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<mods:title id="D1A49563EA94C4D441DE0A01B71A5761">Fossil Darwin wasps (Hymenoptera: Ichneumonidae) from Baltic amber</mods:title>
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<mods:affiliation id="166B2308823C4796E59E46E73B833BD1">Kaliningrad Amber Museum, Marshal Vasilevskiy Square 1, Kaliningrad 236016, Russia</mods:affiliation>
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<mods:affiliation id="3D0215DD119381A2D46936710E6A18E9">Lomonosov Moscow State University, Leninskiye Gory, 1, GSP- 1, Moscow 119991, Russia</mods:affiliation>
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Subfamily
<taxonomicName id="C0DD5DD1FF9AFFF7FE89FB9DFD93FE2E" authority="Kirby, 1837" authorityName="Kirby" authorityYear="1837" box="[266,534,477,503]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="639" phylum="Arthropoda" rank="subFamily" superFamily="Cryptinae">Cryptinae Kirby, 1837</taxonomicName>
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<taxonomicName id="C0DD5DD1FF9AFFF7FF0BF864FF73FDE7" authorityName="Kirby" authorityYear="1837" box="[136,246,548,574]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="639" phylum="Arthropoda" rank="subFamily" subFamily="Cryptinae">Cryptinae</taxonomicName>
is the largest subfamily of
<taxonomicName id="C0DD5DD1FF9AFFF7FDD5F864FC83FDE7" ID-CoL="BCL" authorityName="Latreille" authorityYear="1802" box="[598,774,548,574]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="639" phylum="Arthropoda" rank="family">Ichneumonidae</taxonomicName>
, about 3,100 species have been described, in 275 genera (
<bibRefCitation id="634C5BA3FF9AFFF7FF13F82BFEDAFD5C" author="Broad, G. R. &amp; Shaw, M. R. &amp; Fitton, M. G." box="[144,351,619,646]" pageId="2" pageNumber="639" pagination="411" refId="ref5960" refString="Broad, G. R., Shaw, M. R. &amp; Fitton, M. G. (2018) Ichneumonid wasps (Hymenoptera: Ichneumonidae): their classification and biology. Handbooks for the identification of British Insects, 7 (12), 411 рр." type="journal article" year="2018">
Broad
<emphasis id="35A9FA40FF9AFFF7FF5FF82CFE88FD5C" box="[220,269,619,645]" italics="true" pageId="2" pageNumber="639">et al</emphasis>
., 2018
</bibRefCitation>
). Mostly species are external parasites of insect pupae and cocoons. The earliest
<taxonomicName id="C0DD5DD1FF9AFFF7FDECF8CEFD58FD71" authorityName="Kirby" authorityYear="1837" box="[623,733,654,680]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="639" phylum="Arthropoda" rank="subFamily" subFamily="Cryptinae">Cryptinae</taxonomicName>
are recorded in the Early Eocene (Green River Formation) (
<bibRefCitation id="634C5BA3FF9AFFF7FF0CF895FE21FD29" author="Spasojevic, T. &amp; Wedmann, S. &amp; Klopfstein, S." box="[143,420,725,752]" pageId="2" pageNumber="639" pagination="0197477" refId="ref7941" refString="Spasojevic, T., Wedmann, S. &amp; Klopfstein, S. (2018 b) Seven remarkable new fossil species of parasitoid wasps (Hymenoptera, Ichneumonidae) from the Eocene Messel Pit. PLoS ONE, 13 (6), e 0197477. https: // doi. org / 10.1371 / journal. pone. 0197477" type="journal article" year="2018">
Spasojevic
<emphasis id="35A9FA40FF9AFFF7FE91F896FEC6FD36" box="[274,323,725,751]" italics="true" pageId="2" pageNumber="639">et al</emphasis>
., 2018b
</bibRefCitation>
). A high occurrence and species diversity were noted in the Beds of the Bembridge marls (
<bibRefCitation id="634C5BA3FF9AFFF7FF13F95CFEC1FCEF" author="Khalaim, A. I." box="[144,324,796,822]" pageId="2" pageNumber="639" pagination="344 - 360" refId="ref6631" refString="Khalaim, A. I. (2014) Family Ichneumonidae Latreille, 1802. In: Antropov, A. V., Belokobylskij, S. A., Compton, S. G., Dlussky, G. M., Khalaim, A. I., Kolyada, V. A., Kozlov, M. A., Perfilieva, K. S. &amp; Rasnitsyn, A. P. (2014) The wasps, bees and ants (Insecta: Vespida = Hymenoptera) from the Insect Limestone (Late Eocene) of the Isle of Wight, UK. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 104, 344 - 360. https: // doi. org / 10.1017 / S 1755691014000103" type="journal article" year="2014">Khalaim, 2014</bibRefCitation>
), Florissant (
<bibRefCitation id="634C5BA3FF9AFFF7FE67F95CFDF2FCEF" author="Brues, C. T." box="[484,631,796,822]" pageId="2" pageNumber="639" pagination="491 - 498" refId="ref6012" refString="Brues, C. T. (1906) Fossil parasitic and phytophagous Hymenoptera from Florissant, Colorado. Bulletin of the American Museum of Natural History, 22, 491 - 498." type="journal article" year="1906">Brues, 1906</bibRefCitation>
,
<bibRefCitation id="634C5BA3FF9AFFF7FD09F95CFD43FCEF" author="Brues, C. T." box="[650,710,796,822]" pageId="2" pageNumber="639" pagination="1 - 125" refId="ref6046" refString="Brues, C. T. (1910) The parasitic Hymenoptera of the Tertiary of Florissant, Colorado. Bulletin of the Museum of Comparative Zoology, 54 (1), 1 - 125." type="journal article" year="1910">1910</bibRefCitation>
), and Biamo (
<bibRefCitation id="634C5BA3FF9AFFF7FF65F97FFE12FC80" author="Khalaim, A. I." box="[230,407,831,857]" pageId="2" pageNumber="639" pagination="101 - 112" refId="ref6541" refString="Khalaim, A. I. (2008) Fossil ichneumon wasps (Hymenoptera: Ichneumonidae) from Biamo (Russia), Oligocene. Alavesia, 2, 101 - 112." type="journal article" year="2008">Khalaim, 2008</bibRefCitation>
). However, they are rare in the Early Eocene Tadushi Formation (
<bibRefCitation id="634C5BA3FF9AFFF7FD81F922FF40FC79" author="Tolkanitz, V. I. &amp; Perkovsky, E. E." pageId="2" pageNumber="639" pagination="31 - 34" refId="ref8148" refString="Tolkanitz, V. I. &amp; Perkovsky, E. E. (2018) First record of the late Eocene ichneumon fly Rasnitsynites tarsalis Kasparyan (Ichneumonidae, Townesitinae) in Ukraine confirms correlation of the upper Eocene Lagerstatten. Paleontological Journal, 52 (11), 31 - 34. https: // doi. org / 10.1134 / S 0031030118010136" type="journal article" year="2018">Tolkanitz &amp; Perkovsky, 2018</bibRefCitation>
). The subfamily dominates in inclusions of Baltic amber in terms of abundance, constituting more than 44% of the studied samples. The fossil fauna of Baltic amber is completely unrevised and not a single species has been formally described.
</paragraph>
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