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<mods:title id="D56B435D443D1D3DCB0DB969FE7789F0">Neotropical Agromyzidae (Diptera) of the Mission Géodésique de lÉquateur: Becker (1920) revisited</mods:title>
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<taxonomicName id="4C3F4D44FF89FFCCFF47FF53DAB9FF68" ID-CoL="8BDXS" authority="Becker" authorityName="Becker" box="[151,700,151,178]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="165" phylum="Arthropoda" rank="species" species="nigricornis" subGenus="Cerodontha">
<emphasis id="B94BEAD5FF89FFCCFF47FF53DAB9FF68" bold="true" box="[151,700,151,178]" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFF47FF53D923FF6B" bold="true" box="[151,294,151,177]" italics="true" pageId="8" pageNumber="165">Cerodontha</emphasis>
(
<emphasis id="B94BEAD5FF89FFCCFEE7FF53D9C3FF6B" bold="true" box="[311,454,151,177]" italics="true" pageId="8" pageNumber="165">Cerodontha</emphasis>
)
<emphasis id="B94BEAD5FF89FFCCFE07FF53DA59FF6B" bold="true" box="[471,604,151,177]" italics="true" pageId="8" pageNumber="165">nigricornis</emphasis>
Becker
</emphasis>
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</paragraph>
<paragraph id="8B8036C7FF89FFCCFF47FF7ED92BFF09" blockId="8.[151,700,151,211]" box="[151,302,186,211]" pageId="8" pageNumber="165">
(
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)
</paragraph>
</subSubSection>
<subSubSection id="C325654CFF89FFCCFF47FEC6DC2CFEAD" pageId="8" pageNumber="165" type="reference_group">
<paragraph id="8B8036C7FF89FFCCFF47FEC6DA3FFEC2" blockId="8.[151,833,258,311]" box="[151,570,258,280]" pageId="8" pageNumber="165">
<treatmentCitationGroup id="AB2F11E9FF89FFCCFF47FEC6DA3FFEC2" box="[151,570,258,280]" pageId="8" pageNumber="165">
<taxonomicName id="4C3F4D44FF89FFCCFF47FEC6DA33FEC2" ID-CoL="BDL2T" authority="Becker 1920: 212" authorityName="Becker" authorityPageNumber="212" authorityYear="1920" box="[151,566,258,280]" class="Insecta" family="Agromyzidae" genus="Cerodonta" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="165" phylum="Arthropoda" rank="species" species="nigricornis">
<emphasis id="B94BEAD5FF89FFCCFF47FEC6D97CFEC2" box="[151,377,258,280]" italics="true" pageId="8" pageNumber="165">Cerodonta nigricornis</emphasis>
<treatmentCitation id="0A9E10D6FF89FFCCFEAFFEC6DA33FEC2" author="Becker" box="[383,566,258,280]" page="212" pageId="8" pageNumber="165" year="1920">
<bibRefCitation id="EFAE4B36FF89FFCCFEAFFEC6DA06FEC2" author="Becker" box="[383,515,258,280]" pageId="8" pageNumber="165" refString="Becker, T. (1920) Dipteres brachyceres. In: Mission du Service Geographique de l'Armee pour la mesure d'un arc de Meridien Equatoriel en Amerique du Sud sous le controle scientifique de l'Academie des Sciences, 1899 - 1906. Tome 10. Entomologie. - Botanique. Fascicule 2. - Opiliones. - Dipteres. - Myriapodes. [1919]. Gauthier-Villars, Paris. pp. 163 - 215." type="book chapter" year="1920">Becker 1920</bibRefCitation>
: 212
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8B8036C7FF89FFCCFF47FEE5DB44FEED" blockId="8.[151,833,258,311]" box="[151,833,289,311]" pageId="8" pageNumber="165">
<treatmentCitationGroup id="AB2F11E9FF89FFCCFF47FEE5DB44FEED" box="[151,833,289,311]" pageId="8" pageNumber="165">
<taxonomicName id="4C3F4D44FF89FFCCFF47FEE5DA56FEED" ID-CoL="8BDXS" authority="Spencer 1963: 332" authorityName="Spencer" authorityPageNumber="332" authorityYear="1963" box="[151,595,289,311]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="165" phylum="Arthropoda" rank="species" species="nigricornis">
<emphasis id="B94BEAD5FF89FFCCFF47FEE5D980FEED" box="[151,389,289,311]" italics="true" pageId="8" pageNumber="165">Cerodontha nigricornis</emphasis>
,
<treatmentCitation id="0A9E10D6FF89FFCCFE42FEE5DA56FEED" author="Spencer" box="[402,595,289,311]" page="332" pageId="8" pageNumber="165" year="1963">
<bibRefCitation id="EFAE4B36FF89FFCCFE42FEE5DA25FEED" author="Spencer" box="[402,544,289,311]" pageId="8" pageNumber="165" refString="Spencer, K. A. (1963) A synopsis of the Neotropical Agromyzidae (Diptera). Transactions of the Royal Entomological Society of London, 115, 291 - 389. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1963. tb 00811. x" type="journal article" year="1963">Spencer 1963</bibRefCitation>
: 332
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A9E10D6FF89FFCCFD8DFEE5DB38FEED" author="Sasakawa" box="[605,829,289,311]" page="815" pageId="8" pageNumber="165" year="1992">
<bibRefCitation id="EFAE4B36FF89FFCCFD8DFEE5DB0FFEED" author="Sasakawa" box="[605,778,289,311]" pageId="8" pageNumber="165" refString="Sasakawa, M. (1992 b) The Neotropical Agromyzidae (Diptera). Part 3. New or little-known species from Ecuador. Japanese Journal of Entomology, 60, 815 - 825." type="journal article" year="1992" yearSuffix="b">Sasakawa 1992b</bibRefCitation>
: 815
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8036C7FF89FFCCFF47FE9ADC2CFEAD" blockId="8.[151,1437,350,1890]" box="[151,1065,350,375]" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFF47FE9AD961FEAD" bold="true" box="[151,356,350,375]" pageId="8" pageNumber="165">MNHN specimen</emphasis>
.
<typeStatus id="54848865FF89FFCCFEA3FE9AD9DEFEAD" box="[371,475,350,375]" pageId="8" pageNumber="165" type="holotype">Holotype</typeStatus>
♀:
<collectingCountry id="F3287657FF89FFCCFDD6FE9ADA68FEAD" box="[518,621,350,375]" name="Ecuador" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFDD6FE9ADA68FEAD" bold="true" box="[518,621,350,375]" pageId="8" pageNumber="165">Ecuador</emphasis>
</collectingCountry>
: Casitagua, alt.
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, P. Rivet, 1903.
</paragraph>
</subSubSection>
<subSubSection id="C325654CFF89FFCCFF17FE46DD56FE41" box="[199,1363,386,411]" pageId="8" pageNumber="165" type="materials_examined">
<paragraph id="8B8036C7FF89FFCCFF17FE46DD56FE41" blockId="8.[151,1437,350,1890]" box="[199,1363,386,411]" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFF17FE46DA29FE41" bold="true" box="[199,556,386,411]" pageId="8" pageNumber="165">Additional material examined</emphasis>
.
<emphasis id="B94BEAD5FF89FFCCFDEBFE46DAB1FE41" bold="true" box="[571,692,386,411]" pageId="8" pageNumber="165">
<collectingCountry id="F3287657FF89FFCCFDEBFE46DAB1FE41" box="[571,692,386,411]" name="Argentina" pageId="8" pageNumber="165">Argentina</collectingCountry>
</emphasis>
: Jujuy, La Quiaca,
<date id="FF811007FF89FFCCFC5DFE46DBE1FE41" box="[909,996,386,411]" pageId="8" pageNumber="165" value="1968-10-23">23.x.68</date>
,
<quantity id="4CC79B22FF89FFCCFC3FFE46DC43FE40" box="[1007,1094,386,411]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="165" unit="m" value="3500.0">3500 m</quantity>
, L.E. Pena (
<specimenCount id="9D39FD4EFF89FFCCFB02FE47DCF8FE40" box="[1234,1277,387,411]" pageId="8" pageNumber="165" type="generic">1 ♂</specimenCount>
:
<collectionCode id="ED2EAE02FF89FFCCFAD9FE47DD42FE41" box="[1289,1351,387,411]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="8" pageNumber="165">CNC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C325654CFF89FFCCFF17FE62D8FAFD95" pageId="8" pageNumber="165" type="diagnosis">
<paragraph id="8B8036C7FF89FFCCFF17FE62D8FAFD95" blockId="8.[151,1437,350,1890]" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFF17FE62D944FE65" bold="true" box="[199,321,422,447]" pageId="8" pageNumber="165">Diagnosis.</emphasis>
This species can be distinguished from other Neotropical species of
<taxonomicName id="4C3F4D44FF89FFCCFB83FE62DD72FE65" box="[1107,1399,422,447]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAD5FF89FFCCFB83FE62DD72FE65" box="[1107,1399,422,447]" italics="true" pageId="8" pageNumber="165">Cerodontha (Cerodontha)</emphasis>
</taxonomicName>
by the following combination of characters: the presence of acrostichals; femora yellow only at knees; first flagellomere strongly angulate but without spine; presence of only 1 ors and sometimes 1 weaker ori; orbit with dense patch of white pubescence anteriorly; large size (wing length over 3.0 mm); and the shape of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C325654CFF89FFCCFF17FD9EDD91FA0C" pageId="8" pageNumber="165" type="description">
<paragraph id="8B8036C7FF89FFCCFF17FD9EDA4CFB90" blockId="8.[151,1437,350,1890]" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFF17FD9ED954FDA9" bold="true" box="[199,337,602,627]" pageId="8" pageNumber="165">Description</emphasis>
. Frons width
<quantity id="4CC79B22FF89FFCCFE3EFD9FDA53FDA8" box="[494,598,603,627]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="165" unit="mm" value="0.3">0.30 mm</quantity>
; ratio of frons width to eye width 1.41.5 (measured in dorsal view); orbit 0.220.27 times width of frons (including orbits) at midpoint; parafacial well developed, forming large ring (cheek) below eye; 1 reclinate ors; and 1 fine, pale (almost invisible in male specimen), proclinate and slightly inclinate ori; orbital setulae forming dense white patch of hairs anteriorly (overlooked in original description); first flagellomere elongated, projecting dorsoapically (but without spine), with short pubescence apically; arista slightly shorter than maximum eye height and with short pubescence; gena deep, extended at rear; gena height at midpoint: 0.38 times maximum eye height; upper margin of clypeus narrow, rounded; eye bare or at most with short scattered pubescence. One presutural and three postsutural dorsocentrals; acrostichals few, in 2 rows; prescutellar acrostichal seta absent; scutellum with only one pair of apical setae; notopleuron with 2 setae; postpronotum with 1 strong seta and 1 (in
<typeStatus id="54848865FF89FFCCFED0FC5AD961FC6D" box="[256,356,926,951]" pageId="8" pageNumber="165" type="holotype">holotype</typeStatus>
) to 4 setulae; anepisternum with 1 strong seta on posterior margin a little above midpoint, and no additional seta (
<typeStatus id="54848865FF89FFCCFE97FC06D9A8FC01" box="[327,429,962,987]" pageId="8" pageNumber="165" type="holotype">holotype</typeStatus>
) or 1 small seta (male) underneath; fore femur with 2 strong lateroventral setae near apex, 1 strong lateral seta basally and 1 or 2 strong dorsal setae near apex; fore and mid tibiae without lateral seta; wing length 3.0 mm in male and
<quantity id="4CC79B22FF89FFCCFE18FBCFDA1BFBF8" box="[456,542,1035,1059]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="165" unit="mm" value="3.8">3.8 mm</quantity>
in female; M1+2 ending at wing tip; costa extending to M1+2; last section of CuA1 0.700.77 times length of penultimate.
</paragraph>
<paragraph id="8B8036C7FF89FFCCFF17FB91DA86FAB0" blockId="8.[151,1437,350,1890]" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFF17FB91D91FFBB4" bold="true" box="[199,282,1109,1134]" pageId="8" pageNumber="165">Colour</emphasis>
. Frons completely yellow including most of orbits, except brownish under ors; both vt on brown, but vti at limit of yellow ground; hind margin of eye brown for a short distance beyond vte; face yellow; all antennal segments brown; palpus brown; margin of clypeus brown. Mesonotum and scutellum mat greyish or greyishbrown; postpronotum yellow with central brown patch; postpronotal seta on yellow ground; notopleuron yellow with very small brown marking at anteroventral corner; anepisternum mostly brown excluding yellow posterodorsal corner; katepisternum completely brown; katatergite prominent and yellow; calypter white with margin and fringe pale brown; halter white; legs, including coxae, brown, with knees yellow for about 1/5 of femur length (measurements taken on fore femur).
</paragraph>
<paragraph id="8B8036C7FF89FFCCFF17FAB1DD91FA0C" blockId="8.[151,1437,350,1890]" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFF17FAB1D977FA54" bold="true" box="[199,370,1397,1422]" pageId="8" pageNumber="165">Male genitalia</emphasis>
. Phallus weakly sclerotized, long and straight for most of its length but curving ventrally near apex; distiphallus fused for entire length except narrowly divided behind apex (
<figureCitation id="13042A42FF89FFCCFB96FA5DDC9AFA68" box="[1094,1183,1433,1458]" captionStart="FIGURES 25 27" captionStartId="9.[151,264,1027,1050]" captionTargetBox="[309,1279,204,997]" captionTargetId="figure@9.[300,1286,193,1006]" captionTargetPageId="9" captionText="FIGURES 25 27. Cerodontha nigricornis Becker, (25) phallus, lateral (before dissection), (26) phallus, lateral, (27) phallus, ventral. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/252258/files/figure.png" pageId="8" pageNumber="165">Fig. 27</figureCitation>
); surstylus elongated, projecting ventrally, bearing multiple spines on ventral surface; ejaculatory apodeme small with fan-shaped blade.
</paragraph>
</subSubSection>
<subSubSection id="C325654CFF89FFCCFF17FA25D9C3F8B8" pageId="8" pageNumber="165" type="discussion">
<paragraph id="8B8036C7FF89FFCCFF17FA25D8DBF950" blockId="8.[151,1437,350,1890]" pageId="8" pageNumber="165">
<emphasis id="B94BEAD5FF89FFCCFF17FA25D94DFA20" bold="true" box="[199,328,1505,1530]" pageId="8" pageNumber="165">Comments</emphasis>
. This species was previously known from a single Ecuadorian female specimen (
<bibRefCitation id="EFAE4B36FF89FFCCFAD0FA25DD9DFA20" author="Becker" box="[1280,1432,1505,1530]" pageId="8" pageNumber="165" refString="Becker, T. (1920) Dipteres brachyceres. In: Mission du Service Geographique de l'Armee pour la mesure d'un arc de Meridien Equatoriel en Amerique du Sud sous le controle scientifique de l'Academie des Sciences, 1899 - 1906. Tome 10. Entomologie. - Botanique. Fascicule 2. - Opiliones. - Dipteres. - Myriapodes. [1919]. Gauthier-Villars, Paris. pp. 163 - 215." type="book chapter" year="1920">Becker 1920</bibRefCitation>
;
<bibRefCitation id="EFAE4B36FF89FFCCFF47F9C2D931F9C4" author="Spencer" box="[151,308,1541,1566]" pageId="8" pageNumber="165" refString="Spencer, K. A. (1963) A synopsis of the Neotropical Agromyzidae (Diptera). Transactions of the Royal Entomological Society of London, 115, 291 - 389. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1963. tb 00811. x" type="journal article" year="1963">Spencer 1963</bibRefCitation>
;
<bibRefCitation id="EFAE4B36FF89FFCCFE90F9C1DA05F9C4" author="Sasakawa" box="[320,512,1541,1566]" pageId="8" pageNumber="165" refString="Sasakawa, M. (1992 b) The Neotropical Agromyzidae (Diptera). Part 3. New or little-known species from Ecuador. Japanese Journal of Entomology, 60, 815 - 825." type="journal article" year="1992" yearSuffix="b">Sasakawa 1992b</bibRefCitation>
;
<bibRefCitation id="EFAE4B36FF89FFCCFDDCF9C1DB35F9C4" author="Martinez" box="[524,816,1541,1566]" pageId="8" pageNumber="165" refString="Martinez, M. &amp; Etienne, J. (2002) Liste systematique et biogeographique des Agromyzidae (Diptera) de la region neotropicale. Bolletino di Zoologia agraria e di Bachicoltura, Serie II, 34, 25 - 52." type="journal article" year="2002">Martinez &amp; Etienne 2002</bibRefCitation>
).
<bibRefCitation id="EFAE4B36FF89FFCCFC93F9C1DBE1F9C4" author="Becker" box="[835,996,1541,1566]" pageId="8" pageNumber="165" refString="Becker, T. (1920) Dipteres brachyceres. In: Mission du Service Geographique de l'Armee pour la mesure d'un arc de Meridien Equatoriel en Amerique du Sud sous le controle scientifique de l'Academie des Sciences, 1899 - 1906. Tome 10. Entomologie. - Botanique. Fascicule 2. - Opiliones. - Dipteres. - Myriapodes. [1919]. Gauthier-Villars, Paris. pp. 163 - 215." type="book chapter" year="1920">Becker (1920)</bibRefCitation>
did not explicitly designate a
<typeStatus id="54848865FF89FFCCFAEAF9C1DD99F9C4" box="[1338,1436,1541,1566]" pageId="8" pageNumber="165" type="holotype">holotype</typeStatus>
in the original publication and the specimen does not bear a
<typeStatus id="54848865FF89FFCCFC9EF9EEDB7BF998" box="[846,894,1578,1602]" pageId="8" pageNumber="165">type</typeStatus>
label, but because no other
<typeStatus id="54848865FF89FFCCFB13F9EEDCF6F998" box="[1219,1267,1578,1602]" pageId="8" pageNumber="165">type</typeStatus>
specimens are known to exist, we consider this specimen to be the
<typeStatus id="54848865FF89FFCCFD20F989DB50F9BC" box="[752,853,1613,1638]" pageId="8" pageNumber="165" type="holotype">holotype</typeStatus>
, fixed by monotypy. We have added a label to that effect.
</paragraph>
<paragraph id="8B8036C7FF89FFCCFF17F952D9C3F8B8" blockId="8.[151,1437,350,1890]" pageId="8" pageNumber="165">
One additional specimen was examined from
<collectingCountry id="F3287657FF89FFCCFD15F951DB30F974" box="[709,821,1685,1710]" name="Argentina" pageId="8" pageNumber="165">Argentina</collectingCountry>
and represents the first documented male. This species is most similar to
<taxonomicName id="4C3F4D44FF89FFCCFEB1F97DDAE1F908" authority="Spencer" authorityName="Spencer" box="[353,740,1721,1746]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="165" phylum="Arthropoda" rank="species" species="chilensis" subGenus="Cerodontha">
<emphasis id="B94BEAD5FF89FFCCFEB1F97DD9E3F908" box="[353,486,1721,1746]" italics="true" pageId="8" pageNumber="165">Cerodontha</emphasis>
(
<emphasis id="B94BEAD5FF89FFCCFE26F97EDA15F90B" box="[502,528,1722,1745]" italics="true" pageId="8" pageNumber="165">C.</emphasis>
)
<emphasis id="B94BEAD5FF89FFCCFDF1F97DDA86F908" box="[545,643,1721,1746]" italics="true" pageId="8" pageNumber="165">chilensis</emphasis>
Spencer
</taxonomicName>
and
<taxonomicName id="4C3F4D44FF89FFCCFCCBF97DDCABF908" authority="Spencer" authorityName="Spencer" box="[795,1198,1721,1746]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="165" phylum="Arthropoda" rank="species" species="magellani" subGenus="Cerodontha">
<emphasis id="B94BEAD5FF89FFCCFCCBF97DDBA5F908" box="[795,928,1721,1746]" italics="true" pageId="8" pageNumber="165">Cerodontha</emphasis>
(
<emphasis id="B94BEAD5FF89FFCCFC61F97EDBCFF90B" box="[945,970,1722,1745]" italics="true" pageId="8" pageNumber="165">C.</emphasis>
)
<emphasis id="B94BEAD5FF89FFCCFC0BF97DDC4EF908" box="[987,1099,1721,1746]" italics="true" pageId="8" pageNumber="165">magellani</emphasis>
Spencer
</taxonomicName>
, both described from
<collectingCountry id="F3287657FF89FFCCFF47F919D8D1F92C" box="[151,212,1757,1782]" name="Chile" pageId="8" pageNumber="165">Chile</collectingCountry>
(
<bibRefCitation id="EFAE4B36FF89FFCCFF32F91AD985F92C" author="Spencer" box="[226,384,1758,1782]" pageId="8" pageNumber="165" refString="Spencer, K. A. (1982) Agromyzidae (Diptera) in Chile. Stuttgarter Beitrage zur Naturkunde. Serie A (Biologie), 357, 1 - 55." type="journal article" year="1982">Spencer 1982</bibRefCitation>
), but differs in having a characteristic white patch of proclinate orbital setulae, larger size, and a slightly different curvature of the phallus. Additional material agreeing with the description of this species has also been collected by M. v. Tschirnhaus in the high Andes (
<quantity id="4CC79B22FF89FFCCFC80F8E1DBF5F8E7" box="[848,1008,1829,1854]" metricMagnitude="3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="165" unit="m" value="4500.0" valueMax="5000.0" valueMin="4000.0">40005000 m</quantity>
elevation) of northern
<collectingCountry id="F3287657FF89FFCCFB2AF8E1DD33F8E4" box="[1274,1334,1829,1854]" name="Chile" pageId="8" pageNumber="165">Chile</collectingCountry>
(M. von Tschirnhaus, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>