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<mods:title id="37842FC8FF7C056DDE04556824549B86">Description of the larvae of four Japanese Platambus Thomson, 1859 (Coleoptera: Dytiscidae: Agabinae) with phylogenetic considerations</mods:title>
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<mods:namePart id="EA3870DF5ADF7143551FB097D33D6E14">Okada, Ryohei</mods:namePart>
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<mods:namePart id="80A1CC7F966A6061A4C572F490D04A92">Alarie, Yves</mods:namePart>
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<mods:namePart id="734F3B0DB58BBC54D1D69A017C3FFC83">Michat, Mariano C.</mods:namePart>
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<treatment id="03BC2A5DFF93FFE3ADD3AA76FB13FE22" ID-GBIF-Taxon="159413417" LSID="urn:lsid:plazi:treatment:03BC2A5DFF93FFE3ADD3AA76FB13FE22" httpUri="http://treatment.plazi.org/id/03BC2A5DFF93FFE3ADD3AA76FB13FE22" lastPageId="17" lastPageNumber="418" pageId="13" pageNumber="414">
<subSubSection id="C30FC8C0FF93FFFFADD3AA76FC7DFD2E" box="[151,937,621,648]" pageId="13" pageNumber="414" type="nomenclature">
<paragraph id="8BAA9B4BFF93FFFFADD3AA76FC7DFD2E" blockId="13.[151,937,621,648]" box="[151,937,621,648]" pageId="13" pageNumber="414">
<heading id="D0E22C27FF93FFFFADD3AA76FC7DFD2E" bold="true" box="[151,937,621,648]" fontSize="11" level="1" pageId="13" pageNumber="414" reason="1">
<emphasis id="B9614759FF93FFFFADD3AA76FC7DFD2E" bold="true" box="[151,937,621,648]" pageId="13" pageNumber="414">
Description of the larvae of
<taxonomicName id="4C15E0C8FF93FFFFACB7AA76FC7DFD2E" ID-CoL="4JMRZ" authority="(Sharp, 1884)" baseAuthorityName="Sharp" baseAuthorityYear="1884" box="[499,937,621,648]" class="Insecta" family="Dytiscidae" genus="Platambus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B9614759FF93FFFFACB7AA76FD2DFD21" bold="true" box="[499,761,621,648]" italics="true" pageId="13" pageNumber="414">Platambus fimbriatus</emphasis>
(
<bibRefCitation id="EF84E6BAFF93FFFFAE4CAA76FC75FD2E" author="Sharp, D." box="[776,929,621,647]" pageId="13" pageNumber="414" pagination="439 - 464" refId="ref16064" refString="Sharp, D. (1884) XXI. The water-beetles of Japan. Transactions of the Entomological Society of London, 1884, 439 - 464." type="journal article" year="1884">Sharp, 1884</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FC8C0FF93FFFFADD3AAAFFE51FCF6" pageId="13" pageNumber="414" type="materials_examined">
<paragraph id="8BAA9B4BFF93FFFFADD3AAAFFE51FCF6" blockId="13.[151,1437,692,1979]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFADD3AAAFFE55FD66" bold="true" box="[151,385,692,719]" pageId="13" pageNumber="414">Source of material.</emphasis>
1 instar I, 1 instar II, 10 instar III,
<collectingCountry id="F302DBDBFF93FFFFAE63AAAEFCBFFD66" box="[807,875,693,719]" name="Japan" pageId="13" pageNumber="414">Japan</collectingCountry>
: Echi riv., Tanesato, Notogawa-cho,
<collectingRegion id="49D155A9FF93FFFFA851AAAEFA4CFD66" box="[1301,1432,693,719]" country="Japan" name="Shiga" pageId="13" pageNumber="414">Shiga Pref.</collectingRegion>
,
<date id="FFABBD8BFF93FFFFADD3AAC3FEDBFD5A" box="[151,271,728,755]" pageId="13" pageNumber="414" value="2015-02-22">22.II.2015</date>
;
<date id="FFABBD8BFF93FFFFAC5FAAC3FE7CFD5A" box="[283,424,728,755]" pageId="13" pageNumber="414" value="2015-12-20">20.XII.2015</date>
;
<date id="FFABBD8BFF93FFFFACF7AAC2FDF5FD5A" box="[435,545,729,755]" pageId="13" pageNumber="414" value="2016-01-10">10.I.2016</date>
; R. Okada leg;
<materialsCitation id="3B7D9116FF93FFFFAF95AAC3FD63FCBE" ID-GBIF-Occurrence="2382714967" collectingDate="2017-01-08" collectorName="R. Okada &amp; Larvae" country="Japan" county="Abukuma" location="Saigo-mura" municipality="Tsuruo" pageId="13" pageNumber="414" specimenCount="1" stateProvince="Fukushima">
5 instar I, 1 instar II,
<collectingCountry id="F302DBDBFF93FFFFAE85AAC2FBD1FD5A" box="[961,1029,729,755]" name="Japan" pageId="13" pageNumber="414">Japan</collectingCountry>
:
<collectingCounty id="62CBE3C7FF93FFFFA94AAAC2FBA8FD5A" box="[1038,1148,729,755]" pageId="13" pageNumber="414">Abukuma</collectingCounty>
riv.,
<collectingMunicipality id="6BCE0131FF93FFFFA9F3AAC2FAD3FD5A" box="[1207,1287,729,755]" pageId="13" pageNumber="414">Tsuruo</collectingMunicipality>
,
<location id="8ECACD90FF93FFFFA857AAC2FA4DFD5A" LSID="urn:lsid:plazi:treatment:03BC2A5DFF93FFE3ADD3AA76FB13FE22:8ECACD90FF93FFFFA857AAC2FA4DFD5A" box="[1299,1433,729,755]" country="Japan" county="Abukuma" municipality="Tsuruo" name="Saigo-mura" pageId="13" pageNumber="414" stateProvince="Fukushima">Saigo-mura</location>
,
<collectingRegion id="49D155A9FF93FFFFADD3AAE6FE80FCBE" box="[151,340,765,791]" country="Japan" name="Fukushima" pageId="13" pageNumber="414">Fukushima Pref.</collectingRegion>
,
<date id="FFABBD8BFF93FFFFAC24AAE6FE69FCBE" box="[352,445,765,791]" pageId="13" pageNumber="414" value="2017-01-08">
<collectingDate id="EFEF4463FF93FFFFAC24AAE6FE69FCBE" box="[352,445,765,791]" pageId="13" pageNumber="414" value="2017-01-08">8.I.2017</collectingDate>
</date>
<collectorName id="26E0FE9DFF93FFFFAC81AAE6FDE4FCBE" box="[453,560,765,791]" pageId="13" pageNumber="414">R. Okada</collectorName>
leg.
<collectorName id="26E0FE9DFF93FFFFAF2DAAE6FD63FCBE" box="[617,695,765,791]" pageId="13" pageNumber="414">Larvae</collectorName>
</materialsCitation>
were identified
<emphasis id="B9614759FF93FFFFAE34AAE5FB88FCBE" box="[880,1116,766,791]" italics="true" pageId="13" pageNumber="414">ex societate imaginis</emphasis>
. Both
<taxonomicName id="4C15E0C8FF93FFFFA9EDAAE6FA48FCBE" baseAuthorityName="Sharp" baseAuthorityYear="1884" box="[1193,1436,765,791]" class="Insecta" family="Dytiscidae" genus="Platambus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B9614759FF93FFFFA9EDAAE6FA48FCBE" box="[1193,1436,765,791]" italics="true" pageId="13" pageNumber="414">Platambus fimbriatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C15E0C8FF93FFFFAD82AB39FE89FC92" baseAuthorityName="Sharp" baseAuthorityYear="1873" box="[198,349,802,827]" class="Insecta" family="Dytiscidae" genus="Platambus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B9614759FF93FFFFAD82AB39FE89FC92" box="[198,349,802,827]" italics="true" pageId="13" pageNumber="414">P. pictipennis</emphasis>
</taxonomicName>
occur at these two localities. Larvae of
<taxonomicName id="4C15E0C8FF93FFFFAE5EAB39FC7CFC92" baseAuthorityName="Sharp" baseAuthorityYear="1884" box="[794,936,801,827]" class="Insecta" family="Dytiscidae" genus="Platambus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B9614759FF93FFFFAE5EAB39FC7CFC92" box="[794,936,801,827]" italics="true" pageId="13" pageNumber="414">P. fimbriatus</emphasis>
</taxonomicName>
, however, could readily be recognized using features listed below.
</paragraph>
</subSubSection>
<subSubSection id="C30FC8C0FF93FFFFAD83AB73FA49FA9A" pageId="13" pageNumber="414" type="diagnosis">
<paragraph id="8BAA9B4BFF93FFFFAD83AB73FF06FBD6" blockId="13.[151,1437,692,1979]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AB73FE96FC2A" bold="true" box="[199,322,872,899]" pageId="13" pageNumber="414">Diagnosis.</emphasis>
First instar larva of
<taxonomicName id="4C15E0C8FF93FFFFAF61AB71FD60FC2A" baseAuthorityName="Sharp" baseAuthorityYear="1884" box="[549,692,873,899]" class="Insecta" family="Dytiscidae" genus="Platambus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B9614759FF93FFFFAF61AB71FD60FC2A" box="[549,692,873,899]" italics="true" pageId="13" pageNumber="414">P. fimbriatus</emphasis>
</taxonomicName>
can be distinguished from the other species studied by the following combinations of features: frontoclypeus with less than 10 spatulate setae (
<figureCitation id="132E87CEFF93FFFFA94FAB96FBB4FC0E" box="[1035,1120,909,935]" captionStart="FIGURES 2930" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[315,1271,167,1910]" captionTargetId="figure@14.[315,1273,166,1910]" captionTargetPageId="14" captionText="FIGURES 2930. Platambus fimbriatus (Sharp, 1884), instar I, head capsule: (29) dorsal aspect (colour pattern not represented); (30) ventral aspect. EB, egg bursters; TP, tentorial pits. Numbers and lowercase letters refer to primary setae and pores, respectively. Scale bar = 0.10 mm." httpUri="https://zenodo.org/record/3351194/files/figure.png" pageId="13" pageNumber="414">Fig. 29</figureCitation>
); seta FR3 shorter than seta FR2 (
<figureCitation id="132E87CEFF93FFFFAD93ABAAFEFFFC62" box="[215,299,945,971]" captionStart="FIGURES 2930" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[315,1271,167,1910]" captionTargetId="figure@14.[315,1273,166,1910]" captionTargetPageId="14" captionText="FIGURES 2930. Platambus fimbriatus (Sharp, 1884), instar I, head capsule: (29) dorsal aspect (colour pattern not represented); (30) ventral aspect. EB, egg bursters; TP, tentorial pits. Numbers and lowercase letters refer to primary setae and pores, respectively. Scale bar = 0.10 mm." httpUri="https://zenodo.org/record/3351194/files/figure.png" pageId="13" pageNumber="414">Fig. 29</figureCitation>
); seta PA6 subequal in length to seta PA7 (
<figureCitation id="132E87CEFF93FFFFAE4EABAAFC8AFC62" box="[778,862,945,971]" captionStart="FIGURES 2930" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[315,1271,167,1910]" captionTargetId="figure@14.[315,1273,166,1910]" captionTargetPageId="14" captionText="FIGURES 2930. Platambus fimbriatus (Sharp, 1884), instar I, head capsule: (29) dorsal aspect (colour pattern not represented); (30) ventral aspect. EB, egg bursters; TP, tentorial pits. Numbers and lowercase letters refer to primary setae and pores, respectively. Scale bar = 0.10 mm." httpUri="https://zenodo.org/record/3351194/files/figure.png" pageId="13" pageNumber="414">Fig. 29</figureCitation>
); primary seta LA10 inserted subapically (
<figureCitation id="132E87CEFF93FFFFA87FABAAFA5BFC65" box="[1339,1423,945,972]" captionStart="FIGURES 3137" captionStartId="15.[151,264,1907,1932]" captionTargetBox="[152,1436,252,1820]" captionTargetId="figure@15.[151,1436,249,1820]" captionTargetPageId="15" captionText="FIGURES 3137. Platambus fimbriatus (Sharp, 1884), instar I, head appendages: (3132) antenna, (31) dorsal aspect; (32) ventral aspect; (3334) maxilla, (33) dorsal aspect; (34) ventral aspect; (3536) labium; (35) dorsal aspect; (36) ventral aspect; (37) mandible, dorsal aspect. Numbers and lowercase letters refer to primary setae and pores, respectively; filled squares refer to additional setae. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/3351196/files/figure.png" pageId="13" pageNumber="414">Fig. 36</figureCitation>
); setae TI2, TI3, TI6, TI7, TA6, and TA7 bluntly pointed apically; seta UR4 not inserted contiguously to setae UR2 and UR3; ratio L3/HW &lt;2.90. In addition to color patterns of head capsule (
<figureCitation id="132E87CEFF93FFFFAEB1ABE2FB9CFBBA" box="[1013,1096,1016,1043]" captionStart="FIGURES 5760" captionStartId="21.[151,264,1632,1657]" captionTargetBox="[151,1436,200,1603]" captionTargetId="figure@21.[151,1436,200,1612]" captionTargetPageId="21" captionText="FIGURES 5760. Head capsule of selected species of Platambus Thomson, 1859, instar III, dorsal aspect: (57) P. pictipennis (Sharp, 1873); (58) P. convexus Okada, 2011; (59) P. fimbriatus (Sharp, 1884); (60) P. sawadai (Kamiya, 1932). Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/3351206/files/figure.png" pageId="13" pageNumber="414">Fig. 59</figureCitation>
), last abdominal segment, and urogomphi (
<figureCitation id="132E87CEFF93FFFFAC60AC06FEAAFB9E" box="[292,382,1053,1079]" captionStart="FIGURES 6972" captionStartId="23.[151,264,1546,1571]" captionTargetBox="[178,1411,209,1471]" captionTargetId="figure@23.[151,1436,181,1520]" captionTargetPageId="23" captionText="FIGURES 6972. Last abdominal segment and urogomphi of selected species of Platambus Thomson, 1859, instar III, dorsal aspect: (69) P. pictipennis (Sharp, 1873); (70) P. convexus Okada, 2011; (71) P. fimbriatus (Sharp, 1884); (72) P. sawadai (Kamiya, 1932). Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/3351210/files/figure.png" pageId="13" pageNumber="414">Figs 71</figureCitation>
,
<figureCitation id="132E87CEFF93FFFFACC9AC06FE03FB9E" box="[397,471,1053,1079]" captionStart="FIGURES 7382" captionStartId="24.[151,264,1914,1939]" captionTargetBox="[151,1436,181,1876]" captionTargetId="figure@24.[151,1436,181,1876]" captionTargetPageId="24" captionText="FIGURES 7382. Dorsal habitus of selected species of Platambus Thomson, 1859: (73 75) instar I; (7678) instar II; (79-82) instar III; (73, 76, 79) P. pictipennis (Sharp, 1873); (74, 77, 80) P. fimbriatus (Sharp, 1884); (75, 78, 81) P. sawadai (Kamiya, 1932); (82) P. convexus Okada, 2011. Scale bar = 1.00 mm in (7378), 2.00 mm in (7982)." httpUri="https://zenodo.org/record/3351212/files/figure.png" pageId="13" pageNumber="414">7382</figureCitation>
), the total number of spatulate setae on frontoclypeus, secondary setae on pro- and mesofemur, and the presence/absence of ventral setae on protarsus (
<tableCitation id="C697AEF0FF93FFFFAED2AC5AFC38FBF2" box="[918,1004,1089,1115]" captionStart="TABLE 2" captionStartId="4.[151,239,154,178]" captionTargetPageId="4" captionText="TABLE 2. Number and position of secondary setae (including additional seta on femur) of selected species of Platambus Thomson, 1859: PIC = P. pictipennis (Sharp, 1873); CON = P. convexus Okada, 2011; FIM = P. fimbriatus (Sharp, 1884); SAW = P. sawadai (Kamiya, 1932); A = anterior, AD = anterodorsal, AV = anteroventral, CO = coxa, D = dorsal, Di = distal, FE = femur, PD = posterodorsal, Pr = proximal, PV = posteroventral, TA = tarsus, TI = tibia, TR = trochanter, V = ventral, n = number of specimens examined, Total = total number of secondary setae on segment." httpUri="http://table.plazi.org/id/DF6ACBC3FF9AFFF6ADD3A881FB72FEEB" pageId="13" pageNumber="414" tableUuid="DF6ACBC3FF9AFFF6ADD3A881FB72FEEB">Table 2</tableCitation>
) are helpful at discriminating later instars.
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AC92FA49FA9A" blockId="13.[151,1437,692,1979]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AC92FDB9FB0A" bold="true" box="[199,621,1160,1187]" pageId="13" pageNumber="414">
Instar I (
<figureCitation id="132E87CEFF93FFFFAC00AC93FE04FB0A" box="[324,464,1160,1187]" captionStart-0="FIGURES 2930" captionStart-1="FIGURES 3137" captionStart-2="FIGURES 3842" captionStartId-0="14.[151,264,1930,1955]" captionStartId-1="15.[151,264,1907,1932]" captionStartId-2="16.[151,264,1908,1933]" captionTargetBox-0="[315,1271,167,1910]" captionTargetBox-1="[152,1436,252,1820]" captionTargetBox-2="[258,1329,167,1886]" captionTargetId-0="figure@14.[315,1273,166,1910]" captionTargetId-1="figure@15.[151,1436,249,1820]" captionTargetId-2="figure@16.[257,1330,166,1888]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIGURES 2930. Platambus fimbriatus (Sharp, 1884), instar I, head capsule: (29) dorsal aspect (colour pattern not represented); (30) ventral aspect. EB, egg bursters; TP, tentorial pits. Numbers and lowercase letters refer to primary setae and pores, respectively. Scale bar = 0.10 mm." captionText-1="FIGURES 3137. Platambus fimbriatus (Sharp, 1884), instar I, head appendages: (3132) antenna, (31) dorsal aspect; (32) ventral aspect; (3334) maxilla, (33) dorsal aspect; (34) ventral aspect; (3536) labium; (35) dorsal aspect; (36) ventral aspect; (37) mandible, dorsal aspect. Numbers and lowercase letters refer to primary setae and pores, respectively; filled squares refer to additional setae. Scale bars = 0.10 mm." captionText-2="FIGURES 3842. Platambus fimbriatus (Sharp, 1884), instar I: (3839) metathoracic leg; (38) anterior aspect; (39) posterior aspect; (4041) last abdominal segment; (40) dorsal aspect; (41) ventral aspect; (42) urogomphi, dorsal aspect. Numbers and lowercase letters refer to primary setae and pores, respectively; filled squares refer to additional setae. Scale bars = 0.20 mm in (3839), 0.10 mm in (4042)." httpUri-0="https://zenodo.org/record/3351194/files/figure.png" httpUri-1="https://zenodo.org/record/3351196/files/figure.png" httpUri-2="https://zenodo.org/record/3351198/files/figure.png" pageId="13" pageNumber="414">Figs 2942</figureCitation>
,
<figureCitation id="132E87CEFF93FFFFACA2AC93FDD1FB0A" box="[486,517,1160,1187]" captionStart="FIGURES 7382" captionStartId="24.[151,264,1914,1939]" captionTargetBox="[151,1436,181,1876]" captionTargetId="figure@24.[151,1436,181,1876]" captionTargetPageId="24" captionText="FIGURES 7382. Dorsal habitus of selected species of Platambus Thomson, 1859: (73 75) instar I; (7678) instar II; (79-82) instar III; (73, 76, 79) P. pictipennis (Sharp, 1873); (74, 77, 80) P. fimbriatus (Sharp, 1884); (75, 78, 81) P. sawadai (Kamiya, 1932); (82) P. convexus Okada, 2011. Scale bar = 1.00 mm in (7378), 2.00 mm in (7982)." httpUri="https://zenodo.org/record/3351212/files/figure.png" pageId="13" pageNumber="414">74</figureCitation>
). Color.
</emphasis>
Dorsal surface of head capsule predominantly dark brown; frontoclypeus yellowish anteriorly, dark brown around each egg burster; parietale dark brown, pal- er over occipital region; head appendages pale yellow, A3, A4, MP3, MP2, MP3 and LP2 infuscate apically; thoracic and abdominal terga I to VII pale brown; LAS predominantly dark brown; urogomphi pale yellow basally, dark brown over distal half; legs predominantly pale yellow, dark brown apically.
</paragraph>
</subSubSection>
<subSubSection id="C30FC8C0FF93FFE3AD83AD26FDE3FEB6" lastPageId="17" lastPageNumber="418" pageId="13" pageNumber="414" type="description">
<paragraph id="8BAA9B4BFF93FFFFAD83AD26FB5BFAFE" blockId="13.[151,1437,692,1979]" box="[199,1167,1341,1367]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AD26FEDFFAFE" bold="true" box="[199,267,1341,1367]" pageId="13" pageNumber="414">Body.</emphasis>
Measurements and ratios that characterize the body shape are shown in
<tableCitation id="C697AEF0FF93FFFFA971AD26FB5DFAFE" box="[1077,1161,1341,1367]" captionStart="TABLE 3" captionStartId="5.[151,239,463,487]" captionTargetBox="[154,1429,644,1179]" captionText="TABLE 3. Number and position of additional setae of selected species of Platambus Thomson, 1859: PIC = P. pictipennis (Sharp, 1873); CON = P. convexus Okada, 2011; FIM = P. fimbriatus (Sharp, 1884); SAW = P. sawadai (Kamiya, 1932); AV = anteroventral, PV = posteroventral, Total = total number of additional and secondary setae on segment; n = number of specimens examined." httpUri="http://table.plazi.org/id/DF6ACBC3FF9BFFF7ADD3A9D4FE4DFDFD" pageId="13" pageNumber="414" tableUuid="DF6ACBC3FF9BFFF7ADD3A9D4FE4DFDFD">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AD7AFD6FFA36" blockId="13.[151,1437,692,1979]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AD7AFEDAFAD2" bold="true" box="[199,270,1377,1403]" pageId="13" pageNumber="414">Head.</emphasis>
Head capsule (
<figureCitation id="132E87CEFF93FFFFACFFAD7AFDEBFAD2" box="[443,575,1377,1403]" captionStart="FIGURES 2930" captionStartId="14.[151,264,1930,1955]" captionTargetBox="[315,1271,167,1910]" captionTargetId="figure@14.[315,1273,166,1910]" captionTargetPageId="14" captionText="FIGURES 2930. Platambus fimbriatus (Sharp, 1884), instar I, head capsule: (29) dorsal aspect (colour pattern not represented); (30) ventral aspect. EB, egg bursters; TP, tentorial pits. Numbers and lowercase letters refer to primary setae and pores, respectively. Scale bar = 0.10 mm." httpUri="https://zenodo.org/record/3351194/files/figure.png" pageId="13" pageNumber="414">Figs 2930</figureCitation>
). Rounded, sides slightly diverging posteriorly; frontoclypeus with 10 spatulate setae. Antenna (
<figureCitation id="132E87CEFF93FFFFAC3DAD9EFE2FFA36" box="[377,507,1413,1439]" captionStart="FIGURES 3137" captionStartId="15.[151,264,1907,1932]" captionTargetBox="[152,1436,252,1820]" captionTargetId="figure@15.[151,1436,249,1820]" captionTargetPageId="15" captionText="FIGURES 3137. Platambus fimbriatus (Sharp, 1884), instar I, head appendages: (3132) antenna, (31) dorsal aspect; (32) ventral aspect; (3334) maxilla, (33) dorsal aspect; (34) ventral aspect; (3536) labium; (35) dorsal aspect; (36) ventral aspect; (37) mandible, dorsal aspect. Numbers and lowercase letters refer to primary setae and pores, respectively; filled squares refer to additional setae. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/3351196/files/figure.png" pageId="13" pageNumber="414">Figs 3132</figureCitation>
). A3 finger-like.
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83ADB3FB59F986" blockId="13.[151,1437,692,1979]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83ADB3FE82FA6A" bold="true" box="[199,342,1448,1475]" pageId="13" pageNumber="414">Chaetotaxy.</emphasis>
Seta FR3 shorter than seta FR2; seta PA6 subequal or longer than seta PA7; seta PA13 shorter than PA12; prementum with one additional seta on dorsal surface; setae TI2, TI3, TI6, TI7, TA6, and TA7 bluntly pointed apically; seta AB2 shorter than seta AB9; seta AB15 longer than seta AB11; seta UR4 not inserted contiguously to setae UR2 and UR3; position and number of additional setae on legs are shown in
<tableCitation id="C697AEF0FF93FFFFA977AE0EFB53F986" box="[1075,1159,1557,1583]" captionStart="TABLE 1" captionStartId="3.[151,239,1666,1690]" captionTargetPageId="3" captionText="TABLE 1. Number and position of additional setae of selected species of Platambus Thomson, 1859: PIC = P. pictipennis (Sharp, 1873); CON = P. convexus Okada, 2011; FIM = P. fimbriatus (Sharp, 1884); SAW = P. sawadai (Kamiya, 1932); AV = anteroventral, PV = posteroventral, Total = total number of additional and secondary setae on segment; n = number of specimens examined." httpUri="http://table.plazi.org/id/DF6ACBC3FF9DFFF1ADD3AE99FE4DF8AE" pageId="13" pageNumber="414" tableUuid="DF6ACBC3FF9DFFF1ADD3AE99FE4DF8AE">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AE22FC85F9FA" blockId="13.[151,1437,692,1979]" box="[199,849,1592,1619]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AE22FE4AF9FA" bold="true" box="[199,414,1592,1619]" pageId="13" pageNumber="414">
Instar II (
<figureCitation id="132E87CEFF93FFFFAC78AE23FE45F9FA" box="[316,401,1592,1619]" captionStart="FIGURES 7382" captionStartId="24.[151,264,1914,1939]" captionTargetBox="[151,1436,181,1876]" captionTargetId="figure@24.[151,1436,181,1876]" captionTargetPageId="24" captionText="FIGURES 7382. Dorsal habitus of selected species of Platambus Thomson, 1859: (73 75) instar I; (7678) instar II; (79-82) instar III; (73, 76, 79) P. pictipennis (Sharp, 1873); (74, 77, 80) P. fimbriatus (Sharp, 1884); (75, 78, 81) P. sawadai (Kamiya, 1932); (82) P. convexus Okada, 2011. Scale bar = 1.00 mm in (7378), 2.00 mm in (7982)." httpUri="https://zenodo.org/record/3351212/files/figure.png" pageId="13" pageNumber="414">Fig. 77</figureCitation>
).
</emphasis>
As first-instar larva except as follows:
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AE47FBB1F9DE" blockId="13.[151,1437,692,1979]" box="[199,1125,1628,1655]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AE47FEC4F9DE" bold="true" box="[199,272,1628,1655]" pageId="13" pageNumber="414">Color.</emphasis>
Dorsal surface of LAS yellowish anteriorly, dark brown over posterior half.
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AE9AFB5BF932" blockId="13.[151,1437,692,1979]" box="[199,1167,1665,1691]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AE9AFEDFF932" bold="true" box="[199,267,1665,1691]" pageId="13" pageNumber="414">Body.</emphasis>
Measurements and ratios that characterize the body shape are shown in
<tableCitation id="C697AEF0FF93FFFFA971AE9AFB5DF932" box="[1077,1161,1665,1691]" captionStart="TABLE 3" captionStartId="5.[151,239,463,487]" captionTargetBox="[154,1429,644,1179]" captionText="TABLE 3. Number and position of additional setae of selected species of Platambus Thomson, 1859: PIC = P. pictipennis (Sharp, 1873); CON = P. convexus Okada, 2011; FIM = P. fimbriatus (Sharp, 1884); SAW = P. sawadai (Kamiya, 1932); AV = anteroventral, PV = posteroventral, Total = total number of additional and secondary setae on segment; n = number of specimens examined." httpUri="http://table.plazi.org/id/DF6ACBC3FF9BFFF7ADD3A9D4FE4DFDFD" pageId="13" pageNumber="414" tableUuid="DF6ACBC3FF9BFFF7ADD3A9D4FE4DFDFD">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AEBEFB7EF916" blockId="13.[151,1437,692,1979]" box="[199,1194,1701,1727]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AEBEFEDAF916" bold="true" box="[199,270,1701,1727]" pageId="13" pageNumber="414">Head.</emphasis>
Head capsule. Frontoclypeus with 2021 spatulate setae. Antenna. A3 bulge-like.
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AED3FA49F8AE" blockId="13.[151,1437,692,1979]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AED3FE82F94A" bold="true" box="[199,342,1736,1763]" pageId="13" pageNumber="414">Chaetotaxy.</emphasis>
Position and number of secondary setae on legs are shown in
<tableCitation id="C697AEF0FF93FFFFA950AED2FBBDF94A" box="[1044,1129,1737,1763]" captionStart="TABLE 1" captionStartId="3.[151,239,1666,1690]" captionTargetPageId="3" captionText="TABLE 1. Number and position of additional setae of selected species of Platambus Thomson, 1859: PIC = P. pictipennis (Sharp, 1873); CON = P. convexus Okada, 2011; FIM = P. fimbriatus (Sharp, 1884); SAW = P. sawadai (Kamiya, 1932); AV = anteroventral, PV = posteroventral, Total = total number of additional and secondary setae on segment; n = number of specimens examined." httpUri="http://table.plazi.org/id/DF6ACBC3FF9DFFF1ADD3AE99FE4DF8AE" pageId="13" pageNumber="414" tableUuid="DF6ACBC3FF9DFFF1ADD3AE99FE4DF8AE">Table 1</tableCitation>
; parietale with 4 spine-like secondary setae on lateral margin of each side; anterodorsal secondary setae on TI and TA bluntly pointed apically.
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AF0AFBFCF882" blockId="13.[151,1437,692,1979]" box="[199,1064,1808,1835]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AF0AFD80F882" bold="true" box="[199,596,1808,1835]" pageId="13" pageNumber="414">
Instar III (
<figureCitation id="132E87CEFF93FFFFAC02AF0BFE4AF882" box="[326,414,1808,1835]" captionStart="FIGURES 5760" captionStartId="21.[151,264,1632,1657]" captionTargetBox="[151,1436,200,1603]" captionTargetId="figure@21.[151,1436,200,1612]" captionTargetPageId="21" captionText="FIGURES 5760. Head capsule of selected species of Platambus Thomson, 1859, instar III, dorsal aspect: (57) P. pictipennis (Sharp, 1873); (58) P. convexus Okada, 2011; (59) P. fimbriatus (Sharp, 1884); (60) P. sawadai (Kamiya, 1932). Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/3351206/files/figure.png" pageId="13" pageNumber="414">Figs 59</figureCitation>
,
<figureCitation id="132E87CEFF93FFFFACEFAF0AFE26F882" box="[427,498,1809,1835]" captionStart="FIGURES 6168" captionStartId="22.[151,264,1903,1928]" captionTargetBox="[193,1395,167,1882]" captionTargetId="figure@22.[191,1395,166,1883]" captionTargetPageId="22" captionText="FIGURES 6168. Prothoracic leg of selected species of Platambus Thomson, 1859, instar III: (6162) P. pictipennis (Sharp, 1873), (61) anterior aspect; (62) posterior aspect; (6364) P. convexus Okada, 2011, (63) anterior aspect; (64) posterior aspect; (6566) P. fimbriatus (Sharp, 1884), (65) anterior aspect; (66) posterior aspect; (6768) P. sawadai (Kamiya, 1932), (67) anterior aspect; (68) posterior aspect. Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/3351208/files/figure.png" pageId="13" pageNumber="414">6566</figureCitation>
,
<figureCitation id="132E87CEFF93FFFFACBBAF0AFDC8F882" box="[511,540,1809,1835]" captionStart="FIGURES 6972" captionStartId="23.[151,264,1546,1571]" captionTargetBox="[178,1411,209,1471]" captionTargetId="figure@23.[151,1436,181,1520]" captionTargetPageId="23" captionText="FIGURES 6972. Last abdominal segment and urogomphi of selected species of Platambus Thomson, 1859, instar III, dorsal aspect: (69) P. pictipennis (Sharp, 1873); (70) P. convexus Okada, 2011; (71) P. fimbriatus (Sharp, 1884); (72) P. sawadai (Kamiya, 1932). Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/3351210/files/figure.png" pageId="13" pageNumber="414">71</figureCitation>
,
<figureCitation id="132E87CEFF93FFFFAF6DAF0BFD9CF882" box="[553,584,1808,1835]" captionStart="FIGURES 7382" captionStartId="24.[151,264,1914,1939]" captionTargetBox="[151,1436,181,1876]" captionTargetId="figure@24.[151,1436,181,1876]" captionTargetPageId="24" captionText="FIGURES 7382. Dorsal habitus of selected species of Platambus Thomson, 1859: (73 75) instar I; (7678) instar II; (79-82) instar III; (73, 76, 79) P. pictipennis (Sharp, 1873); (74, 77, 80) P. fimbriatus (Sharp, 1884); (75, 78, 81) P. sawadai (Kamiya, 1932); (82) P. convexus Okada, 2011. Scale bar = 1.00 mm in (7378), 2.00 mm in (7982)." httpUri="https://zenodo.org/record/3351212/files/figure.png" pageId="13" pageNumber="414">80</figureCitation>
).
</emphasis>
As second-instar larva except as follows:
</paragraph>
<paragraph id="8BAA9B4BFF93FFFFAD83AF2FFC47F812" blockId="13.[151,1437,692,1979]" pageId="13" pageNumber="414">
<emphasis id="B9614759FF93FFFFAD83AF2FFEC4F8E6" bold="true" box="[199,272,1844,1871]" pageId="13" pageNumber="414">Color.</emphasis>
Dorsal surface of head capsule yellowish; parietale yellowish with a narrow V-shaped dark brown marking across capsule at about mid-length and some brown maculae posteriorly; pro- and mesotergum and abdominal segments yellowish with several brown maculae, metanotum predominantly dark brown; LAS yellowish anteriorly, dark brown over distal 1/3 to 1/2; urogomphi predominantly yellow.
</paragraph>
<caption id="DF6ACBC3FF90FFFCADD3AF91FE22F843" httpUri="https://zenodo.org/record/3351194/files/figure.png" pageId="14" pageNumber="415" startId="14.[151,264,1930,1955]" targetBox="[315,1271,167,1910]" targetPageId="14">
<paragraph id="8BAA9B4BFF90FFFCADD3AF91FE22F843" blockId="14.[151,1437,1930,2027]" pageId="14" pageNumber="415">
<emphasis id="B9614759FF90FFFCADD3AF91FE83F80A" bold="true" box="[151,343,1930,1955]" pageId="14" pageNumber="415">FIGURES 2930.</emphasis>
<taxonomicName id="4C15E0C8FF90FFFCAC24AF90FD02F80A" authority="(Sharp, 1884)" authorityName="Sharp" authorityYear="1884" baseAuthorityName="Sharp" baseAuthorityYear="1884" box="[352,726,1930,1955]" class="Insecta" family="Dytiscidae" genus="Platambus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B9614759FF90FFFCAC24AF90FDE9F80A" box="[352,573,1931,1955]" italics="true" pageId="14" pageNumber="415">Platambus fimbriatus</emphasis>
(
<bibRefCitation id="EF84E6BAFF90FFFCAF09AF91FD1BF80A" author="Sharp, D." box="[589,719,1930,1955]" pageId="14" pageNumber="415" pagination="439 - 464" refId="ref16064" refString="Sharp, D. (1884) XXI. The water-beetles of Japan. Transactions of the Entomological Society of London, 1884, 439 - 464." type="journal article" year="1884">Sharp, 1884</bibRefCitation>
)
</taxonomicName>
, instar I, head capsule: (29) dorsal aspect (colour pattern not represented); (30) ventral aspect. EB, egg bursters; TP, tentorial pits. Numbers and lowercase letters refer to primary setae and pores, respectively. Scale bar = 0.10 mm.
</paragraph>
</caption>
<caption id="DF6ACBC3FF91FFFDADD3AF68FD94F85E" httpUri="https://zenodo.org/record/3351196/files/figure.png" pageId="15" pageNumber="416" startId="15.[151,264,1907,1932]" targetBox="[152,1436,252,1820]" targetPageId="15">
<paragraph id="8BAA9B4BFF91FFFDADD3AF68FD94F85E" blockId="15.[151,1436,1907,2040]" pageId="15" pageNumber="416">
<emphasis id="B9614759FF91FFFDADD3AF68FE82F825" bold="true" box="[151,342,1907,1932]" pageId="15" pageNumber="416">FIGURES 3137.</emphasis>
<taxonomicName id="4C15E0C8FF91FFFDAC1AAF68FD06F822" authority="(Sharp, 1884)" authorityName="Sharp" authorityYear="1884" baseAuthorityName="Sharp" baseAuthorityYear="1884" box="[350,722,1907,1932]" class="Insecta" family="Dytiscidae" genus="Platambus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B9614759FF91FFFDAC1AAF68FDEFF822" box="[350,571,1907,1931]" italics="true" pageId="15" pageNumber="416">Platambus fimbriatus</emphasis>
(
<bibRefCitation id="EF84E6BAFF91FFFDAF0EAF68FD18F822" author="Sharp, D." box="[586,716,1907,1932]" pageId="15" pageNumber="416" pagination="439 - 464" refId="ref16064" refString="Sharp, D. (1884) XXI. The water-beetles of Japan. Transactions of the Entomological Society of London, 1884, 439 - 464." type="journal article" year="1884">Sharp, 1884</bibRefCitation>
)
</taxonomicName>
, instar I, head appendages: (3132) antenna, (31) dorsal aspect; (32) ventral aspect; (3334) maxilla, (33) dorsal aspect; (34) ventral aspect; (3536) labium; (35) dorsal aspect; (36) ventral aspect; (37) mandible, dorsal aspect. Numbers and lowercase letters refer to primary setae and pores, respectively; filled squares refer to additional setae. Scale bars = 0.10 mm.
</paragraph>
</caption>
<caption id="DF6ACBC3FF8EFFE2ADD3AF6FFE12F850" httpUri="https://zenodo.org/record/3351198/files/figure.png" pageId="16" pageNumber="417" startId="16.[151,264,1908,1933]" targetBox="[258,1329,167,1886]" targetPageId="16">
<paragraph id="8BAA9B4BFF8EFFE2ADD3AF6FFE12F850" blockId="16.[151,1437,1908,2041]" pageId="16" pageNumber="417">
<emphasis id="B9614759FF8EFFE2ADD3AF6FFE81F824" bold="true" box="[151,341,1908,1933]" pageId="16" pageNumber="417">FIGURES 3842.</emphasis>
<taxonomicName id="4C15E0C8FF8EFFE2AC18AF6EFD18F824" authority="(Sharp, 1884)" authorityName="Sharp" authorityYear="1884" baseAuthorityName="Sharp" baseAuthorityYear="1884" box="[348,716,1908,1933]" class="Insecta" family="Dytiscidae" genus="Platambus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="fimbriatus">
<emphasis id="B9614759FF8EFFE2AC18AF6EFDE3F824" box="[348,567,1909,1933]" italics="true" pageId="16" pageNumber="417">Platambus fimbriatus</emphasis>
(
<bibRefCitation id="EF84E6BAFF8EFFE2AF02AF6FFD11F824" author="Sharp, D." box="[582,709,1908,1933]" pageId="16" pageNumber="417" pagination="439 - 464" refId="ref16064" refString="Sharp, D. (1884) XXI. The water-beetles of Japan. Transactions of the Entomological Society of London, 1884, 439 - 464." type="journal article" year="1884">Sharp, 1884</bibRefCitation>
)
</taxonomicName>
, instar I: (3839) metathoracic leg; (38) anterior aspect; (39) posterior aspect; (4041) last abdominal segment; (40) dorsal aspect; (41) ventral aspect; (42) urogomphi, dorsal aspect. Numbers and lowercase letters refer to primary setae and pores, respectively; filled squares refer to additional setae. Scale bars = 0.20 mm in (3839), 0.10 mm in (4042).
</paragraph>
</caption>
<paragraph id="8BAA9B4BFF8FFFE3AD83A882FB5BFF1A" blockId="17.[151,1436,153,395]" box="[199,1167,153,179]" pageId="17" pageNumber="418">
<emphasis id="B9614759FF8FFFE3AD83A882FEDFFF1A" bold="true" box="[199,267,153,179]" pageId="17" pageNumber="418">Body.</emphasis>
Measurements and ratios that characterize the body shape are shown in
<tableCitation id="C697AEF0FF8FFFE3A971A882FB5DFF1A" box="[1077,1161,153,179]" captionStart="TABLE 3" captionStartId="5.[151,239,463,487]" captionTargetBox="[154,1429,644,1179]" captionText="TABLE 3. Number and position of additional setae of selected species of Platambus Thomson, 1859: PIC = P. pictipennis (Sharp, 1873); CON = P. convexus Okada, 2011; FIM = P. fimbriatus (Sharp, 1884); SAW = P. sawadai (Kamiya, 1932); AV = anteroventral, PV = posteroventral, Total = total number of additional and secondary setae on segment; n = number of specimens examined." httpUri="http://table.plazi.org/id/DF6ACBC3FF9BFFF7ADD3A9D4FE4DFDFD" pageId="17" pageNumber="418" tableUuid="DF6ACBC3FF9BFFF7ADD3A9D4FE4DFDFD">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8BAA9B4BFF8FFFE3AD83A8A6FC2DFF7E" blockId="17.[151,1436,153,395]" box="[199,1017,188,215]" pageId="17" pageNumber="418">
<emphasis id="B9614759FF8FFFE3AD83A8A6FEDAFF7E" bold="true" box="[199,270,189,215]" pageId="17" pageNumber="418">Head.</emphasis>
Head capsule (
<figureCitation id="132E87CEFF8FFFE3ACFEA8A6FDDBFF7E" box="[442,527,188,215]" captionStart="FIGURES 5760" captionStartId="21.[151,264,1632,1657]" captionTargetBox="[151,1436,200,1603]" captionTargetId="figure@21.[151,1436,200,1612]" captionTargetPageId="21" captionText="FIGURES 5760. Head capsule of selected species of Platambus Thomson, 1859, instar III, dorsal aspect: (57) P. pictipennis (Sharp, 1873); (58) P. convexus Okada, 2011; (59) P. fimbriatus (Sharp, 1884); (60) P. sawadai (Kamiya, 1932). Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/3351206/files/figure.png" pageId="17" pageNumber="418">Fig. 59</figureCitation>
). Frontoclypeus with 2631 spatulate setae.
</paragraph>
<paragraph id="8BAA9B4BFF8FFFE3AD83A8FBFDE3FEB6" blockId="17.[151,1436,153,395]" pageId="17" pageNumber="418">
<emphasis id="B9614759FF8FFFE3AD83A8FBFE84FF52" bold="true" box="[199,336,224,251]" pageId="17" pageNumber="418">Chaetotaxy</emphasis>
. Position and number of secondary setae on legs are shown in
<tableCitation id="C697AEF0FF8FFFE3A945A8FAFB80FF52" box="[1025,1108,225,251]" captionStart="TABLE 1" captionStartId="3.[151,239,1666,1690]" captionTargetPageId="3" captionText="TABLE 1. Number and position of additional setae of selected species of Platambus Thomson, 1859: PIC = P. pictipennis (Sharp, 1873); CON = P. convexus Okada, 2011; FIM = P. fimbriatus (Sharp, 1884); SAW = P. sawadai (Kamiya, 1932); AV = anteroventral, PV = posteroventral, Total = total number of additional and secondary setae on segment; n = number of specimens examined." httpUri="http://table.plazi.org/id/DF6ACBC3FF9DFFF1ADD3AE99FE4DF8AE" pageId="17" pageNumber="418" tableUuid="DF6ACBC3FF9DFFF1ADD3AE99FE4DF8AE">Table 1</tableCitation>
; parietale with 68 spine-like secondary setae along lateral margin.
</paragraph>
</subSubSection>
<subSubSection id="C30FC8C0FF8FFFE3AD83A933FB13FE22" pageId="17" pageNumber="418" type="biology_ecology">
<paragraph id="8BAA9B4BFF8FFFE3AD83A933FB13FE22" blockId="17.[151,1436,153,395]" pageId="17" pageNumber="418">
<emphasis id="B9614759FF8FFFE3AD83A933FEFEFEEA" bold="true" box="[199,298,296,323]" pageId="17" pageNumber="418">Habitat.</emphasis>
Larvae occur along quiet margins of moderate sized rivers flowing on sandy bottom. First-instar larvae are mainly collected along the banks under stones in shallow water. Whereas also collected in shallow parts, both second- and third- instar larvae appear to be more abundant in dry grass deposits (
<figureCitation id="132E87CEFF8FFFE3A922A96AFB6DFE22" box="[1126,1209,368,395]" captionStart="FIGURES 8388" captionStartId="25.[151,264,1842,1867]" captionTargetBox="[151,1436,189,1808]" captionTargetId="figure@25.[151,1436,189,1808]" captionTargetPageId="25" captionText="FIGURES 8388. Collecting localities of the larvae of Platambus Thomson, 1859: (8385) Echi river, Shiga Prefecture, Western Japan, (83) moderate sized river, with withered grass hanging; (84) river bank with gravel bed; (85) shallow area with dry grass deposited; (86) Nishinopporo-sawa river, Hokkaido Prefecture, Northern Japan, small stream flowing under broadleaved trees; (8788) Happudani river, Shiga Prefecture, Western Japan, (87) mountain stream; (88) gutters with constantly flowing clear water." httpUri="https://zenodo.org/record/3351214/files/figure.png" pageId="17" pageNumber="418">Fig. 85</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>