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<document id="FFBCC10A0BF60F86AF880CF2C3D2C8A9" ID-DOI="10.11646/zootaxa.4103.3.8" ID-GBIF-Dataset="f9d4e9cc-6e22-4cba-b7ab-76a02bcbec74" ID-ISSN="1175-5326" ID-Zenodo-Dep="263832" ID-ZooBank="A2F6D402-EB14-47F6-954E-410D788B5B47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460530234840" checkinUser="plazi" docAuthor="Watanabe, Kyohei" docDate="2016" docId="0396C962A805CA37FF30F8B120B6C813" docLanguage="en" docName="zootaxa.4103.3.8.pdf.imf" docOrigin="Zootaxa 4103 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pion" docType="key" docVersion="9" lastPageNumber="294" masterDocId="FFAFB11AA801CA32FFA7FFFC252ACB06" masterDocTitle="First record of the genus Pion Schiødte, 1839 (Hymenoptera: Ichneumonidae: Ctenopelmatinae), from Japan, with description of a new species" masterLastPageNumber="294" masterPageNumber="289" pageNumber="293" updateTime="1698315690337" updateUser="plazi">
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<mods:title id="B725E4F39FD55CB9A7F1CE95F841F77C">First record of the genus Pion Schiødte, 1839 (Hymenoptera: Ichneumonidae: Ctenopelmatinae), from Japan, with description of a new species</mods:title>
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<mods:namePart id="CD320C4FB471451EF723A859D7CF3E62">Watanabe, Kyohei</mods:namePart>
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<treatment id="0396C962A805CA37FF30F8B120B6C813" ID-DOI="http://doi.org/10.5281/zenodo.6075409" ID-GBIF-Taxon="125042965" ID-Zenodo-Dep="6075409" LSID="urn:lsid:plazi:treatment:0396C962A805CA37FF30F8B120B6C813" httpUri="http://treatment.plazi.org/id/0396C962A805CA37FF30F8B120B6C813" lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293">
<subSubSection id="C3252BFFA805CA36FF30F8B12422CCE1" pageId="4" pageNumber="293" type="nomenclature">
<paragraph id="8B807874A805CA36FF30F8B127A9CC61" blockId="4.[151,643,1869,1895]" box="[151,643,1869,1895]" pageId="4" pageNumber="293">
<heading id="D0C8CF18A805CA36FF30F8B127A9CC61" bold="true" box="[151,643,1869,1895]" fontSize="11" level="1" pageId="4" pageNumber="293" reason="1">
<emphasis id="B94BA466A805CA36FF30F8B127A9CC61" bold="true" box="[151,643,1869,1895]" pageId="4" pageNumber="293">
Key to world species* of the genus
<taxonomicName id="4C3F03F7A805CA36FDEDF8B127A9CC61" ID-CoL="6QQB" ID-ENA="65268" box="[586,643,1869,1895]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B94BA466A805CA36FDEDF8B127A9CC61" bold="true" box="[586,643,1869,1895]" italics="true" pageId="4" pageNumber="293">Pion</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B807874A805CA36FF30F8682422CCE1" blockId="4.[151,1436,1940,2023]" pageId="4" pageNumber="293">
*
<taxonomicName id="4C3F03F7A805CA36FF03F86927CBCCAC" authority="Constantineanu &amp; Constantineanu, 1970" authorityName="Constantineanu &amp; Constantineanu" authorityYear="1970" box="[164,737,1940,1962]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="interstitialis">
<emphasis id="B94BA466A805CA36FF03F8692412CCAC" box="[164,312,1940,1962]" italics="true" pageId="4" pageNumber="293">P. interstitialis</emphasis>
Constantineanu &amp; Constantineanu, 1970
</taxonomicName>
, and
<taxonomicName id="4C3F03F7A805CA36FCBEF8692169CCAC" authority="Jaennicke, 1867" authorityName="Jaennicke" authorityYear="1867" box="[793,1091,1940,1962]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="schenkii">
<emphasis id="B94BA466A805CA36FCBEF86926ACCCAC" box="[793,902,1940,1962]" italics="true" pageId="4" pageNumber="293">P. schenkii</emphasis>
(Jaennicke, 1867)
</taxonomicName>
are not included because there is little useful information for identification in their original descriptions and I could not be examine the
<typeStatus id="5484C6D6A805CA36FB6AF8482028CCCF" box="[1229,1282,1972,1993]" pageId="4" pageNumber="293">types</typeStatus>
or determined specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3252BFFA804CA37FF30FF6B20B6C813" pageId="5" pageNumber="294" type="key">
<keyStep id="30CB60D1A804CA37FF30FF6B20B1CA02" pageId="5" pageNumber="294">
<paragraph id="8B807874A804CA37FF30FF6B20B6CBCC" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FF6B20B6CBCC" pageId="5" pageNumber="294">
1.
<collectingCountry id="F32838E4A804CA37FF46FF6B243BCBAA" box="[225,273,151,172]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
femur black in both sexes. Face black in male. Apex of metasoma red in female. Lateromedian longitudinal carina of propodeum parallel. Western Palaearctic region...................................... ...
<taxonomicName id="4C3F03F7A804CA37FB23FF4920B6CBCC" ID-CoL="4J2PR" authority="Schiodte, 1839" authorityName="Schiodte" authorityYear="1839" box="[1156,1436,181,202]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="B94BA466A804CA37FB23FF49202FCBCC" box="[1156,1285,181,202]" italics="true" pageId="5" pageNumber="294">Pion nigripes</emphasis>
<bibRefCitation id="EFAE0585A804CA37FAABFF4920B6CBCC" author="Schiodte" box="[1292,1436,181,202]" pageId="5" pageNumber="294" refString="Schiodte, G. (1839) Beretning om Resultaterne af en i Sommeren 1838 foretagen entomologisk Undersogelse af det sydlige Sjaelland, en Deel af Laaland, og Bornholm. Naturhistorisk Tidskrift, 2, 309 - 394." type="journal article" year="1839">Schiødte, 1839</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B807874A804CA37FF30FF2E20B1CA02" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FF2E20B1CA02" pageId="5" pageNumber="294">
-.
<collectingCountry id="F32838E4A804CA37FF46FF2E243BCBE1" box="[225,273,210,231]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
femur various in colouration. Face yellow in male (
<figureCitation id="130464F1A804CA37FCAEFF2E267BCBE1" box="[777,849,210,231]" captionStart="FIGURES 8 10" captionStartId="4.[151,264,1345,1368]" captionTargetBox="[151,1436,193,1323]" captionTargetId="figure@4.[151,1436,193,1324]" captionTargetPageId="4" captionText="FIGURES 8 10. Pion japonicum sp. nov. 8, female (holotype), dorsal habitus; 9, female (paratype), lateral habitus; 10, male (paratype), lateral habitus. Scale bar: 2.0 mm." httpUri="https://zenodo.org/record/263834/files/figure.png" pageId="5" pageNumber="294">Fig. 10</figureCitation>
). Apex of metasoma red or black (
<figureCitation id="130464F1A804CA37FB07FF2E202FCBE1" box="[1184,1285,210,231]" captionStart="FIGURES 8 10" captionStartId="4.[151,264,1345,1368]" captionTargetBox="[151,1436,193,1323]" captionTargetId="figure@4.[151,1436,193,1324]" captionTargetPageId="4" captionText="FIGURES 8 10. Pion japonicum sp. nov. 8, female (holotype), dorsal habitus; 9, female (paratype), lateral habitus; 10, male (paratype), lateral habitus. Scale bar: 2.0 mm." httpUri="https://zenodo.org/record/263834/files/figure.png" pageId="5" pageNumber="294">Figs 810</figureCitation>
). Lateromedian longitudinal carina of propodeum parallel or convergent anteriorly............................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CB60D1A804CA37FF30FEF120B1CA7C" pageId="5" pageNumber="294">
<paragraph id="8B807874A804CA37FF30FEF120B1CA39" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FEF120B1CA39" pageId="5" pageNumber="294">
2. Lateromedian longitudinal carina of propodeum distinctly convergent anteriorly. The spiracle of T1 situated at basal
<quantity id="4CC7D591A804CA37FAC5FEF120B6CA24" box="[1378,1436,269,290]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="294" unit="in" value="0.5">0.5 in</quantity>
female.............................................................................................. 3
</keyLead>
</paragraph>
<paragraph id="8B807874A804CA37FF30FEBB20B1CA7C" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FEBB20B1CA7C" pageId="5" pageNumber="294">
-. Lateromedian longitudinal carina of propodeum parallel (
<figureCitation id="130464F1A804CA37FCA0FEBB266BCA5A" box="[775,833,327,348]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1876,1899]" captionTargetBox="[167,1420,195,1844]" captionTargetId="figure@3.[151,1436,193,1856]" captionTargetPageId="3" captionText="FIGURES 1 7. Pion japonicum sp. nov., 1 3, female (holotype): 4 7, male (paratype). 1, head, frontal view, surface sculptures omitted except for left half of clypeus; 2, propodeum, dorsal view, surface sculptures omitted except for left half of dorsal face; 3, T 1, dorsal view; 4, subgenital plate, ventral view; 5, 6, genital capsule, dorsal (5) and ventral (6) view; 7, right aedeagus, lateral view." httpUri="https://zenodo.org/record/263833/files/figure.png" pageId="5" pageNumber="294">Fig. 2</figureCitation>
) or nearly so. The spiracle of T1 situated at basal
<quantity id="4CC7D591A804CA37FAB9FEBB207FCA5A" box="[1310,1365,327,348]" metricMagnitude="-2" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="294" unit="in" value="0.4">0.4 in</quantity>
female.................................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CB60D1A804CA37FF30FE7E20B1CAD4" pageId="5" pageNumber="294">
<paragraph id="8B807874A804CA37FF30FE7E20B1CAB2" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FE7E20B1CAB2" pageId="5" pageNumber="294">
3.
<collectingCountry id="F32838E4A804CA37FF46FE7E243BCA91" box="[225,273,386,407]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
femur entirely red in female, reddish-yellow to yellow with black base and apex in male. Oriental region (Southern part of
<collectingCountry id="F32838E4A804CA37FF5CFE63241DCAB2" box="[251,311,415,436]" name="China" pageId="5" pageNumber="294">China</collectingCountry>
).................................................................... ..
<taxonomicName id="4C3F03F7A804CA37FB26FE5C20B1CAB2" authority="Sheng, 2011" authorityName="Sheng" authorityYear="2011" box="[1153,1435,415,437]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="yifengensis">
<emphasis id="B94BA466A804CA37FB26FE5C2035CAB3" box="[1153,1311,415,437]" italics="true" pageId="5" pageNumber="294">Pion yifengensis</emphasis>
Sheng, 2011
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B807874A804CA37FF30FE4120B1CAD4" blockId="5.[151,1436,151,789]" box="[151,1435,445,466]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FE4120B1CAD4" box="[151,1435,445,466]" pageId="5" pageNumber="294">
-.
<collectingCountry id="F32838E4A804CA37FF46FE41243BCAD4" box="[225,273,445,466]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
femur red with black base and apex. Male unknown. Western Palaearctic region..........
<taxonomicName id="4C3F03F7A804CA37FB32FE4120B1CAD4" ID-CoL="4J2PV" authority="Bauer, 1958" authorityName="Bauer" authorityYear="1958" box="[1173,1435,445,466]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="stammeri">
<emphasis id="B94BA466A804CA37FB32FE41200BCAD4" box="[1173,1313,445,466]" italics="true" pageId="5" pageNumber="294">Pion stammeri</emphasis>
Bauer, 1958
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CB60D1A804CA37FF30FE2620B1C941" pageId="5" pageNumber="294">
<paragraph id="8B807874A804CA37FF30FE2620B6C90A" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FE2620B6C90A" pageId="5" pageNumber="294">
4. T2-T7 (or sometimes posterior part of T1) red and apex of metasoma red.
<collectingCountry id="F32838E4A804CA37FC6BFE2626D6CAE9" box="[972,1020,474,495]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
femur reddish-brown.
<collectingCountry id="F32838E4A804CA37FB46FE26203BCAE9" box="[1249,1297,474,495]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
tibia reddishbrown with black apex. Western Palaearctic region.................................
<taxonomicName id="4C3F03F7A804CA37FBF1FE0420B6C90A" ID-CoL="4J2PP" authority="Gravenhorst, 1829" authorityName="Gravenhorst" authorityYear="1829" box="[1110,1436,503,525]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="fortipes">
<emphasis id="B94BA466A804CA37FBF1FE0421F8C90B" box="[1110,1234,503,525]" italics="true" pageId="5" pageNumber="294">Pion fortipes</emphasis>
(Gravenhorst, 1829)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B807874A804CA37FF30FDE920B1C941" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FDE920B1C941" pageId="5" pageNumber="294">
-. T2 (or sometimes posterior part of T1)-T4 (or sometimes T5) red and apex of metasoma black (
<figureCitation id="130464F1A804CA37FBCAFDE921E9C92C" box="[1133,1219,533,554]" captionStart="FIGURES 8 10" captionStartId="4.[151,264,1345,1368]" captionTargetBox="[151,1436,193,1323]" captionTargetId="figure@4.[151,1436,193,1324]" captionTargetPageId="4" captionText="FIGURES 8 10. Pion japonicum sp. nov. 8, female (holotype), dorsal habitus; 9, female (paratype), lateral habitus; 10, male (paratype), lateral habitus. Scale bar: 2.0 mm." httpUri="https://zenodo.org/record/263834/files/figure.png" pageId="5" pageNumber="294">Figs 8, 9</figureCitation>
).
<collectingCountry id="F32838E4A804CA37FB73FDE9202EC92C" box="[1236,1284,533,554]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
femur and tibia various in colouration. Eastern Palaearctic region............................................................ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CB60D1A804CA37FF30FDB320B6C813" pageId="5" pageNumber="294">
<paragraph id="8B807874A804CA37FF30FDB320B6C999" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FDB320B6C999" pageId="5" pageNumber="294">
5. Lateromedian longitudinal carina almost parallel but slightly convergent anteriorly in female. Malar space 0.5 times as long as basal width of mandible. Antenna with 3034 flagellomeres.
<collectingCountry id="F32838E4A804CA37FCE7FD91265AC984" box="[832,880,621,642]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
femur black with a large red area in female.
<collectingCountry id="F32838E4A804CA37FA9FFD912042C984" box="[1336,1384,621,642]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
tibia entirely black in female. Male unknown. Northern part of
<collectingCountry id="F32838E4A804CA37FCAAFD76266CC999" box="[781,838,650,671]" name="China" pageId="5" pageNumber="294">China</collectingCountry>
............
<taxonomicName id="4C3F03F7A804CA37FC43FD7620B6C999" ID-CoL="4J2PT" authority="Chen, Sheng &amp; Mao, 1998" authorityName="Chen, Sheng &amp; Mao" authorityYear="1998" box="[996,1436,650,671]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="qinyuanensis">
<emphasis id="B94BA466A804CA37FC43FD7621BEC999" box="[996,1172,650,671]" italics="true" pageId="5" pageNumber="294">Pion qinyuanensis</emphasis>
Chen, Sheng &amp; Mao, 1998
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B807874A804CA37FF30FD5B20B6C813" blockId="5.[151,1436,151,789]" pageId="5" pageNumber="294">
<keyLead id="30CEDB41A804CA37FF30FD5B20B6C813" pageId="5" pageNumber="294">
-. Lateromedian longitudinal carina completely parallel, not convergent anteriorly in female (
<figureCitation id="130464F1A804CA37FBFEFD5B21BFC9BA" box="[1113,1173,679,700]" captionStart="FIGURES 1 7" captionStartId="3.[151,264,1876,1899]" captionTargetBox="[167,1420,195,1844]" captionTargetId="figure@3.[151,1436,193,1856]" captionTargetPageId="3" captionText="FIGURES 1 7. Pion japonicum sp. nov., 1 3, female (holotype): 4 7, male (paratype). 1, head, frontal view, surface sculptures omitted except for left half of clypeus; 2, propodeum, dorsal view, surface sculptures omitted except for left half of dorsal face; 3, T 1, dorsal view; 4, subgenital plate, ventral view; 5, 6, genital capsule, dorsal (5) and ventral (6) view; 7, right aedeagus, lateral view." httpUri="https://zenodo.org/record/263833/files/figure.png" pageId="5" pageNumber="294">Fig. 2</figureCitation>
). Malar space 0.6 times as long as basal width of mandible in female. Antenna with 2829 flagellomeres.
<collectingCountry id="F32838E4A804CA37FC7EFD392123C9DC" box="[985,1033,709,730]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
femur black in female (rarely tinged with red) (
<figureCitation id="130464F1A804CA37FEB1FD1E247BC9F1" box="[278,337,738,759]" captionStart="FIGURES 8 10" captionStartId="4.[151,264,1345,1368]" captionTargetBox="[151,1436,193,1323]" captionTargetId="figure@4.[151,1436,193,1324]" captionTargetPageId="4" captionText="FIGURES 8 10. Pion japonicum sp. nov. 8, female (holotype), dorsal habitus; 9, female (paratype), lateral habitus; 10, male (paratype), lateral habitus. Scale bar: 2.0 mm." httpUri="https://zenodo.org/record/263834/files/figure.png" pageId="5" pageNumber="294">Fig. 9</figureCitation>
), black with a ventral longitudinal stripe in male (
<figureCitation id="130464F1A804CA37FC9AFD1E26ACC9F1" box="[829,902,738,759]" captionStart="FIGURES 8 10" captionStartId="4.[151,264,1345,1368]" captionTargetBox="[151,1436,193,1323]" captionTargetId="figure@4.[151,1436,193,1324]" captionTargetPageId="4" captionText="FIGURES 8 10. Pion japonicum sp. nov. 8, female (holotype), dorsal habitus; 9, female (paratype), lateral habitus; 10, male (paratype), lateral habitus. Scale bar: 2.0 mm." httpUri="https://zenodo.org/record/263834/files/figure.png" pageId="5" pageNumber="294">Fig. 10</figureCitation>
).
<collectingCountry id="F32838E4A804CA37FC3FFD1E26E2C9F1" box="[920,968,738,759]" name="India" pageId="5" pageNumber="294">Hind</collectingCountry>
tibia yellowish-brown basally, black apically in female (
<figureCitation id="130464F1A804CA37FE88FD032440C812" box="[303,362,767,788]" captionStart="FIGURES 8 10" captionStartId="4.[151,264,1345,1368]" captionTargetBox="[151,1436,193,1323]" captionTargetId="figure@4.[151,1436,193,1324]" captionTargetPageId="4" captionText="FIGURES 8 10. Pion japonicum sp. nov. 8, female (holotype), dorsal habitus; 9, female (paratype), lateral habitus; 10, male (paratype), lateral habitus. Scale bar: 2.0 mm." httpUri="https://zenodo.org/record/263834/files/figure.png" pageId="5" pageNumber="294">Fig. 9</figureCitation>
).
<collectingCountry id="F32838E4A804CA37FEDBFD032499C812" box="[380,435,767,788]" name="Japan" pageId="5" pageNumber="294">Japan</collectingCountry>
.......................................................
<taxonomicName id="4C3F03F7A804CA37FBEAFCFC2063C812" authority="Watanabe" authorityName="Watanabe" box="[1101,1353,767,789]" class="Insecta" family="Ichneumonidae" genus="Pion" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="japonicum" status="sp. nov.">
<emphasis id="B94BA466A804CA37FBEAFCFC21CFC813" box="[1101,1253,768,789]" italics="true" pageId="5" pageNumber="294">Pion japonicum</emphasis>
Watanabe
</taxonomicName>
<emphasis id="B94BA466A804CA37FAE8FD0320B6C813" bold="true" box="[1359,1436,767,789]" pageId="5" pageNumber="294">
<taxonomicNameLabel id="A278191DA804CA37FAE8FD0320B6C813" box="[1359,1436,767,789]" pageId="5" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>