pageId="105" pageNumber="106">Fig. 92d</figureCitation>
|
|
). Supporting bundle not projecting or slightly so, composed of spindles, up to
|
|
<quantity id="4CEDABE42348FFB9FEFB8357FE7BF9E2" box="[345,442,1593,1619]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="105" pageNumber="106" unit="mm" value="1.8">1.80 mm</quantity>
|
|
long, with simple tubercles and one spiny end (
|
|
<figureCitation id="132E1A842348FFB9FC638356FBE1F9E2" box="[961,1056,1592,1619]" captionStart="FIGURE 92" captionStartId="113.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@113.[178,1409,181,1919]" captionTargetPageId="113" captionText="FIGURE 92. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, abaxial point spindles; b, lateral point spindles; c, adaxial and tentacle scales; d, polyp stalk rods; e, supporting bundle spindle, partly; f-g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk. Scale at a also applies to e, g; scale at b also applies to c, f, h." figureDoi="http://doi.org/10.5281/zenodo.3765115" httpUri="https://zenodo.org/record/3765115/files/figure.png" pageId="105" pageNumber="106">Fig. 92e</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="8BAA06012348FFB9FF658332FAF6F92B" blockId="105.[151,1437,1160,2015]" pageId="105" pageNumber="106">
|
|
<emphasis id="B961DA132348FFB9FF658332FE1DF9C6" bold="true" box="[199,476,1628,1655]" pageId="105" pageNumber="106">Surface layer top stalk.</emphasis>
|
|
Capstans, derivatives of capstans, spindles, and unilaterally spinose spindles, all with simple and complex tubercles (
|
|
<figureCitation id="132E1A842348FFB9FE4883EEFDFBF92A" box="[490,570,1664,1691]" captionStart="FIGURE 92" captionStartId="113.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@113.[178,1409,181,1919]" captionTargetPageId="113" captionText="FIGURE 92. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, abaxial point spindles; b, lateral point spindles; c, adaxial and tentacle scales; d, polyp stalk rods; e, supporting bundle spindle, partly; f-g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk. Scale at a also applies to e, g; scale at b also applies to c, f, h." figureDoi="http://doi.org/10.5281/zenodo.3765115" httpUri="https://zenodo.org/record/3765115/files/figure.png" pageId="105" pageNumber="106">Fig. 92</figureCitation>
|
|
f-g). Capstans
|
|
<quantity id="4CEDABE42348FFB9FD7583EEFCB0F92A" box="[727,881,1664,1691]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="105" pageNumber="106" unit="mm" value="0.125" valueMax="0.2" valueMin="0.05">0.05-0.20 mm</quantity>
|
|
long; spindles up to about
|
|
<quantity id="4CEDABE42348FFB9FB3583EEFB39F92A" box="[1175,1272,1664,1691]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="105" pageNumber="106" unit="mm" value="0.6">0.60 mm</quantity>
|
|
long.
|
|
</paragraph>
|
|
<paragraph id="8BAA06012348FFB9FF6583CAFA5AF953" blockId="105.[151,1437,1160,2015]" pageId="105" pageNumber="106">
|
|
<emphasis id="B961DA132348FFB9FF6583CAFE1AF90E" bold="true" box="[199,475,1700,1727]" pageId="105" pageNumber="106">Surface layer base stalk</emphasis>
|
|
. Capstans, thorny derivatives of capstans, spindles, and unilaterally spinose spindles, all with simple and complex tubercles (
|
|
<figureCitation id="132E1A842348FFB9FD8383A6FD4BF953" box="[545,650,1736,1762]" captionStart="FIGURE 92" captionStartId="113.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@113.[178,1409,181,1919]" captionTargetPageId="113" captionText="FIGURE 92. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, abaxial point spindles; b, lateral point spindles; c, adaxial and tentacle scales; d, polyp stalk rods; e, supporting bundle spindle, partly; f-g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk. Scale at a also applies to e, g; scale at b also applies to c, f, h." figureDoi="http://doi.org/10.5281/zenodo.3765115" httpUri="https://zenodo.org/record/3765115/files/figure.png" pageId="105" pageNumber="106">Figs. 92h</figureCitation>
|
|
,
|
|
<figureCitation id="132E1A842348FFB9FD3783A7FD7EF952" box="[661,703,1737,1763]" captionStart="FIGURE 93" captionStartId="114.[151,250,1248,1273]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure@114.[151,1435,181,1222]" captionTargetPageId="114" captionText="FIGURE 93. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, sclerites of surface layer base of stalk; b-c, spindles of interior base of stalk; d, tuberculation of interior base stalk spindle. Scale at a also applies to b." figureDoi="http://doi.org/10.5281/zenodo.3765117" httpUri="https://zenodo.org/record/3765117/files/figure.png" pageId="105" pageNumber="106">93a</figureCitation>
|
|
). Capstans
|
|
<quantity id="4CEDABE42348FFB9FC9E83A6FC14F952" box="[828,981,1736,1763]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="105" pageNumber="106" unit="mm" value="0.1" valueMax="0.15" valueMin="0.05">0.05-0.15 mm</quantity>
|
|
long; spindles up to about
|
|
<quantity id="4CEDABE42348FFB9FB5983A6FA9CF952" box="[1275,1373,1736,1763]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="105" pageNumber="106" unit="mm" value="0.5">0.50 mm</quantity>
|
|
long.
|
|
</paragraph>
|
|
<paragraph id="8BAA06012348FFB9FF658382FC85F89A" blockId="105.[151,1437,1160,2015]" pageId="105" pageNumber="106">
|
|
<emphasis id="B961DA132348FFB9FF658382FEA2F8B6" bold="true" box="[199,355,1772,1799]" pageId="105" pageNumber="106">Interior stalk</emphasis>
|
|
. Spindles, up to
|
|
<quantity id="4CEDABE42348FFB9FDBF8383FD9CF8B6" box="[541,605,1773,1799]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="105" pageNumber="106" unit="mm" value="1.0">1 mm</quantity>
|
|
long, with pointed ends (
|
|
<figureCitation id="132E1A842348FFB9FCD58382FC14F8B6" box="[887,981,1772,1799]" captionStart="FIGURE 93" captionStartId="114.[151,250,1248,1273]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure@114.[151,1435,181,1222]" captionTargetPageId="114" captionText="FIGURE 93. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, sclerites of surface layer base of stalk; b-c, spindles of interior base of stalk; d, tuberculation of interior base stalk spindle. Scale at a also applies to b." figureDoi="http://doi.org/10.5281/zenodo.3765117" httpUri="https://zenodo.org/record/3765117/files/figure.png" pageId="105" pageNumber="106">Fig. 93c</figureCitation>
|
|
), and simple and complex tubercles (
|
|
<figureCitation id="132E1A842348FFB9FAD28382FF05F89A" captionStart="FIGURE 93" captionStartId="114.[151,250,1248,1273]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure@114.[151,1435,181,1222]" captionTargetPageId="114" captionText="FIGURE 93. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, sclerites of surface layer base of stalk; b-c, spindles of interior base of stalk; d, tuberculation of interior base stalk spindle. Scale at a also applies to b." figureDoi="http://doi.org/10.5281/zenodo.3765117" httpUri="https://zenodo.org/record/3765117/files/figure.png" pageId="105" pageNumber="106">Fig. 93b</figureCitation>
|
|
). The smaller spindles often have side branches (
|
|
<figureCitation id="132E1A842348FFB9FD7A827EFCF9F89A" box="[728,824,1808,1835]" captionStart="FIGURE 93" captionStartId="114.[151,250,1248,1273]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure@114.[151,1435,181,1222]" captionTargetPageId="114" captionText="FIGURE 93. Litophyton horseshoeensis, n. sp., holotype NTM C2256; a, sclerites of surface layer base of stalk; b-c, spindles of interior base of stalk; d, tuberculation of interior base stalk spindle. Scale at a also applies to b." figureDoi="http://doi.org/10.5281/zenodo.3765117" httpUri="https://zenodo.org/record/3765117/files/figure.png" pageId="105" pageNumber="106">Fig. 93b</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30F558A2348FFB9FF65825BFD07F8FF" box="[199,710,1844,1871]" pageId="105" pageNumber="106" type="etymology">
|
|
<paragraph id="8BAA06012348FFB9FF65825BFD07F8FF" blockId="105.[151,1437,1160,2015]" box="[199,710,1844,1871]" pageId="105" pageNumber="106">
|
|
<emphasis id="B961DA132348FFB9FF65825BFE84F8FE" bold="true" box="[199,325,1845,1871]" pageId="105" pageNumber="106">Etymology</emphasis>
|
|
. Named after the
|
|
<typeStatus id="54AEB8A32348FFB9FDAB825AFDA9F8FF" box="[521,616,1844,1870]" pageId="105" pageNumber="106" type="holotype">holotype</typeStatus>
|
|
locality.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30F558A2348FFB9FF658236FAE9F8C3" box="[199,1320,1880,1907]" pageId="105" pageNumber="106" type="distribution">
|
|
<paragraph id="8BAA06012348FFB9FF658236FAE9F8C3" blockId="105.[151,1437,1160,2015]" box="[199,1320,1880,1907]" pageId="105" pageNumber="106">
|
|
<emphasis id="B961DA132348FFB9FF658236FE94F8C2" bold="true" box="[199,341,1880,1907]" pageId="105" pageNumber="106">Distribution</emphasis>
|
|
. Great Barrier Reef: Horseshoe reef, Sweet-lip reef, Bills reef, Central reef, Davies reef.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C30F558A2348FFB9FF658213FD27F86F" pageId="105" pageNumber="106" type="discussion">
|
|
<paragraph id="8BAA06012348FFB9FF658213FB56F827" blockId="105.[151,1437,1160,2015]" box="[199,1175,1916,1943]" pageId="105" pageNumber="106">
|
|
<emphasis id="B961DA132348FFB9FF658213FEF8F826" bold="true" box="[199,313,1917,1943]" pageId="105" pageNumber="106">Remarks.</emphasis>
|
|
NTM C1379 has somewhat longer internal stalk spindles, up to
|
|
<quantity id="4CEDABE42348FFB9FC558213FB99F826" box="[1015,1112,1917,1943]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="105" pageNumber="106" unit="mm" value="1.2">1.20 mm</quantity>
|
|
long.
|
|
</paragraph>
|
|
<paragraph id="8BAA06012348FFB9FF6582CFFD27F86F" blockId="105.[151,1437,1160,2015]" pageId="105" pageNumber="106">
|
|
<taxonomicName id="4C157D822348FFB9FF6582CFFE4EF80B" authorityName="Van" authorityYear="2020" box="[199,399,1952,1979]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="105" pageNumber="106" phylum="Cnidaria" rank="species" species="horseshoeensis">
|
|
<emphasis id="B961DA132348FFB9FF6582CFFF16F80B" box="[199,215,1953,1978]" italics="true" pageId="105" pageNumber="106">L</emphasis>
|
|
.
|
|
<emphasis id="B961DA132348FFB9FF4782CEFE4EF80B" box="[229,399,1952,1978]" italics="true" pageId="105" pageNumber="106">horseshoeensis</emphasis>
|
|
</taxonomicName>
|
|
resembles
|
|
<taxonomicName id="4C157D822348FFB9FDAE82CFFC6CF80A" authority="(Kukenthal, 1895) (Ternate)" baseAuthorityName="Kukenthal" baseAuthorityYear="1895" box="[524,941,1952,1979]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="105" pageNumber="106" phylum="Cnidaria" rank="species" species="debilis">
|
|
<emphasis id="B961DA132348FFB9FDAE82CFFDB2F80B" box="[524,627,1952,1978]" italics="true" pageId="105" pageNumber="106">L. debilis</emphasis>
|
|
(
|
|
<bibRefCitation id="EF847BF02348FFB9FD2382CEFCFAF80A" author="Kukenthal" box="[641,827,1952,1979]" firstAuthor="Kukenthal" pageId="105" pageNumber="106" pagination="426 - 436" refId="ref42159" refString="Kukenthal, W. (1895) Alcyonaceen von Ternate. Fam. Nephthyidae Verrill. Zoologische Anzeiger, 488, 426 - 436." type="journal article" year="1895">Kükenthal, 1895</bibRefCitation>
|
|
) (
|
|
<collectingCountry id="F30246912348FFB9FCF082CFFC66F80A" box="[850,935,1953,1979]" name="Indonesia" pageId="105" pageNumber="106">Ternate</collectingCountry>
|
|
)
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="132E1A842348FFB9FC1F82CEFBE7F80B" box="[957,1062,1952,1978]" captionStart="FIGURE 69" captionStartId="88.[151,248,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@88.[178,1409,181,1919]" captionTargetPageId="88" captionText="FIGURE 69. Litophyton type specimens: a-c, L. brachiatum n. sp., a, holotype NTM C4154, b, paratype NTM C1473, c, paratype NTM C4179; d, L. bayeri (Verseveldt, 1966), syntypes RMNH 2239; e, L. debilis (Kükenthal, 1895) holotype SMF 64; f, L.burfordensis n. sp., holotype NTM C3171. All natural size." figureDoi="http://doi.org/10.5281/zenodo.3765067" httpUri="https://zenodo.org/record/3765067/files/figure.png" pageId="105" pageNumber="106">Figs. 69e</figureCitation>
|
|
,
|
|
<figureCitation id="132E1A842348FFB9FB9082CFFB91F80A" box="[1074,1104,1953,1979]" captionStart="FIGURE 77" captionStartId="96.[151,248,1910,1935]" captionTargetBox="[193,1394,186,1882]" captionTargetId="figure@96.[190,1397,181,1884]" captionTargetPageId="96" captionText="FIGURE 77. Litophyton debilis (Kükenthal, 1895), holotype SMF 64; a, lateral view polyp armature; b, abaxial view of polyp armature; c, abaxial point spindles; d, adaxial and tentacle scales; e, polyp stalk rods; f, supporting bundle spindle, partly; g, sclerites of surface layer top of stalk; h, sclerites of surface layer base of stalk; i-j, spindles of interior base of stalk. Scale at f also applies to c, g-i; scale at d also applies to e." figureDoi="http://doi.org/10.5281/zenodo.3765085" httpUri="https://zenodo.org/record/3765085/files/figure.png" pageId="105" pageNumber="106">77</figureCitation>
|
|
), but has many thorny derivatives of capstans in the surface of the base of the stalk.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |