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<document id="5549865D78C0BE774322EAC435970107" ID-DOI="10.11646/zootaxa.4027.3.2" ID-GBIF-Dataset="89c5bda8-87ad-4f90-a154-2de036e9c71e" ID-ISSN="1175-5326" ID-Zenodo-Dep="241473" ID-ZooBank="8B613537-E7D0-4BDB-B168-943C5B54B0BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461282934901" checkinUser="plazi" docAuthor="Behounek, G., Han, H. L. &amp; Kononenko, V. S." docDate="2015" docId="039687F0FFF4FFF0FF2AFB75B0C8FB64" docLanguage="en" docName="zt04027p365.pdf" docOrigin="Zootaxa 4027 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belciana" docType="treatment" docVersion="8" lastPageNumber="347" masterDocId="FFAFFF88FFF1FFF6FFBDFF81B55EFF83" masterDocTitle="A revision of the genus Belciana Walker, 1862 with description of three new species (Lepidoptera, Noctuidae: Pantheinae) from East and South East Asia. Revision of Pantheinae, contribution XII" masterLastPageNumber="365" masterPageNumber="341" pageNumber="346" updateTime="1698614449975" updateUser="plazi">
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<mods:title id="9879F6D760FF2D27A7F80E97038EF776">A revision of the genus Belciana Walker, 1862 with description of three new species (Lepidoptera, Noctuidae: Pantheinae) from East and South East Asia. Revision of Pantheinae, contribution XII</mods:title>
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<treatment id="039687F0FFF4FFF0FF2AFB75B0C8FB64" ID-DOI="http://doi.org/10.5281/zenodo.6116199" ID-GBIF-Taxon="119640895" ID-Zenodo-Dep="6116199" LSID="urn:lsid:plazi:treatment:039687F0FFF4FFF0FF2AFB75B0C8FB64" httpUri="http://treatment.plazi.org/id/039687F0FFF4FFF0FF2AFB75B0C8FB64" lastPageId="6" lastPageNumber="347" pageId="5" pageNumber="346">
<subSubSection id="C325656DFFF4FFF3FF2AFB75B447FAAC" pageId="5" pageNumber="346" type="nomenclature">
<paragraph id="8B8036E6FFF4FFF3FF2AFB75B497FA8D" blockId="5.[151,457,1268,1327]" box="[151,457,1268,1294]" pageId="5" pageNumber="346">
<heading id="D0C8818AFFF4FFF3FF2AFB75B497FA8D" bold="true" box="[151,457,1268,1294]" fontSize="11" level="1" pageId="5" pageNumber="346" reason="1">
<emphasis id="B94BEAF4FFF4FFF3FF2AFB75B497FA8D" bold="true" box="[151,457,1268,1294]" pageId="5" pageNumber="346">
<taxonomicName id="4C3F4D65FFF4FFF3FF2AFB75B45DFA8D" ID-CoL="923P6" box="[151,259,1268,1294]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAF4FFF4FFF3FF2AFB75B45DFA8D" bold="true" box="[151,259,1268,1294]" italics="true" pageId="5" pageNumber="346">Belciana</emphasis>
</taxonomicName>
sp. (Irian Jaya)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8036E6FFF4FFF3FF2AFA97B447FAAC" blockId="5.[151,457,1268,1327]" box="[151,281,1302,1327]" pageId="5" pageNumber="346">
(
<figureCitation id="13042A63FFF4FFF3FF22FA97B5B9FAAD" box="[159,231,1302,1327]" captionStart="FIGURES 1 8" captionStartId="16.[151,263,1838,1861]" captionTargetBox="[154,1436,181,1808]" captionTargetId="figure@16.[154,1439,181,1809]" captionTargetPageId="16" captionText="FIGURES 1 8. Belciana spp., adults. 1. B. biformis, male, Sumatra, slide GB / ZSM N 4159 (ZSM); 2. B. biformis, male, Sumatra, slide GB / ZSM N 4160 (ZSM); 3. B. biformis, female, Sumatra, slide GB 7359 (GB); 4. B. biformis, female, Sumatra, slide GB / ZSM N 4198 (ZSM); 5. B. hemodoides, male, Philippines, Palawan, slide GB 7389 (GB); 6. B. hemodoides, female, Philippines, Palawan, slide GB / ZSM N 4286 (ZSM); 7. B. sp., female, Irian Jaya, slide GB 7363 (GB); 8. B. sulawesiana sp. n., male, holotype, Sulawesi, slide GB 7381 (GB)." httpUri="https://zenodo.org/record/241474/files/figure.png" pageId="5" pageNumber="346">Figs 7</figureCitation>
,
<figureCitation id="13042A63FFF4FFF3FF4EFA97B44EFAAC" box="[243,272,1302,1327]" captionStart="FIGURES 41 45" captionStartId="23.[151,263,1841,1864]" captionTargetBox="[158,1426,223,1811]" captionTargetId="figure@23.[151,1436,193,1821]" captionTargetPageId="23" captionText="FIGURES 41 45. Belciana and Diphteroides spp., male (41) and female (42 45) genitalia. 41. B. scorpio, Thailand, slide GB / ZSM N 4215 (ZSM); 42. D. kendricki, Irian Jaya, slide GB 7372 (GB); 43. B. biformis, Sumatra, slide GB / ZSM N 4158 (ZSM); 44. B. hemodoides, Borneo, slide GB 7389 (GB); 45. B. sp., Irian Jaya, slide GB 7363 (GB). Most distinctive characters are marked by lines." httpUri="https://zenodo.org/record/241481/files/figure.png" pageId="5" pageNumber="346">45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C325656DFFF4FFF3FF2AFADFB4A1FA3C" pageId="5" pageNumber="346" type="materials_examined">
<paragraph id="8B8036E6FFF4FFF3FF2AFADFB4A1FA3C" blockId="5.[151,1436,1374,1975]" pageId="5" pageNumber="346">
<emphasis id="B94BEAF4FFF4FFF3FF2AFADFB4DBFAF4" bold="true" box="[151,389,1374,1399]" pageId="5" pageNumber="346">Material examined.</emphasis>
<collectionCode id="ED2EAE23FFF4FFF3FE2DFADEB775FAF4" box="[400,555,1375,1399]" pageId="5" pageNumber="346">
<collectingCountry id="F3287676FFF4FFF3FE2DFADEB775FAF4" box="[400,555,1375,1399]" name="Indonesia" pageId="5" pageNumber="346">INDONESIA</collectingCountry>
</collectionCode>
,
<collectionCode id="ED2EAE23FFF4FFF3FD81FADEB722FAF5" box="[572,636,1375,1398]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="5" pageNumber="346">NEW</collectionCode>
<collectionCode id="ED2EAE23FFF4FFF3FD3AFADEB7A9FAF4" box="[647,759,1375,1399]" pageId="5" pageNumber="346">
<collectingCountry id="F3287676FFF4FFF3FD3AFADEB7A9FAF4" box="[647,759,1375,1399]" name="Guinea-Bissau" pageId="5" pageNumber="346">GUINEA</collectingCountry>
</collectionCode>
:
<specimenCount id="9D39FD6FFFF4FFF3FCBBFADEB630FAF4" box="[774,878,1374,1399]" pageId="5" pageNumber="346" type="female">1 female</specimenCount>
, Reg. Irian Jaya, Arfak Mts., Nabire,
<quantity id="4CC79B03FFF4FFF3FA89FADEB021FAF4" box="[1332,1407,1374,1399]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="346" unit="km" value="12.0">12 km</quantity>
E Samabusa Lagari,
<quantity id="4CC79B03FFF4FFF3FED1FA03B4F7FA19" box="[364,425,1410,1435]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="346" unit="m" value="50.0">50 m</quantity>
, (primary forest),
<geoCoordinate id="EE0B5021FFF4FFF3FDC0FA03B7BFFA18" box="[637,737,1410,1435]" direction="south" orientation="latitude" pageId="5" pageNumber="346" precision="925" value="-3.3666668">03°22 S</geoCoordinate>
, 135°29 E, Brechlin R. &amp; Cerny K. leg.,
<date id="FF811026FFF4FFF3FB7AFA03B00CFA18" box="[1223,1362,1410,1435]" pageId="5" pageNumber="346" value="1993-12-06" valueMax="1993-12-08" valueMin="1993-12-06">6-8.xii.1993</date>
, slide, GB7363 female, coll. GB/
<collectionCode id="ED2EAE23FFF4FFF3FE03FA26B4A4FA3C" box="[446,506,1447,1471]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="346">ZSM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C325656DFFF4FFF3FF7AFA4BB429F8A4" pageId="5" pageNumber="346" type="diagnosis">
<paragraph id="8B8036E6FFF4FFF3FF7AFA4BB01EF880" blockId="5.[151,1436,1374,1975]" pageId="5" pageNumber="346">
<emphasis id="B94BEAF4FFF4FFF3FF7AFA4BB4D0FA60" bold="true" box="[199,398,1482,1507]" pageId="5" pageNumber="346">Diagnosis. Adult</emphasis>
(
<figureCitation id="13042A63FFF4FFF3FE22FA4BB4BBFA60" box="[415,485,1482,1507]" captionStart="FIGURES 1 8" captionStartId="16.[151,263,1838,1861]" captionTargetBox="[154,1436,181,1808]" captionTargetId="figure@16.[154,1439,181,1809]" captionTargetPageId="16" captionText="FIGURES 1 8. Belciana spp., adults. 1. B. biformis, male, Sumatra, slide GB / ZSM N 4159 (ZSM); 2. B. biformis, male, Sumatra, slide GB / ZSM N 4160 (ZSM); 3. B. biformis, female, Sumatra, slide GB 7359 (GB); 4. B. biformis, female, Sumatra, slide GB / ZSM N 4198 (ZSM); 5. B. hemodoides, male, Philippines, Palawan, slide GB 7389 (GB); 6. B. hemodoides, female, Philippines, Palawan, slide GB / ZSM N 4286 (ZSM); 7. B. sp., female, Irian Jaya, slide GB 7363 (GB); 8. B. sulawesiana sp. n., male, holotype, Sulawesi, slide GB 7381 (GB)." httpUri="https://zenodo.org/record/241474/files/figure.png" pageId="5" pageNumber="346">Fig. 7</figureCitation>
). Wingspan
<quantity id="4CC79B03FFF4FFF3FDCBFA4AB794FA61" box="[630,714,1483,1507]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="346" unit="mm" value="39.0">39 mm</quantity>
. This unknown species is close and similar in appearance to
<taxonomicName id="4C3F4D65FFF4FFF3FA39FA4DB5AAF984" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF4FFF3FA39FA4DB5AAF984" italics="true" pageId="5" pageNumber="346">B. biformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D65FFF4FFF3FE8DFA6EB486F984" box="[304,472,1518,1543]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF4FFF3FE8DFA6EB486F984" box="[304,472,1518,1543]" italics="true" pageId="5" pageNumber="346">B. hemodoides</emphasis>
</taxonomicName>
, but differs from both by the structure of female genitalia, particularly by shape of antrum and antevaginal plate, shape of sclerotised area in joining of ductus with corpus bursae and by shape of corpus bursae.
<emphasis id="B94BEAF4FFF4FFF3FEFBF9B7B751F9CC" bold="true" box="[326,527,1590,1615]" pageId="5" pageNumber="346">Female genitalia</emphasis>
(
<figureCitation id="13042A63FFF4FFF3FD9CF9B7B727F9CC" box="[545,633,1590,1615]" captionStart="FIGURES 41 45" captionStartId="23.[151,263,1841,1864]" captionTargetBox="[158,1426,223,1811]" captionTargetId="figure@23.[151,1436,193,1821]" captionTargetPageId="23" captionText="FIGURES 41 45. Belciana and Diphteroides spp., male (41) and female (42 45) genitalia. 41. B. scorpio, Thailand, slide GB / ZSM N 4215 (ZSM); 42. D. kendricki, Irian Jaya, slide GB 7372 (GB); 43. B. biformis, Sumatra, slide GB / ZSM N 4158 (ZSM); 44. B. hemodoides, Borneo, slide GB 7389 (GB); 45. B. sp., Irian Jaya, slide GB 7363 (GB). Most distinctive characters are marked by lines." httpUri="https://zenodo.org/record/241481/files/figure.png" pageId="5" pageNumber="346">Fig. 45</figureCitation>
). Ovipositor quadrangular; apophyses anteriores and posteriores ones moderate long and equal in length, rather strong; antrum much narrower than in
<taxonomicName id="4C3F4D65FFF4FFF3FBA0F9DDB1C7F9F0" box="[1053,1177,1626,1651]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF4FFF3FBA0F9DDB1C7F9F0" box="[1053,1177,1626,1651]" italics="true" pageId="5" pageNumber="346">B. biformis</emphasis>
</taxonomicName>
, but broader than in
<taxonomicName id="4C3F4D65FFF4FFF3FA39F9DDB443F914" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF4FFF3FA39F9DDB443F914" italics="true" pageId="5" pageNumber="346">B. hemodoides</emphasis>
</taxonomicName>
, strongly split; lateral sides of the antevaginal plate form two acute sclerotised protuberances; ductus bursae longer and broader than in
<taxonomicName id="4C3F4D65FFF4FFF3FDA4F925B7C8F938" box="[537,662,1698,1723]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF4FFF3FDA4F925B7C8F938" box="[537,662,1698,1723]" italics="true" pageId="5" pageNumber="346">B. biformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D65FFF4FFF3FD73F925B62DF938" box="[718,883,1698,1723]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF4FFF3FD73F925B62DF938" box="[718,883,1698,1723]" italics="true" pageId="5" pageNumber="346">B. hemodoides</emphasis>
</taxonomicName>
; the area in joining of ductus with corpus bursae less sclerotised, but more ribbed and more amorphous compared with two those species; corpus bursae broad, constricted to its caudal part, with prominent extension in caudal part, much bigger than in
<emphasis id="B94BEAF4FFF4FFF3FB29F96DB01EF880" box="[1172,1344,1770,1795]" italics="true" pageId="5" pageNumber="346">
<taxonomicName id="4C3F4D65FFF4FFF3FB29F96DB063F880" box="[1172,1341,1770,1795]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="hemodoides">B. hemodoides</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B8036E6FFF4FFF3FF7AF88FB429F8A4" blockId="5.[151,1436,1374,1975]" box="[199,375,1806,1831]" pageId="5" pageNumber="346">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="C325656DFFF4FFF0FF7AF8B3B447FF50" lastPageId="6" lastPageNumber="347" pageId="5" pageNumber="346" type="distribution">
<paragraph id="8B8036E6FFF4FFF3FF7AF8B3B58EF8EC" blockId="5.[151,1436,1374,1975]" pageId="5" pageNumber="346">
<emphasis id="B94BEAF4FFF4FFF3FF7AF8B3B742F8C8" bold="true" box="[199,540,1842,1867]" pageId="5" pageNumber="346">Distribution and bionomics.</emphasis>
Irian Jaya (New
<collectingCountry id="F3287676FFF4FFF3FD57F8B3B61EF8C8" box="[746,832,1842,1867]" name="Guinea-Bissau" pageId="5" pageNumber="346">Guinea</collectingCountry>
). The species was collected in the primary forest in May.
</paragraph>
<paragraph id="8B8036E6FFF4FFF3FF7AF8FBB792F834" blockId="5.[151,1436,1374,1975]" pageId="5" pageNumber="346">
<emphasis id="B94BEAF4FFF4FFF3FF7AF8FBB458F810" bold="true" box="[199,262,1914,1939]" pageId="5" pageNumber="346">Note.</emphasis>
Althouth the species substantively differs by female genitalia from its allies, we cannot describe it before collecting more material sufficient for description.
</paragraph>
<paragraph id="8B8036E6FFF7FFF0FF2AFF16B447FF50" blockId="6.[151,485,151,211]" pageId="6" pageNumber="347">
<taxonomicName id="4C3F4D65FFF7FFF0FF2AFF16B4C0FF32" box="[151,414,151,177]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="sulawesiana" status="sp. nov.">
<emphasis id="B94BEAF4FFF7FFF0FF2AFF16B4C0FF32" bold="true" box="[151,414,151,177]" italics="true" pageId="6" pageNumber="347">Belciana sulawesiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A278578FFFF7FFF0FE18FF19B4BBFF32" box="[421,485,152,177]" pageId="6" pageNumber="347" rank="species">
<emphasis id="B94BEAF4FFF7FFF0FE18FF19B4BBFF32" bold="true" box="[421,485,152,177]" pageId="6" pageNumber="347">sp. n.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13042A63FFF7FFF0FF22FF3BB5B9FF50" box="[159,231,186,211]" captionStart="FIGURES 1 8" captionStartId="16.[151,263,1838,1861]" captionTargetBox="[154,1436,181,1808]" captionTargetId="figure@16.[154,1439,181,1809]" captionTargetPageId="16" captionText="FIGURES 1 8. Belciana spp., adults. 1. B. biformis, male, Sumatra, slide GB / ZSM N 4159 (ZSM); 2. B. biformis, male, Sumatra, slide GB / ZSM N 4160 (ZSM); 3. B. biformis, female, Sumatra, slide GB 7359 (GB); 4. B. biformis, female, Sumatra, slide GB / ZSM N 4198 (ZSM); 5. B. hemodoides, male, Philippines, Palawan, slide GB 7389 (GB); 6. B. hemodoides, female, Philippines, Palawan, slide GB / ZSM N 4286 (ZSM); 7. B. sp., female, Irian Jaya, slide GB 7363 (GB); 8. B. sulawesiana sp. n., male, holotype, Sulawesi, slide GB 7381 (GB)." httpUri="https://zenodo.org/record/241474/files/figure.png" pageId="6" pageNumber="347">Figs 8</figureCitation>
,
<figureCitation id="13042A63FFF7FFF0FF4EFF3AB44EFF50" box="[243,272,187,211]" captionStart="FIGURES 32 34" captionStartId="20.[151,263,1841,1864]" captionTargetBox="[156,1432,196,1814]" captionTargetId="figure@20.[151,1436,193,1821]" captionTargetPageId="20" captionText="FIGURES 32 34. Belciana spp., male genitalia. 32. B. biformis, Sumatra, slide GB / ZSM N 4159 (ZSM); 33. B. hemodoides, Philippines, Mindoro, slide GB 7380 (GB); 34. B. sulawesiana sp. n., holotype, Sulawesi GB 7381 (GB). Most distinctive characters are marked by lines." httpUri="https://zenodo.org/record/241478/files/figure.png" pageId="6" pageNumber="347">34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C325656DFFF7FFF0FF2AFE83B149FEBC" pageId="6" pageNumber="347" type="materials_examined">
<paragraph id="8B8036E6FFF7FFF0FF2AFE83B149FEBC" blockId="6.[151,1437,258,1255]" pageId="6" pageNumber="347">
<emphasis id="B94BEAF4FFF7FFF0FF2AFE83B463FE98" bold="true" box="[151,317,258,283]" pageId="6" pageNumber="347">
<typeStatus id="54848844FFF7FFF0FF2AFE83B58CFE98" box="[151,210,258,283]" pageId="6" pageNumber="347">Type</typeStatus>
material
</emphasis>
.
<emphasis id="B94BEAF4FFF7FFF0FEF7FE83B49EFE98" bold="true" box="[330,448,258,283]" pageId="6" pageNumber="347">
<typeStatus id="54848844FFF7FFF0FEF7FE83B4E5FE98" box="[330,443,258,283]" pageId="6" pageNumber="347" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED2EAE23FFF7FFF0FE7AFE82B73CFE98" box="[455,610,259,283]" pageId="6" pageNumber="347">
<collectingCountry id="F3287676FFF7FFF0FE7AFE82B73CFE98" box="[455,610,259,283]" name="Indonesia" pageId="6" pageNumber="347">INDONESIA</collectingCountry>
</collectionCode>
, SULAWESI: male, Reg. Sulawesi Selatan, Puncak, Palopo,
<quantity id="4CC79B03FFF7FFF0FAB3FE83B0C9FE99" box="[1294,1431,258,283]" metricMagnitude="2" metricUnit="m" metricValue="9.75" metricValueMax="13.0" metricValueMin="6.5" pageId="6" pageNumber="347" unit="m" value="975.0" valueMax="1300.0" valueMin="650.0">650-1300 m</quantity>
, 02°55 S ,
<geoCoordinate id="EE0B5021FFF7FFF0FEB8FEA7B429FEBD" box="[261,375,294,319]" direction="east" orientation="longitude" pageId="6" pageNumber="347" precision="925" value="120.083336">120°05 E</geoCoordinate>
, Brechlin R. ex coll.,
<date id="FF811026FFF7FFF0FDD6FEA6B7E5FEBC" box="[619,699,295,319]" pageId="6" pageNumber="347" value="1997-10">x.1997</date>
, slide GB7381, coll. GB/
<collectionCode id="ED2EAE23FFF7FFF0FC6BFEA6B14CFEBC" box="[982,1042,295,319]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="347">ZSM</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C325656DFFF7FFF0FF7AFECBB720FE70" pageId="6" pageNumber="347" type="diagnosis">
<paragraph id="8B8036E6FFF7FFF0FF7AFECBB720FE70" blockId="6.[151,1437,258,1255]" pageId="6" pageNumber="347">
<emphasis id="B94BEAF4FFF7FFF0FF7AFECBB41FFEE0" bold="true" box="[199,321,330,355]" pageId="6" pageNumber="347">Diagnosis.</emphasis>
The new species is close to
<taxonomicName id="4C3F4D65FFF7FFF0FD37FECDB659FEE0" box="[650,775,330,355]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FD37FECDB659FEE0" box="[650,775,330,355]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D65FFF7FFF0FCFCFECDB6B6FEE0" box="[833,1000,330,355]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF7FFF0FCFCFECDB6B6FEE0" box="[833,1000,330,355]" italics="true" pageId="6" pageNumber="347">B. hemodoides</emphasis>
</taxonomicName>
, while the ground wing colouration is more similar to
<taxonomicName id="4C3F4D65FFF7FFF0FEF4FEEEB49BFE04" box="[329,453,366,391]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FEF4FEEEB49BFE04" box="[329,453,366,391]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
. Externally it differs from the last by darker blackish subbasal field, smaller medial and subterminal patches in costal area and by expressed orbicular as white spot and reniform as white spot with black dot. In male genitalia it differs from both species by broader valva, with broader sclerotised part, broader harpe and structure of aedeagus and arming of vesica.
</paragraph>
</subSubSection>
<subSubSection id="C325656DFFF7FFF0FF7AFE7FB73DFB1C" pageId="6" pageNumber="347" type="description">
<paragraph id="8B8036E6FFF7FFF0FF7AFE7FB776FC20" blockId="6.[151,1437,258,1255]" pageId="6" pageNumber="347">
<emphasis id="B94BEAF4FFF7FFF0FF7AFE7FB4FBFD94" bold="true" box="[199,421,510,535]" pageId="6" pageNumber="347">Description. Adult</emphasis>
(
<figureCitation id="13042A63FFF7FFF0FE08FE7FB4A4FD94" box="[437,506,510,535]" captionStart="FIGURES 1 8" captionStartId="16.[151,263,1838,1861]" captionTargetBox="[154,1436,181,1808]" captionTargetId="figure@16.[154,1439,181,1809]" captionTargetPageId="16" captionText="FIGURES 1 8. Belciana spp., adults. 1. B. biformis, male, Sumatra, slide GB / ZSM N 4159 (ZSM); 2. B. biformis, male, Sumatra, slide GB / ZSM N 4160 (ZSM); 3. B. biformis, female, Sumatra, slide GB 7359 (GB); 4. B. biformis, female, Sumatra, slide GB / ZSM N 4198 (ZSM); 5. B. hemodoides, male, Philippines, Palawan, slide GB 7389 (GB); 6. B. hemodoides, female, Philippines, Palawan, slide GB / ZSM N 4286 (ZSM); 7. B. sp., female, Irian Jaya, slide GB 7363 (GB); 8. B. sulawesiana sp. n., male, holotype, Sulawesi, slide GB 7381 (GB)." httpUri="https://zenodo.org/record/241474/files/figure.png" pageId="6" pageNumber="347">Fig. 8</figureCitation>
). Wingspan
<quantity id="4CC79B03FFF7FFF0FD35FE7EB782FD95" box="[648,732,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="347" unit="mm" value="43.0">43 mm</quantity>
. Tegulae brown; patagia pale bluish-green, the thorax dorsally between patagia bluish-green like
<taxonomicName id="4C3F4D65FFF7FFF0FD9DFDA5B7C1FDB8" box="[544,671,546,571]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FD9DFDA5B7C1FDB8" box="[544,671,546,571]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
and
<emphasis id="B94BEAF4FFF7FFF0FD67FDA5B6D7FDB8" box="[730,905,546,571]" italics="true" pageId="6" pageNumber="347">
<taxonomicName id="4C3F4D65FFF7FFF0FD67FDA5B6DBFDB8" box="[730,901,546,571]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="hemodoides">B. hemodoides</taxonomicName>
.
</emphasis>
Ground colour of forewing pale bluish-green, like in
<taxonomicName id="4C3F4D65FFF7FFF0FF4EFDC6B428FDDC" box="[243,374,582,607]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FF4EFDC6B428FDDC" box="[243,374,582,607]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
; subbasal field filled with black-brown, broader and darker than in
<taxonomicName id="4C3F4D65FFF7FFF0FB06FDC6B061FDDC" box="[1211,1343,582,607]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FB06FDC6B061FDDC" box="[1211,1343,582,607]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D65FFF7FFF0FA39FDC6B443FD00" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF7FFF0FA39FDC6B443FD00" italics="true" pageId="6" pageNumber="347">B. hemodoides</emphasis>
</taxonomicName>
; medial field with diffused patches of darker green, and diffused whitish spots; medial line distinct blackish, not reach costal mark, which is less expressed than in
<taxonomicName id="4C3F4D65FFF7FFF0FC37FD0EB154FD24" box="[906,1034,654,679]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FC37FD0EB154FD24" box="[906,1034,654,679]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D65FFF7FFF0FBF5FD0EB1AFFD24" box="[1096,1265,654,679]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF7FFF0FBF5FD0EB1AFFD24" box="[1096,1265,654,679]" italics="true" pageId="6" pageNumber="347">B. hemodoides</emphasis>
</taxonomicName>
; orbicular and reniform stigmata expressed as diffused whitish spots, reniform with blackish dot in bottom; submedial and antemedial lines as diffused row of whitish spots; subterminal line blackish bordered with whitish inwardly; brown apical-subapical patch more contrast and broader, quadrangular, compared with
<taxonomicName id="4C3F4D65FFF7FFF0FB8CFD7DB1EEFC90" box="[1073,1200,762,787]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FB8CFD7DB1EEFC90" box="[1073,1200,762,787]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
, not connected with subtornal brown patch, which is smaller than in
<taxonomicName id="4C3F4D65FFF7FFF0FD78FC9EB61BFCB4" box="[709,837,798,823]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FD78FC9EB61BFCB4" box="[709,837,798,823]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D65FFF7FFF0FC3DFC9EB179FCB4" box="[896,1063,798,823]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF7FFF0FC3DFC9EB179FCB4" box="[896,1063,798,823]" italics="true" pageId="6" pageNumber="347">B. hemodoides</emphasis>
</taxonomicName>
; terminal line as a row of black vertical streaks; cilia brownish-grey, pale in base. Hindwing dark greyish outwardly, pale greyish-yellow inwardly, with traceable discal spot, medial band and blackish interrupted terminal line with thin whitish bordering; cilia brownish-grey with yellowish-grey.
</paragraph>
<paragraph id="8B8036E6FFF7FFF0FF7AFC2FB740FBF8" blockId="6.[151,1437,258,1255]" pageId="6" pageNumber="347">
<emphasis id="B94BEAF4FFF7FFF0FF7AFC2FB42FFC44" bold="true" box="[199,369,942,967]" pageId="6" pageNumber="347">Male genitalia</emphasis>
(
<figureCitation id="13042A63FFF7FFF0FE3DFC2FB48AFC44" box="[384,468,942,967]" captionStart="FIGURES 32 34" captionStartId="20.[151,263,1841,1864]" captionTargetBox="[156,1432,196,1814]" captionTargetId="figure@20.[151,1436,193,1821]" captionTargetPageId="20" captionText="FIGURES 32 34. Belciana spp., male genitalia. 32. B. biformis, Sumatra, slide GB / ZSM N 4159 (ZSM); 33. B. hemodoides, Philippines, Mindoro, slide GB 7380 (GB); 34. B. sulawesiana sp. n., holotype, Sulawesi GB 7381 (GB). Most distinctive characters are marked by lines." httpUri="https://zenodo.org/record/241478/files/figure.png" pageId="6" pageNumber="347">Fig. 34</figureCitation>
). Compared with
<taxonomicName id="4C3F4D65FFF7FFF0FD21FC2EB646FC44" box="[668,792,942,967]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FD21FC2EB646FC44" box="[668,792,942,967]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3F4D65FFF7FFF0FCF3FC2EB6ACFC44" box="[846,1010,942,967]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF7FFF0FCF3FC2EB6ACFC44" box="[846,1010,942,967]" italics="true" pageId="6" pageNumber="347">B. hemodoides</emphasis>
</taxonomicName>
the new species has somewhat longer uncus, which is more solid than in
<taxonomicName id="4C3F4D65FFF7FFF0FDA0FC55B7C7FC68" box="[541,665,978,1003]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="B94BEAF4FFF7FFF0FDA0FC55B7C7FC68" box="[541,665,978,1003]" italics="true" pageId="6" pageNumber="347">B. biformis</emphasis>
</taxonomicName>
, broader valva with less developed coremata, wider sclerotised area, shorter and more flat juxta. Aedeagus with dorsal branch of carina bearing very short extension much smaller than in
<taxonomicName id="4C3F4D65FFF7FFF0FF08FB9DB402FBB0" box="[181,348,1050,1075]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF7FFF0FF08FB9DB402FBB0" box="[181,348,1050,1075]" italics="true" pageId="6" pageNumber="347">B. hemodoides</emphasis>
</taxonomicName>
; vesica basally bulbous, apically tubular, with 68 small cornuti, irregular tiny cornuti and one separated cornutus, all are smaller than in
<taxonomicName id="4C3F4D65FFF7FFF0FDCBFBBEB642FBD4" box="[630,796,1086,1111]" class="Insecta" family="Noctuidae" genus="Belciana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="hemodoides">
<emphasis id="B94BEAF4FFF7FFF0FDCBFBBEB642FBD4" box="[630,796,1086,1111]" italics="true" pageId="6" pageNumber="347">B. hemodoides</emphasis>
</taxonomicName>
; ventral branch of carina with small patch of tiny spines more dense than in related species.
</paragraph>
<paragraph id="8B8036E6FFF7FFF0FF7AFB07B73DFB1C" blockId="6.[151,1437,258,1255]" box="[199,611,1158,1183]" pageId="6" pageNumber="347">
<emphasis id="B94BEAF4FFF7FFF0FF7AFB07B4CDFB1C" bold="true" box="[199,403,1158,1183]" pageId="6" pageNumber="347">Female genitalia.</emphasis>
Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C325656DFFF7FFF0FF7AFB2BB063FB40" box="[199,1341,1194,1219]" pageId="6" pageNumber="347" type="etymology">
<paragraph id="8B8036E6FFF7FFF0FF7AFB2BB063FB40" blockId="6.[151,1437,258,1255]" box="[199,1341,1194,1219]" pageId="6" pageNumber="347">
<emphasis id="B94BEAF4FFF7FFF0FF7AFB2BB410FB40" bold="true" box="[199,334,1194,1219]" pageId="6" pageNumber="347">Etymology.</emphasis>
The species name is given from the name of the
<typeStatus id="54848844FFF7FFF0FCCEFB2AB6FDFB40" box="[883,931,1195,1219]" pageId="6" pageNumber="347">type</typeStatus>
locality, Sulawesi Island,
<collectingCountry id="F3287676FFF7FFF0FB74FB2BB064FB40" box="[1225,1338,1194,1219]" name="Indonesia" pageId="6" pageNumber="347">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C325656DFFF7FFF0FF7AFB4FB0C8FB64" box="[199,1430,1230,1255]" pageId="6" pageNumber="347" type="distribution">
<paragraph id="8B8036E6FFF7FFF0FF7AFB4FB0C8FB64" blockId="6.[151,1437,258,1255]" box="[199,1430,1230,1255]" pageId="6" pageNumber="347">
<emphasis id="B94BEAF4FFF7FFF0FF7AFB4FB74BFB64" bold="true" box="[199,533,1230,1255]" pageId="6" pageNumber="347">Distribution and bionomics.</emphasis>
<collectingCountry id="F3287676FFF7FFF0FDA1FB4FB7D2FB64" box="[540,652,1230,1255]" name="Indonesia" pageId="6" pageNumber="347">Indonesia</collectingCountry>
, Sulawesi Isl. The species was collected in primary forest in October.
</paragraph>
</subSubSection>
</treatment>
</document>