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<document id="A4CCFA26D639338F0A245B6384A95F6A" ID-DOI="10.5281/zenodo.202575" ID-GBIF-Dataset="e6fc0416-99c9-495a-af89-35115e9a1675" ID-ISSN="1175-5326" ID-Zenodo-Dep="202575" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460302833884" checkinUser="plazi" docAuthor="Pietrykowska-Tudruj, Ewa &amp; Staniec, Bernard" docDate="2011" docId="7E75879EFF93FFE46ACC004D0B1EFC19" docLanguage="en" docName="zt02865p067.pdf" docOrigin="Zootaxa 2865" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Philonthus debilis Gravenhorst" docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="824CFFE6FF94FFE36A5B01220E7FFFC9" masterDocTitle="The morphology of the pupae of six species of Philonthina (Coleoptera, Staphylinidae, Staphylinini) with taxonomic remarks" masterLastPageNumber="67" masterPageNumber="53" pageNumber="60" updateTime="1698252825721" updateUser="plazi">
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<mods:title id="4EFF163C6EE3671C7E8BF82069805881">The morphology of the pupae of six species of Philonthina (Coleoptera, Staphylinidae, Staphylinini) with taxonomic remarks</mods:title>
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<mods:namePart id="50BA19B7D3FB39FEF4D13F0F1240C86C">Pietrykowska-Tudruj, Ewa</mods:namePart>
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<mods:namePart id="B360BD50054005FB66362928E592310F">Staniec, Bernard</mods:namePart>
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<mods:date id="D0F91D98A65B9F9E418DBD9D62B02783">2011</mods:date>
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<treatment id="7E75879EFF93FFE46ACC004D0B1EFC19" ID-DOI="http://doi.org/10.5281/zenodo.6183493" ID-GBIF-Taxon="119400037" ID-Zenodo-Dep="6183493" LSID="urn:lsid:plazi:treatment:7E75879EFF93FFE46ACC004D0B1EFC19" httpUri="http://treatment.plazi.org/id/7E75879EFF93FFE46ACC004D0B1EFC19" lastPageNumber="60" pageId="7" pageNumber="60">
<subSubSection id="BEC66503FF93FFE46ACC004D0B1EFC19" pageId="7" pageNumber="60" type="nomenclature">
<paragraph id="F6633688FF93FFE46ACC004D0C65FE41" blockId="7.[151,659,367,428]" box="[151,538,367,392]" pageId="7" pageNumber="60">
<heading id="AD2B81E4FF93FFE46ACC004D0C65FE41" bold="true" box="[151,538,367,392]" fontSize="10" level="2" pageId="7" pageNumber="60" reason="2">
<taxonomicName id="31DC4D0BFF93FFE46ACC004D0C65FE41" ID-CoL="4G9TS" authority="Gravenhorst" authorityName="Gravenhorst" box="[151,538,367,392]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="C4A8EA9AFF93FFE46ACC004D0C65FE41" bold="true" box="[151,538,367,392]" pageId="7" pageNumber="60">
<emphasis id="C4A8EA9AFF93FFE46ACC004D0F17FE41" bold="true" box="[151,360,367,392]" italics="true" pageId="7" pageNumber="60">Philonthus debilis</emphasis>
(Gravenhorst)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F6633688FF93FFE46ACC00B60CECFE65" blockId="7.[151,659,367,428]" box="[151,659,404,428]" pageId="7" pageNumber="60">
((
<figureCitation id="6EE72A0DFF93FFE46AF200B60F56FE65" box="[169,297,404,428]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1969,1992]" captionTargetBox="[217,1335,194,1938]" captionTargetId="figure@8.[209,1368,193,1939]" captionTargetPageId="8" captionText="FIGURES 13 18. Pupa of Philonthus debilis (13 15) and P. decorus (16 18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect." httpUri="https://zenodo.org/record/202578/files/figure.png" pageId="7" pageNumber="60">Figs 1315</figureCitation>
,
<figureCitation id="6EE72A0DFF93FFE46B6E00B60F02FE65" box="[309,381,404,428]" captionStart="FIGURES 19 40" captionStartId="9.[151,264,1885,1908]" captionTargetBox="[232,1328,193,1860]" captionTargetId="figure@9.[216,1345,193,1863]" captionTargetPageId="9" captionText="FIGURES 19 40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21 25), P. carbonarius (26 32), P. debilis (33 35) and P. decorus (36 40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19 A, 26 A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39 A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34 A) and spine (32, 35, 38)." httpUri="https://zenodo.org/record/202579/files/figure.png" pageId="7" pageNumber="60">3335</figureCitation>
, 4949B, 56, 60, 6870)
</paragraph>
<paragraph id="F6633688FF93FFE46ACC00FE0B1EFC19" blockId="7.[151,1436,476,976]" pageId="7" pageNumber="60">
Body length:
<quantity id="31249B6DFF93FFE46B6A00FE0FA7FE3D" box="[305,472,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="7" pageNumber="60" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.203.30 mm</quantity>
; body width:
<quantity id="31249B6DFF93FFE4682F00FE0D64FE3D" box="[628,795,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.365" metricValueMax="1.38" metricValueMin="1.35" pageId="7" pageNumber="60" unit="mm" value="1.365" valueMax="1.38" valueMin="1.35">1.351.38 mm</quantity>
; head width:
<quantity id="31249B6DFF93FFE469EF00FE0A24FE3D" box="[948,1115,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="7.15" metricValueMax="7.7" metricValueMin="6.6" pageId="7" pageNumber="60" unit="mm" value="0.715" valueMax="0.77" valueMin="0.66">0.660.77 mm</quantity>
; pronotum width:
<quantity id="31249B6DFF93FFE46F7700FE0EB9FDDE" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="9.8" metricValueMin="9.0" pageId="7" pageNumber="60" unit="mm" value="0.94" valueMax="0.98" valueMin="0.9">0.900.98 mm</quantity>
; 1.1 x as broad as long; body rather slender and relatively weakly sclerotized. Head almost as long as wide (
<figureCitation id="6EE72A0DFF93FFE46AC403060F5BFDF5" box="[159,292,548,572]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1969,1992]" captionTargetBox="[217,1335,194,1938]" captionTargetId="figure@8.[209,1368,193,1939]" captionTargetPageId="8" captionText="FIGURES 13 18. Pupa of Philonthus debilis (13 15) and P. decorus (16 18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect." httpUri="https://zenodo.org/record/202578/files/figure.png" pageId="7" pageNumber="60">Figs 13, 14</figureCitation>
). Antennae reaching to middle of elytra length as in
<taxonomicName id="31DC4D0BFF93FFE469D803070A69FDF5" box="[899,1046,548,572]" class="Insecta" family="Staphylinidae" genus="Bisnius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="fimetarius">
<emphasis id="C4A8EA9AFF93FFE469D803070A69FDF5" box="[899,1046,548,572]" italics="true" pageId="7" pageNumber="60">B. fimetarius</emphasis>
</taxonomicName>
(
<figureCitation id="6EE72A0DFF93FFE46E7C03060A0EFDF5" box="[1063,1137,548,572]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1962,1985]" captionTargetBox="[201,1371,187,1936]" captionTargetId="figure@4.[199,1376,187,1937]" captionTargetPageId="4" captionText="FIGURES 1 6. Pupa of Bisnius fimentarius (1 3) and Gabrius osseticus (4 6). (1, 4) Ventral aspect, (2, 5) lateral aspect, (3, 6) dorsal aspect." httpUri="https://zenodo.org/record/202576/files/figure.png" pageId="7" pageNumber="60">Figs 2</figureCitation>
,
<figureCitation id="6EE72A0DFF93FFE46E2403060AE1FDF5" box="[1151,1182,548,572]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1969,1992]" captionTargetBox="[217,1335,194,1938]" captionTargetId="figure@8.[209,1368,193,1939]" captionTargetPageId="8" captionText="FIGURES 13 18. Pupa of Philonthus debilis (13 15) and P. decorus (16 18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect." httpUri="https://zenodo.org/record/202578/files/figure.png" pageId="7" pageNumber="60">14</figureCitation>
). Margin of pronotum with 8 or 10 projections (5/5, 4/4, projections on sides in individuals). Wings slightly protruding on the posterior margin of 1st (morphologically 3rd) clearly visible, abdominal segment (
<figureCitation id="6EE72A0DFF93FFE46998034D0A37FD4E" box="[963,1096,623,647]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1969,1992]" captionTargetBox="[217,1335,194,1938]" captionTargetId="figure@8.[209,1368,193,1939]" captionTargetPageId="8" captionText="FIGURES 13 18. Pupa of Philonthus debilis (13 15) and P. decorus (16 18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect." httpUri="https://zenodo.org/record/202578/files/figure.png" pageId="7" pageNumber="60">Figs 13, 14</figureCitation>
). Each fore, middle, and hind tibiae with 45, 9, and 4 protuberances respectively. Tarsi of hind legs reaching or protruding the middle of 4th (morphologically 6th) clearly visible abdominal segment length (
<figureCitation id="6EE72A0DFF93FFE4692E039E0D84FD1D" box="[885,1019,700,724]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1969,1992]" captionTargetBox="[217,1335,194,1938]" captionTargetId="figure@8.[209,1368,193,1939]" captionTargetPageId="8" captionText="FIGURES 13 18. Pupa of Philonthus debilis (13 15) and P. decorus (16 18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect." httpUri="https://zenodo.org/record/202578/files/figure.png" pageId="7" pageNumber="60">Figs 13, 14</figureCitation>
). Abdominal segments IIIV almost equal in width, then narrowed to the terminal segment of the body (
<figureCitation id="6EE72A0DFF93FFE469F903C20D88FD31" box="[930,1015,736,760]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1969,1992]" captionTargetBox="[217,1335,194,1938]" captionTargetId="figure@8.[209,1368,193,1939]" captionTargetPageId="8" captionText="FIGURES 13 18. Pupa of Philonthus debilis (13 15) and P. decorus (16 18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect." httpUri="https://zenodo.org/record/202578/files/figure.png" pageId="7" pageNumber="60">Fig. 15</figureCitation>
). Abdominal segments IIIVIII each with a pair of setiform projections (6 pairs in total) on sides (
<figureCitation id="6EE72A0DFF93FFE4690A02260DABFCD5" box="[849,980,772,796]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1969,1992]" captionTargetBox="[217,1335,194,1938]" captionTargetId="figure@8.[209,1368,193,1939]" captionTargetPageId="8" captionText="FIGURES 13 18. Pupa of Philonthus debilis (13 15) and P. decorus (16 18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect." httpUri="https://zenodo.org/record/202578/files/figure.png" pageId="7" pageNumber="60">Figs 1315</figureCitation>
). Segments IIIVII 1.21.4 x as long as projections (
<figureCitation id="6EE72A0DFF93FFE46B79020A0FDBFC89" box="[290,420,808,832]" captionStart="FIGURES 13 18" captionStartId="8.[151,264,1969,1992]" captionTargetBox="[217,1335,194,1938]" captionTargetId="figure@8.[209,1368,193,1939]" captionTargetPageId="8" captionText="FIGURES 13 18. Pupa of Philonthus debilis (13 15) and P. decorus (16 18). (13, 16) Ventral aspect, (14, 17) lateral aspect, (15, 18) dorsal aspect." httpUri="https://zenodo.org/record/202578/files/figure.png" pageId="7" pageNumber="60">Figs 1315</figureCitation>
); projections of segment VIII usually curved, at least as long as segment (
<figureCitation id="6EE72A0DFF93FFE46EBA020A0B1BFC89" box="[1249,1380,808,832]" captionStart="FIGURES 19 40" captionStartId="9.[151,264,1885,1908]" captionTargetBox="[232,1328,193,1860]" captionTargetId="figure@9.[216,1345,193,1863]" captionTargetPageId="9" captionText="FIGURES 19 40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21 25), P. carbonarius (26 32), P. debilis (33 35) and P. decorus (36 40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19 A, 26 A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39 A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34 A) and spine (32, 35, 38)." httpUri="https://zenodo.org/record/202579/files/figure.png" pageId="7" pageNumber="60">Figs 33, 34</figureCitation>
). All abdominal projections on sides bearing sparse cuticular processes (
<figureCitation id="6EE72A0DFF93FFE469DD026E0DA6FCAD" box="[902,985,844,868]" captionStart="FIGURES 19 40" captionStartId="9.[151,264,1885,1908]" captionTargetBox="[232,1328,193,1860]" captionTargetId="figure@9.[216,1345,193,1863]" captionTargetPageId="9" captionText="FIGURES 19 40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21 25), P. carbonarius (26 32), P. debilis (33 35) and P. decorus (36 40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19 A, 26 A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39 A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34 A) and spine (32, 35, 38)." httpUri="https://zenodo.org/record/202579/files/figure.png" pageId="7" pageNumber="60">Fig. 35</figureCitation>
). Abdominal segments with microstructure as in
<figureCitation id="6EE72A0DFF93FFE46B5F02520F2AFC41" box="[260,341,880,904]" captionStart="FIGURES 19 40" captionStartId="9.[151,264,1885,1908]" captionTargetBox="[232,1328,193,1860]" captionTargetId="figure@9.[216,1345,193,1863]" captionTargetPageId="9" captionText="FIGURES 19 40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21 25), P. carbonarius (26 32), P. debilis (33 35) and P. decorus (36 40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19 A, 26 A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39 A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34 A) and spine (32, 35, 38)." httpUri="https://zenodo.org/record/202579/files/figure.png" pageId="7" pageNumber="60">Fig. 34</figureCitation>
A. Female pupa with two well-developed ventral, elongated, straight prolongations (Fig. 49); ventral and terminal prolongation with tiny cuticular processes (Figs 49A, 49B). The first pair of functional spiracles distinctly narrowed apically (
<figureCitation id="6EE72A0DFF93FFE46B84029A0C21FC19" box="[479,606,952,976]" captionStart="FIGURES 52 70" captionStartId="11.[151,264,1916,1939]" captionTargetBox="[257,1311,181,1889]" captionTargetId="figure@11.[256,1312,181,1890]" captionTargetPageId="11" captionText="FIGURES 52 70. Pupa of Bisnius fimentarius (52, 61, 62), Gabrius osseticus (53, 58, 63, 64), Philonthus atratus (54, 59, 65), P. carbonarius (55, 66), P. debilis (56, 60, 68 70) and P. d e c o r u s (57, 67). (52 60) The first pair of functional spiracles; (61 70) atrophied spiracles." httpUri="https://zenodo.org/record/202580/files/figure.png" pageId="7" pageNumber="60">Figs 56, 60</figureCitation>
) with reticulate microstructure; atrophied spiracles as in
<figureCitation id="6EE72A0DFF93FFE46E86029A0B21FC19" box="[1245,1374,952,976]" captionStart="FIGURES 52 70" captionStartId="11.[151,264,1916,1939]" captionTargetBox="[257,1311,181,1889]" captionTargetId="figure@11.[256,1312,181,1890]" captionTargetPageId="11" captionText="FIGURES 52 70. Pupa of Bisnius fimentarius (52, 61, 62), Gabrius osseticus (53, 58, 63, 64), Philonthus atratus (54, 59, 65), P. carbonarius (55, 66), P. debilis (56, 60, 68 70) and P. d e c o r u s (57, 67). (52 60) The first pair of functional spiracles; (61 70) atrophied spiracles." httpUri="https://zenodo.org/record/202580/files/figure.png" pageId="7" pageNumber="60">Figs 6870</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>