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<document id="93C05DF76145C5E09BBA3230546A88A2" ID="10.11646/zootaxa.4200.4.7" ID-DOI="10.11646/zootaxa.4200.4.7" ID-GBIF-Dataset="c207b55d-3ec0-4787-8c81-74e315d14eb5" ID-ISSN="1175-5326" ID-Zenodo-Dep="191703" ID-ZooBank="1F2DA504-24BD-49C6-A7F3-A26B2132D6EE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480581841182" checkinUser="plazi" docAuthor="Stonis, Jonas R., Remeikis, Andrius, Diškus, Arūnas &amp; Gerulaitis, Virginijus" docDate="2016" docId="03A4145CFFAF0B6E7EE6FABB8EDCFEE8" docLanguage="en" docName="zootaxa.4200.4.7.pdf" docOrigin="Zootaxa 4200 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stigmella varispinella Diskus &amp; Stonis, sp. nov." docType="treatment" docVersion="7" lastPageNumber="569" masterDocId="FF9D6C24FFAB0B667E71FFC48D2DFFF5" masterDocTitle="The Ando-Patagonian Stigmella magnispinella group (Lepidoptera, Nepticulidae) with description of new species from Ecuador, Peru and Argentina" masterLastPageNumber="579" masterPageNumber="561" pageNumber="565" updateTime="1698395402645" updateUser="plazi">
<mods:mods id="5D75289BB0813B53CFD5C5B41A238434" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0A987EC3BC5570FA104310FD7BFE3AEF">The Ando-Patagonian Stigmella magnispinella group (Lepidoptera, Nepticulidae) with description of new species from Ecuador, Peru and Argentina</mods:title>
</mods:titleInfo>
<mods:name id="FCBBB171BDA1DF834344FF905E7DA5E6" type="personal">
<mods:role id="BB5FB7F17E31753356EA662865075DB5">
<mods:roleTerm id="DDFA2A8A11A09147FE6BBEE589EC7B7B">Author</mods:roleTerm>
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<mods:namePart id="CD6C30E5D2DCBB8051BB9D33F1CEF907">Stonis, Jonas R.</mods:namePart>
</mods:name>
<mods:name id="BB0B177C61E2A9630F54619A793F3C73" type="personal">
<mods:role id="A6EFEB7CCE0B183D35AC1E436BD0B0AC">
<mods:roleTerm id="168843B1B78EB4B86AD987C20DC42DB2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81B2F90739E2DEFC2250EDFFDCD2ADC8">Remeikis, Andrius</mods:namePart>
</mods:name>
<mods:name id="49BE75B5202038A4D1CB2141CF968EEF" type="personal">
<mods:role id="43326E293BAB9003CFBBD7BE66AA6A46">
<mods:roleTerm id="78304B38B498EA7CA6DCA03515835D3B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CECB493AD2C025F4C3BEAE6C64817AAE">Diškus, Arūnas</mods:namePart>
</mods:name>
<mods:name id="ABB1540E0D60E79B626BD6A030E4FC62" type="personal">
<mods:role id="7EFCF42A26AD54918B77DD3B9A81B762">
<mods:roleTerm id="660D353AB7E15F22C119D85D826D483D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B2BC4AEDD97C49ABB62EC115F9046638">Gerulaitis, Virginijus</mods:namePart>
</mods:name>
<mods:typeOfResource id="354ADCF882D55E520A337A4F9CEFCEDB">text</mods:typeOfResource>
<mods:relatedItem id="9BE14F69D33FC12F8E9583D2B8274579" type="host">
<mods:titleInfo id="177D10CFB643B08F0AF947DEDB073497">
<mods:title id="BBA3F3E33BCA7D5FCF4B7235D0E2994C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BB30DE9E8C330407A3B007B20CCF030F">
<mods:date id="BA2AA4FA97A27424CFBE05A818839C7A">2016</mods:date>
<mods:detail id="E95FB9BD7F19C3FB038928B3227663FE" type="volume">
<mods:number id="86367C1E3D1C428FE6601ABD83ABF59B">4200</mods:number>
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<mods:detail id="A6084D1142F059AFF060B04FF8D588FE" type="issue">
<mods:number id="E9D7438E96B9B776FD280D7B55B611AD">4</mods:number>
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<mods:start id="013BB4D9E7A986399572B462419E8500">561</mods:start>
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<mods:classification id="40919AE5879272ABB82C733CC149D398">journal article</mods:classification>
<mods:identifier id="87CCD35B29D3CD76152229E57123C5D8" type="DOI">10.11646/zootaxa.4200.4.7</mods:identifier>
<mods:identifier id="F600C076CA79871E8F59DEB0285541E4" type="GBIF-Dataset">c207b55d-3ec0-4787-8c81-74e315d14eb5</mods:identifier>
<mods:identifier id="EA639254020517539F0F52D8F131DD47" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="74ACD43E09F12590A5E7A99837538EDF" type="Zenodo-Dep">191703</mods:identifier>
<mods:identifier id="AA9AC843A00FB6BAB80F894D56A01A25" type="ZooBank">1F2DA504-24BD-49C6-A7F3-A26B2132D6EE</mods:identifier>
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<treatment id="03A4145CFFAF0B6E7EE6FABB8EDCFEE8" ID-DOI="http://doi.org/10.5281/zenodo.6079526" ID-GBIF-Taxon="124936449" ID-Zenodo-Dep="6079526" LSID="urn:lsid:plazi:treatment:03A4145CFFAF0B6E7EE6FABB8EDCFEE8" httpUri="http://treatment.plazi.org/id/03A4145CFFAF0B6E7EE6FABB8EDCFEE8" lastPageId="8" lastPageNumber="569" pageId="4" pageNumber="565">
<subSubSection id="C317F6C1FFAF0B627EE6FABB8C73FA4E" pageId="4" pageNumber="565" type="nomenclature">
<paragraph id="8BB2A54AFFAF0B627EE6FABB8FFAFA6F" blockId="4.[151,727,1407,1467]" box="[151,727,1407,1434]" pageId="4" pageNumber="565">
<heading id="D0FA1226FFAF0B627EE6FABB8FFAFA6F" bold="true" box="[151,727,1407,1434]" fontSize="11" level="1" pageId="4" pageNumber="565" reason="1">
<emphasis id="B9797958FFAF0B627EE6FABB8FFAFA6F" bold="true" box="[151,727,1407,1434]" pageId="4" pageNumber="565">
<taxonomicName id="4C0DDEC9FFAF0B627EE6FABB8F41FA6F" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" box="[151,620,1407,1434]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="species" species="varispinella" status="sp. nov.">
<emphasis id="B9797958FFAF0B627EE6FABB8CB0FA6C" bold="true" box="[151,413,1407,1433]" italics="true" pageId="4" pageNumber="565">Stigmella varispinella</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<taxonomicNameLabel id="A24AC423FFAF0B627C08FA448FFAFA6F" box="[633,727,1408,1434]" pageId="4" pageNumber="565" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB2A54AFFAF0B627EE6FA668C73FA4E" blockId="4.[151,727,1407,1467]" box="[151,350,1442,1467]" pageId="4" pageNumber="565">
(
<figureCitation id="1336B9CFFFAF0B627EEEFA668DCAFA4E" box="[159,231,1442,1467]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[156,1400,193,1832]" captionTargetId="figure@2.[156,1430,193,1913]" captionTargetPageId="2" captionText="FIGURE 2. Currently known distribution of the species of the Stigmella magnispinella group." httpUri="https://zenodo.org/record/191705/files/figure.png" pageId="4" pageNumber="565">Figs 2</figureCitation>
,
<figureCitation id="1336B9CFFFAF0B627E82FA678C03FA4E" box="[243,302,1443,1467]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURES 6 – 12" captionStartId-0="5.[151,250,1904,1926]" captionStartId-1="6.[151,250,1765,1787]" captionStartId-2="7.[151,264,1753,1776]" captionTargetBox-0="[216,1362,338,1812]" captionTargetBox-1="[211,1367,232,1688]" captionTargetBox-2="[226,1354,199,1725]" captionTargetId-0="figure@5.[207,1379,301,1883]" captionTargetId-1="figure@6.[207,1379,193,1770]" captionTargetId-2="figure@7.[224,1358,197,1729]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 4. Diagnostics of the species of the Stigmella magnispinella group (male genitalia: phallus)." captionText-1="FIGURE 5. Diagnostics of species of the Stigmella magnispinella group (male genitalia: capsule)." captionText-2="FIGURES 6 – 12. Male genitalia of Stigmella varispinella Diškus &amp; Stonis, sp. nov., holotype, genitalia slide no. AD 810. 6 – 8, capsule with phallus removed; 9, uncus; 10, basal connection of valvae; 11, 12, phallus and cornuti (ZMUC)." httpUri-0="https://zenodo.org/record/191707/files/figure.png" httpUri-1="https://zenodo.org/record/191708/files/figure.png" httpUri-2="https://zenodo.org/record/191709/files/figure.png" pageId="4" pageNumber="565">4–12</figureCitation>
,
<figureCitation id="1336B9CFFFAF0B627F48FA678C7BFA4E" box="[313,342,1443,1467]" captionStart="FIGURES 36 – 41" captionStartId="16.[151,264,1810,1833]" captionTargetBox="[167,1448,193,1754]" captionTargetId="figure@16.[167,1452,193,1790]" captionTargetPageId="16" captionText="FIGURES 36 – 41. Male adults of the species of the Stigmella magnispinella group. 36, S. olekarsholti Remeikis, Diškus &amp; Stonis, sp. nov., holotype; 37, S. varispinella Diškus &amp; Stonis, sp. nov., holotype; 38, 39, S. magnispinella Remeikis &amp; Stonis, sp. nov., holotype; 40, S. dolia Remeikis &amp; Stonis, sp. nov., holotype; 41, S. patagonica Remeikis &amp; Stonis, sp. nov., paratype (ZMUC)." httpUri="https://zenodo.org/record/191715/files/figure.png" pageId="4" pageNumber="565">37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C317F6C1FFAF0B627EE6FA2E8FE6F9BE" pageId="4" pageNumber="565" type="materials_examined">
<paragraph id="8BB2A54AFFAF0B627EE6FA2E8FE6F9BE" blockId="4.[151,1437,1514,2007]" pageId="4" pageNumber="565">
<materialsCitation id="3B65AF17FFAF0B627EE6FA2E8FEAF9BE" ID-GBIF-Occurrence="1413076059" collectingDate="2005-01-26" collectionCode="ZMUC" collectorName="A. Diskus &amp; J. R. Stonis" country="Ecuador" elevation="3025" latitude="-1.8780556" location="Pallatanga" longLatPrecision="21" longitude="-78.90305" pageId="4" pageNumber="565" specimenCount="811" specimenCount-male="811" stateProvince="Chimborazo Province" typeStatus="holotype">
<emphasis id="B9797958FFAF0B627EE6FA2E8C60F9F6" bold="true" box="[151,333,1514,1539]" pageId="4" pageNumber="565">
<typeStatus id="54B61BE8FFAF0B627EE6FA2E8DFFF9F6" box="[151,210,1514,1539]" pageId="4" pageNumber="565">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B61BE8FFAF0B627F2DFA2E8CE4F9F6" box="[348,457,1514,1539]" pageId="4" pageNumber="565" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D0B6EC3FFAF0B627FADFA2F8CDEF9F7" box="[476,499,1515,1538]" pageId="4" pageNumber="565" type="male">♂</specimenCount>
,
<collectingCountry id="F31AE5DAFFAF0B627C79FA2F8FBFF9F6" box="[520,658,1515,1539]" name="Ecuador" pageId="4" pageNumber="565">ECUADOR</collectingCountry>
,
<collectingRegion id="49C96BA8FFAF0B627CD7FA2E8E8AF9F6" box="[678,935,1514,1539]" country="Ecuador" name="Chimborazo" pageId="4" pageNumber="565">Chimborazo Province</collectingRegion>
, about
<quantity id="4CF508AFFFAF0B627A79FA2F897AF9F6" box="[1032,1111,1514,1539]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="565" unit="km" value="30.0">30 km</quantity>
NE
<location id="8ED2F391FFAF0B627AEAFA2E883EF9F6" LSID="urn:lsid:plazi:treatment:03A4145CFFAF0B6E7EE6FABB8EDCFEE8:8ED2F391FFAF0B627AEAFA2E883EF9F6" box="[1179,1299,1514,1539]" country="Ecuador" latitude="-1.8780556" longLatPrecision="21" longitude="-78.90305" name="Pallatanga" pageId="4" pageNumber="565" stateProvince="Chimborazo Province">Pallatanga</location>
,
<geoCoordinate id="EE39C38DFFAF0B627B57FA2E88BAF9F6" box="[1318,1431,1514,1539]" direction="south" orientation="latitude" pageId="4" pageNumber="565" precision="15" value="-1.8780556">1°52'41&quot;S</geoCoordinate>
,
<geoCoordinate id="EE39C38DFFAF0B627EE6F9CA8C30F9D2" box="[151,285,1550,1575]" direction="west" orientation="longitude" pageId="4" pageNumber="565" precision="15" value="-78.90305">78°54'11&quot;W</geoCoordinate>
,
<elevation id="00204279FFAF0B627F58F9CA8CC2F9D3" box="[297,495,1550,1575]" metricMagnitude="3" metricUnit="m" metricValue="3.025" pageId="4" pageNumber="565" unit="m" value="3025.0">
elevation
<quantity id="4CF508AFFFAF0B627FE6F9CA8CC2F9D3" box="[407,495,1550,1575]" metricMagnitude="3" metricUnit="m" metricValue="3.025" pageId="4" pageNumber="565" unit="m" value="3025.0">3025 m</quantity>
</elevation>
, leaf-mining larva
<collectedFrom id="3811E6B7FFAF0B627CB4F9CB8E4EF9D2" box="[709,867,1551,1575]" pageId="4" pageNumber="565">
on
<taxonomicName id="4C0DDEC9FFAF0B627C99F9CB8E4EF9D2" box="[744,867,1551,1575]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="565" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
</collectedFrom>
plant
<collectingDate id="EFF77A62FFAF0B627DDBF9CA893AF9D2" box="[938,1047,1550,1575]" pageId="4" pageNumber="565" value="2005-01-26">
<date id="FFB3838AFFAF0B627DDBF9CA893AF9D2" box="[938,1047,1550,1575]" pageId="4" pageNumber="565" value="2005-01-26">26.i.2005</date>
</collectingDate>
), field card no. 4832,
<collectorName id="26F8C09CFFAF0B627B7FF9CB8852F9D2" box="[1294,1407,1550,1575]" pageId="4" pageNumber="565">A. Diškus</collectorName>
&amp;
<collectorName id="26F8C09CFFAF0B627EE6F9F78C35F9BE" box="[151,280,1586,1611]" pageId="4" pageNumber="565">J. R. Stonis</collectorName>
, genitalia slide no. AD
<specimenCount id="9D0B6EC3FFAF0B627C6AF9F78F74F9BE" box="[539,601,1587,1611]" pageId="4" pageNumber="565" type="male">810♂</specimenCount>
(
<collectionCode id="ED1C3D8FFFAF0B627C19F9F78F92F9BE" box="[616,703,1587,1611]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="565">ZMUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C317F6C1FFAF0B627EB6F9928972F943" pageId="4" pageNumber="565" type="diagnosis">
<paragraph id="8BB2A54AFFAF0B627EB6F9928972F943" blockId="4.[151,1437,1514,2007]" pageId="4" pageNumber="565">
<emphasis id="B9797958FFAF0B627EB6F9928C6CF99A" bold="true" box="[199,321,1622,1647]" pageId="4" pageNumber="565">Diagnosis.</emphasis>
The combination of a long, lobate vinculum, distinctive basal connection between valvae, and the blue-purple iridescence of the forewing distinguishes
<taxonomicName id="4C0DDEC9FFAF0B627C85F9BF8EBBF966" box="[756,918,1659,1683]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="species" species="varispinella" status="sp. nov.">
<emphasis id="B9797958FFAF0B627C85F9BF8EBBF966" box="[756,918,1659,1683]" italics="true" pageId="4" pageNumber="565">S. varispinella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24AC423FFAF0B627DECF9BE8EDAF966" box="[925,1015,1658,1683]" pageId="4" pageNumber="565" rank="species">
<emphasis id="B9797958FFAF0B627DECF9BE8EDAF966" bold="true" box="[925,1015,1658,1683]" pageId="4" pageNumber="565">sp. nov.</emphasis>
</taxonomicNameLabel>
from the most similar
<taxonomicName id="4C0DDEC9FFAF0B627A8BF9BF88B1F966" box="[1274,1436,1659,1683]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="species" species="olekarsholti" status="sp. nov.">
<emphasis id="B9797958FFAF0B627A8BF9BF88B1F966" box="[1274,1436,1659,1683]" italics="true" pageId="4" pageNumber="565">S. olekarsholti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24AC423FFAF0B627EE6F95A8DDDF942" box="[151,240,1694,1719]" pageId="4" pageNumber="565" rank="species">
<emphasis id="B9797958FFAF0B627EE6F95A8DDDF942" bold="true" box="[151,240,1694,1719]" pageId="4" pageNumber="565">sp. nov.</emphasis>
</taxonomicNameLabel>
(see also the pictorial key to the
<taxonomicName id="4C0DDEC9FFAF0B627C13F95B8E30F943" box="[610,797,1694,1718]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="565" phylum="Arthropoda" rank="species" species="magnispinella">
<emphasis id="B9797958FFAF0B627C13F95B8E30F943" box="[610,797,1694,1718]" italics="true" pageId="4" pageNumber="565">S. magnispinella</emphasis>
</taxonomicName>
group provided in figs 4, 5).
</paragraph>
</subSubSection>
<subSubSection id="C317F6C1FFAF0B617EB6F9068FB3F811" lastPageId="7" lastPageNumber="568" pageId="4" pageNumber="565" type="description">
<paragraph id="8BB2A54AFFAF0B637EB6F9068E35FF20" blockId="4.[151,1437,1514,2007]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" lastPageNumber="566" pageId="4" pageNumber="565">
<emphasis id="B9797958FFAF0B627EB6F9068C29F92E" bold="true" box="[199,260,1730,1755]" pageId="4" pageNumber="565">Male</emphasis>
(
<figureCitation id="1336B9CFFFAF0B627F62F9068C45F92E" box="[275,360,1730,1755]" captionStart="FIGURES 36 – 41" captionStartId="16.[151,264,1810,1833]" captionTargetBox="[167,1448,193,1754]" captionTargetId="figure@16.[167,1452,193,1790]" captionTargetPageId="16" captionText="FIGURES 36 – 41. Male adults of the species of the Stigmella magnispinella group. 36, S. olekarsholti Remeikis, Diškus &amp; Stonis, sp. nov., holotype; 37, S. varispinella Diškus &amp; Stonis, sp. nov., holotype; 38, 39, S. magnispinella Remeikis &amp; Stonis, sp. nov., holotype; 40, S. dolia Remeikis &amp; Stonis, sp. nov., holotype; 41, S. patagonica Remeikis &amp; Stonis, sp. nov., paratype (ZMUC)." httpUri="https://zenodo.org/record/191715/files/figure.png" pageId="4" pageNumber="565">Fig. 37</figureCitation>
). Forewing length about 2.0 mm; wingspan about 4.5 mm. Head: palpi glossy, brownish cream; frontal tuft pale ferruginous to ochreous cream (at certain angle of view concolorous with scape and collar); collar and scape ochreous cream; antenna distinctly longer than half the length of forewing; flagellum with 33–35 segments, dark grey-brown with golden gloss. Thorax and tegula very dark grey-brown with golden gloss. Forewing dark grey-brown with golden gloss and with two very slender, ill-defined, silvery shiny fasciae: postmedian and apical; most scales of forewing apex with strong blue and light purple iridescence while scales in forewing basal 2/3 with weak blue and purple iridescence; fringe grey-brown with some golden gloss; underside of forewing brown-black, without spots or androconia. Hindwing grey to dark grey on upper side and underside, without androconia or spots; its fringe pale grey to dark grey. Legs glossy, greyish cream; hindlegs with dark grey scales on upper side. Abdomen black-grey on upper side.
</paragraph>
<paragraph id="8BB2A54AFFAE0B637EB6FF1B8C8FFF0D" blockId="5.[151,1436,151,248]" box="[199,418,223,248]" pageId="5" pageNumber="566">
<emphasis id="B9797958FFAE0B637EB6FF1B8C33FF0D" bold="true" box="[199,286,223,248]" pageId="5" pageNumber="566">Female</emphasis>
. Unknown.
</paragraph>
<caption id="DF72F5C2FFAE0B637EE6F8B4898CF873" box="[151,1185,1904,1926]" httpUri="https://zenodo.org/record/191707/files/figure.png" pageId="5" pageNumber="566" targetBox="[216,1362,338,1812]" targetPageId="5">
<paragraph id="8BB2A54AFFAE0B637EE6F8B4898CF873" blockId="5.[151,1185,1904,1926]" box="[151,1185,1904,1926]" pageId="5" pageNumber="566">
<emphasis id="B9797958FFAE0B637EE6F8B48C3FF870" bold="true" box="[151,274,1904,1926]" pageId="5" pageNumber="566">FIGURE 4.</emphasis>
Diagnostics of the species of the
<taxonomicName id="4C0DDEC9FFAE0B637C16F8B48E71F873" box="[615,860,1904,1926]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="magnispinella">
<emphasis id="B9797958FFAE0B637C16F8B48E71F873" box="[615,860,1904,1926]" italics="true" pageId="5" pageNumber="566">Stigmella magnispinella</emphasis>
</taxonomicName>
group (male genitalia: phallus).
</paragraph>
</caption>
<caption id="DF72F5C2FFAD0B607EE6F9218952F90E" box="[151,1151,1765,1787]" httpUri="https://zenodo.org/record/191708/files/figure.png" pageId="6" pageNumber="567" targetBox="[211,1367,232,1688]" targetPageId="6">
<paragraph id="8BB2A54AFFAD0B607EE6F9218952F90E" blockId="6.[151,1151,1765,1787]" box="[151,1151,1765,1787]" pageId="6" pageNumber="567">
<emphasis id="B9797958FFAD0B607EE6F9218C3FF90E" bold="true" box="[151,274,1765,1787]" pageId="6" pageNumber="567">FIGURE 5.</emphasis>
Diagnostics of species of the
<taxonomicName id="4C0DDEC9FFAD0B607C33F9218E1BF90E" box="[578,822,1765,1787]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="species" species="magnispinella">
<emphasis id="B9797958FFAD0B607C33F9218E1BF90E" box="[578,822,1765,1787]" italics="true" pageId="6" pageNumber="567">Stigmella magnispinella</emphasis>
</taxonomicName>
group (male genitalia: capsule).
</paragraph>
</caption>
<paragraph id="8BB2A54AFFAD0B607EB6F8E78F3EF81A" blockId="6.[151,1436,1827,2031]" pageId="6" pageNumber="567">
<emphasis id="B9797958FFAD0B607EB6F8E78C5BF8C9" bold="true" box="[199,374,1827,1852]" pageId="6" pageNumber="567">Male genitalia</emphasis>
(
<figureCitation id="1336B9CFFFAD0B607FF8F8E78F2EF8CE" box="[393,515,1827,1852]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURES 6 – 12" captionStartId-0="5.[151,250,1904,1926]" captionStartId-1="6.[151,250,1765,1787]" captionStartId-2="7.[151,264,1753,1776]" captionTargetBox-0="[216,1362,338,1812]" captionTargetBox-1="[211,1367,232,1688]" captionTargetBox-2="[226,1354,199,1725]" captionTargetId-0="figure@5.[207,1379,301,1883]" captionTargetId-1="figure@6.[207,1379,193,1770]" captionTargetId-2="figure@7.[224,1358,197,1729]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 4. Diagnostics of the species of the Stigmella magnispinella group (male genitalia: phallus)." captionText-1="FIGURE 5. Diagnostics of species of the Stigmella magnispinella group (male genitalia: capsule)." captionText-2="FIGURES 6 – 12. Male genitalia of Stigmella varispinella Diškus &amp; Stonis, sp. nov., holotype, genitalia slide no. AD 810. 6 – 8, capsule with phallus removed; 9, uncus; 10, basal connection of valvae; 11, 12, phallus and cornuti (ZMUC)." httpUri-0="https://zenodo.org/record/191707/files/figure.png" httpUri-1="https://zenodo.org/record/191708/files/figure.png" httpUri-2="https://zenodo.org/record/191709/files/figure.png" pageId="6" pageNumber="567">Figs 4–12</figureCitation>
). Capsule longer (285–290 µm) than wide (140 µm). Uncus with two distally tapered lateral lobes (
<figureCitation id="1336B9CFFFAD0B607FF9F8828CE1F8AA" box="[392,460,1862,1887]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1753,1776]" captionTargetBox="[226,1354,199,1725]" captionTargetId="figure@7.[224,1358,197,1729]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Male genitalia of Stigmella varispinella Diškus &amp; Stonis, sp. nov., holotype, genitalia slide no. AD 810. 6 – 8, capsule with phallus removed; 9, uncus; 10, basal connection of valvae; 11, 12, phallus and cornuti (ZMUC)." httpUri="https://zenodo.org/record/191709/files/figure.png" pageId="6" pageNumber="567">Fig. 9</figureCitation>
). Gnathos with two stout caudal processes. Valva (
<figureCitation id="1336B9CFFFAD0B607A77F8828947F8AA" box="[1030,1130,1862,1887]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1753,1776]" captionTargetBox="[226,1354,199,1725]" captionTargetId="figure@7.[224,1358,197,1729]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Male genitalia of Stigmella varispinella Diškus &amp; Stonis, sp. nov., holotype, genitalia slide no. AD 810. 6 – 8, capsule with phallus removed; 9, uncus; 10, basal connection of valvae; 11, 12, phallus and cornuti (ZMUC)." httpUri="https://zenodo.org/record/191709/files/figure.png" pageId="6" pageNumber="567">Figs 6–8</figureCitation>
) 150 µm long, with bulged inner lobe and one slender apical processes; transtilla with large sublateral processes (
<figureCitation id="1336B9CFFFAD0B607A2EF8AF89E8F876" box="[1119,1221,1899,1924]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1753,1776]" captionTargetBox="[226,1354,199,1725]" captionTargetId="figure@7.[224,1358,197,1729]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Male genitalia of Stigmella varispinella Diškus &amp; Stonis, sp. nov., holotype, genitalia slide no. AD 810. 6 – 8, capsule with phallus removed; 9, uncus; 10, basal connection of valvae; 11, 12, phallus and cornuti (ZMUC)." httpUri="https://zenodo.org/record/191709/files/figure.png" pageId="6" pageNumber="567">Figs 6, 7</figureCitation>
). Juxta indistintive or absent. Ventral plate of vinculum very long, with large lateral lobes. Phallus (
<figureCitation id="1336B9CFFFAD0B607A60F84A89A2F852" box="[1041,1167,1934,1959]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1753,1776]" captionTargetBox="[226,1354,199,1725]" captionTargetId="figure@7.[224,1358,197,1729]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Male genitalia of Stigmella varispinella Diškus &amp; Stonis, sp. nov., holotype, genitalia slide no. AD 810. 6 – 8, capsule with phallus removed; 9, uncus; 10, basal connection of valvae; 11, 12, phallus and cornuti (ZMUC)." httpUri="https://zenodo.org/record/191709/files/figure.png" pageId="6" pageNumber="567">Figs 11, 12</figureCitation>
) 175–180 µm long, 45– 50 µm wide; vesica with a specific set of cornuti comprised of two groups of small spine-like cornuti and one very large horn-like cornutus (
<figureCitation id="1336B9CFFFAD0B607FC2F8128F2BF81A" box="[435,518,2006,2031]" captionStart="FIGURES 6 – 12" captionStartId="7.[151,264,1753,1776]" captionTargetBox="[226,1354,199,1725]" captionTargetId="figure@7.[224,1358,197,1729]" captionTargetPageId="7" captionText="FIGURES 6 – 12. Male genitalia of Stigmella varispinella Diškus &amp; Stonis, sp. nov., holotype, genitalia slide no. AD 810. 6 – 8, capsule with phallus removed; 9, uncus; 10, basal connection of valvae; 11, 12, phallus and cornuti (ZMUC)." httpUri="https://zenodo.org/record/191709/files/figure.png" pageId="6" pageNumber="567">Fig. 12</figureCitation>
).
</paragraph>
<caption id="DF72F5C2FFAC0B617EE6F91D89C5F8FB" httpUri="https://zenodo.org/record/191709/files/figure.png" pageId="7" pageNumber="568" targetBox="[226,1354,199,1725]" targetPageId="7">
<paragraph id="8BB2A54AFFAC0B617EE6F91D89C5F8FB" blockId="7.[151,1436,1753,1806]" pageId="7" pageNumber="568">
<emphasis id="B9797958FFAC0B617EE6F91D8C6AF91A" bold="true" box="[151,327,1753,1776]" pageId="7" pageNumber="568">FIGURES 6–12.</emphasis>
Male genitalia of
<taxonomicName id="4C0DDEC9FFAC0B617C73F91E8EA2F905" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" box="[514,911,1754,1776]" class="Insecta" family="Nepticulidae" genus="Stigmella" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="568" phylum="Arthropoda" rank="species" species="varispinella" status="sp. nov.">
<emphasis id="B9797958FFAC0B617C73F91E8FCDF905" box="[514,736,1754,1776]" italics="true" pageId="7" pageNumber="568">Stigmella varispinella</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<taxonomicNameLabel id="A24AC423FFAC0B617DEBF91E8EC7F91A" box="[922,1002,1754,1775]" pageId="7" pageNumber="568" rank="species">
<emphasis id="B9797958FFAC0B617DEBF91E8EC7F91A" bold="true" box="[922,1002,1754,1775]" pageId="7" pageNumber="568">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, genitalia slide no. AD810. 6–8, capsule with phallus removed; 9, uncus; 10, basal connection of valvae; 11, 12, phallus and cornuti (ZMUC).
</paragraph>
</caption>
<paragraph id="8BB2A54AFFAC0B617EB6F8FF8FB3F811" blockId="7.[151,1436,1851,2020]" pageId="7" pageNumber="568">
<emphasis id="B9797958FFAC0B617EB6F8FF8C6EF8A1" bold="true" box="[199,323,1851,1876]" pageId="7" pageNumber="568">Bionomics</emphasis>
. Larvae mine leaves of in January (the host, an
<taxonomicName id="4C0DDEC9FFAC0B617D27F8F88EFCF8A1" box="[854,977,1852,1876]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="568" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
plant, has not been properly documented and identified and remains unknown. However, we consider that the host-plant may belong to the genus
<taxonomicName id="4C0DDEC9FFAC0B617B4DF8A48DF8F869" authority="Cass." authorityName="Cass." class="Magnoliopsida" family="Asteraceae" genus="Gynoxys" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="568" phylum="Tracheophyta" rank="genus">
<emphasis id="B9797958FFAC0B617B4DF8A488B1F882" box="[1340,1436,1888,1911]" italics="true" pageId="7" pageNumber="568">Gynoxys</emphasis>
Cass.
</taxonomicName>
,
<taxonomicName id="4C0DDEC9FFAC0B617E92F8408C4EF869" box="[227,355,1924,1948]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="568" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
). Leaf-mine is combined: in the beginning, the gallery is very slender; further it widened to a large elongate blotch (the leaf-mine has not been properly recorded).
<typeStatus id="54B61BE8FFAC0B617DD0F8638924F835" box="[929,1033,1959,1984]" pageId="7" pageNumber="568" type="holotype">Holotype</typeStatus>
adult emerged in February. Cocoon yellowish white. Otherwise biology unknown.
</paragraph>
</subSubSection>
<subSubSection id="C317F6C1FFA30B6E7EB6FF538DDFFF21" pageId="8" pageNumber="569" type="distribution">
<paragraph id="8BB2A54AFFA30B6E7EB6FF538DDFFF21" blockId="8.[151,1436,151,285]" pageId="8" pageNumber="569">
<emphasis id="B9797958FFA30B6E7EB6FF538C74FF45" bold="true" box="[199,345,151,176]" pageId="8" pageNumber="569">Distribution</emphasis>
. Known from single locality in the Andes (
<collectingCountry id="F31AE5DAFFA30B6E7D35FF538E8AFF45" box="[836,935,151,176]" name="Ecuador" pageId="8" pageNumber="569">Ecuador</collectingCountry>
:
<collectingRegion id="49C96BA8FFA30B6E7DC2FF5389A3FF45" box="[947,1166,151,176]" country="Ecuador" name="Pichincha" pageId="8" pageNumber="569">Pichincha Province</collectingRegion>
) at elevation of
<quantity id="4CF508AFFFA30B6E7B36FF5388B1FF5A" box="[1351,1436,151,176]" metricMagnitude="3" metricUnit="m" metricValue="3.025" pageId="8" pageNumber="569" unit="m" value="3025.0">3025 m</quantity>
(
<figureCitation id="1336B9CFFFA30B6E7EEEFF788DC8FF21" box="[159,229,188,213]" captionStart="FIGURE 2" captionStartId="2.[151,250,1935,1957]" captionTargetBox="[156,1400,193,1832]" captionTargetId="figure@2.[156,1430,193,1913]" captionTargetPageId="2" captionText="FIGURE 2. Currently known distribution of the species of the Stigmella magnispinella group." httpUri="https://zenodo.org/record/191705/files/figure.png" pageId="8" pageNumber="569">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C317F6C1FFA30B6E7EB6FF1B8EDCFEE8" pageId="8" pageNumber="569" type="etymology">
<paragraph id="8BB2A54AFFA30B6E7EB6FF1B8EDCFEE8" blockId="8.[151,1436,151,285]" pageId="8" pageNumber="569">
<emphasis id="B9797958FFA30B6E7EB6FF1B8C6BFF0D" bold="true" box="[199,326,223,248]" pageId="8" pageNumber="569">Etymology</emphasis>
. The species name is derived from Latin
<emphasis id="B9797958FFA30B6E7D65FF248E77FF02" box="[788,858,224,247]" italics="true" pageId="8" pageNumber="569">varius</emphasis>
(varied, diverse) and
<emphasis id="B9797958FFA30B6E7A3FFF2489A6FF02" box="[1102,1163,224,247]" italics="true" pageId="8" pageNumber="569">spina</emphasis>
(a spine, thorn, bone) in reference to the combination of small and large cornuti in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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