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<mods:title id="792E506615FD988434B7B4093A7F6E64">A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World</mods:title>
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<treatment id="7E1B4F4519AEFFB963774B4B4BF074E0" ID-GBIF-Taxon="156200921" LSID="urn:lsid:plazi:treatment:7E1B4F4519AEFFB963774B4B4BF074E0" httpUri="http://treatment.plazi.org/id/7E1B4F4519AEFFB963774B4B4BF074E0" lastPageId="28" lastPageNumber="103" pageId="26" pageNumber="101">
<subSubSection id="A2DCB081A84DA40A78F0ADE124F7C979" pageId="26" pageNumber="101" type="nomenclature">
<paragraph id="29F9ED8FE122298E2F5044155D7B7F2D" pageId="26" pageNumber="101">
<taxonomicName id="D61C551CDF2B44F68D258F01BD1AA007" ID-CoL="62PRG" ID-ENA="2448399" authority="Barr, 1962" authorityName="Barr" authorityYear="1962" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="genus">Cymatoderella Barr, 1962</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8FF4470DA459F84BA4CED2822D767183" pageId="26" pageNumber="101" type="type species">
<paragraph id="453E7E97E6AC51B75AC09EED6F046992" pageId="26" pageNumber="101">Type species.</paragraph>
<paragraph id="62ACF3915EA966C7BE869789B3CC2C34" pageId="26" pageNumber="101">
<taxonomicName id="7FCD4442C076243F0030910FC75026CE" class="Insecta" family="Cleridae" genus="Tillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tillus collaris" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="species" species="collaris">Tillus collaris</taxonomicName>
Spinola, 1844, original designation.
</paragraph>
</subSubSection>
<subSubSection id="D6B19A9E001BF3924DCF3A468C6B5602" pageId="26" pageNumber="101" type="distribution">
<paragraph id="09CF37083EFBBAC1FD3D073D7AD33AA0" pageId="26" pageNumber="101">Distribution.</paragraph>
<paragraph id="E7215006D633D17F8A55C441A20F4353" pageId="26" pageNumber="101">Shown in Fig. 21H.</paragraph>
</subSubSection>
<subSubSection id="F42F6C00EBBFB07AC66FD6B7FA11FDBD" pageId="26" pageNumber="101" type="differential diagnosis">
<paragraph id="449A009E96F79EADF94482178F6DC32E" pageId="26" pageNumber="101">Differential diagnosis.</paragraph>
<paragraph id="23E8F60FB369635EC45762A447E82733" pageId="26" pageNumber="101">
<taxonomicName id="04D2D5824D67B17BF86653D53557680D" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
is most similar to various
<taxonomicName id="64C328A6E7272E16F14D31B9AF8FC69B" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="genus">Cymatodera</taxonomicName>
species of moderate dimensions. The two genera can be recognized based on the size of the ommatidia.
<taxonomicName id="8B1AB9936C0D85603804772F2C962ED7" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
species have the diameter of the ommatidia somewhat small (Fig. 6F) compared to
<taxonomicName id="6BAB6B2D66BA0CCC2ABFD6D416809E35" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="genus">Cymatodera</taxonomicName>
species (Fig. 12A). Additionally, the bicolored composition of the integument in
<taxonomicName id="4BE20A74FE232F471290777D0CC86ABD" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
, with a testaceous to ferrugineous coloration on the head and pronotum and a piceous tone on the rest of the body (Fig. 3
<normalizedToken id="E477059BDD2DA30A393ADB1F53CCD37E" originalValue="BD">B-D</normalizedToken>
) will serve to separate these genera.
<taxonomicName id="68359E454029B711451ABA332F1E2732" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera bicolor" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="species" species="bicolor">Cymatodera bicolor</taxonomicName>
(Say) is the only species in the genus with a similar color pattern, but it has an elongate and narrow body shape (Fig. 5E), not a robust one, as observed in
<taxonomicName id="6432164E5CFE18CFB63B4AF42B9AE21E" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="26" pageNumber="101" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
(Fig. 3
<normalizedToken id="533863093EAF93B49DFFD421FAB2D663" originalValue="BD">B-D</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection id="59637EE95739574B06FEA563BA51FEFE" lastPageId="27" lastPageNumber="102" pageId="26" pageNumber="101" type="redescription">
<paragraph id="2A6D448F9277F6A6913F3C5E5DD33951" pageId="26" pageNumber="101">Redescription.</paragraph>
<paragraph id="BD1351F101CF138CB8FE600F00C09F13" lastPageId="27" lastPageNumber="102" pageId="26" pageNumber="101">
Size: 3-7 mm. Body: Small, relatively robust individuals. Color: Pronotum bicolored, testaceous to ferruginous in the median region and piceous on
<pageBreakToken id="B7345412FE0BFF19156A682D644DD823" pageId="27" pageNumber="102" start="start">the</pageBreakToken>
margins to uniformly testaceous to ferruginous; legs, antennae, thorax, elytra and abdominal segments piceous; head and mouthparts can be testaceous, ferruginous, or with an array of piceous tones; for
<taxonomicName id="029AA1FB21603880E52ABAA05447E8D1" lsidName="C. patagoniae" pageId="27" pageNumber="102" rank="species" species="patagoniae">C. patagoniae</taxonomicName>
, visible ventrites 4-6 can be testaceous to ferruginous. Form: Small sized individuals, body short, robust, elytra subparallel to moderately expanded posteriorly.
</paragraph>
<paragraph id="3BA5E14551B839EAF7FD5E5F78CCAD37" pageId="27" pageNumber="102">
Head: Eyes medium sized, taller than wide, bulging laterally, emarginate (Fig. 6F); sculpturing variously impressed; vestiture variable; antennal insertion located in front of emargination; clypeus emarginate medially; antennae with 11 antennomeres; sexual dimorphism slightly difficult to observe in the last abdominal segment; terminal maxillary palpi cylindrical; terminal labial palpi securiform (Fig. 3
<normalizedToken id="29919D8ABFF3FC5856044B6C9D67DE45" originalValue="BD">B-D</normalizedToken>
).
</paragraph>
<paragraph id="22CC3CBAE3A7436F0EB92F36EB6B2966" pageId="27" pageNumber="102">Thorax: Pronotum narrower than elytral base; widest at middle; sides constricted subapically; more strongly constricted behind middle; disc convex; anterior depression feebly indicated; antescutelar impression absent; posterior margin conspicuously constricted transversally. Prosternum smooth, variously punctate and vested. Mesoventrite wider than long; shiny, variously punctate. Metaventrite convex, smooth, shiny, moderately clothed.</paragraph>
<paragraph id="EE938D17216EA88AA3F931F9E72FF4DF" pageId="27" pageNumber="102">Legs: Femora swollen; tibia slender; rugose to rugulose; tibial spur formula 2-2-2, pulvillar formula 4-4-4.</paragraph>
<paragraph id="EB411DADBD8708ACA688D438FA121053" pageId="27" pageNumber="102">Elytra: Broad; robust; gradually expanded behind middle; humeri strongly indicated; elongate; surface convex, expanded behind middle; moderately to coarsely sculptured; sculpturing arranged in regular striae; elytral declivity somewhat steep; epipleural fold complete, narrowing toward apex; pygidium concealed in dorsal view.</paragraph>
<paragraph id="96318486A23812E8A1368A830354B66E" pageId="27" pageNumber="102">
Abdomen: Six visible ventrites. First visible segment shiny; smooth; 1.5
<normalizedToken id="C848D908843401DC7410965FBF13DE3B" originalValue="×">x</normalizedToken>
longer than remaining segments. Ventrites 2-4 subquadrate; smooth; shiny; variously impressed and clothed; lateral margins parallel; posterior margins truncate. Fifth visible ventrite subquadrate; variously vested; lateral margins oblique; posterior margin truncate. Sixth visible ventrite subtriangular, displaying a degree of sexual dimorphism; lateral margins strongly oblique; posterior margin rounded to moderately emarginate. Fifth tergite subquadrate; posterior margin truncate. Sixth ventrite subtriangular.
</paragraph>
</subSubSection>
<subSubSection id="EC02F7A33FDA38E060C06B1D7184F95A" pageId="27" pageNumber="102" type="remarks">
<paragraph id="F85CA95830DF371807A3104A4AC4FE1D" pageId="27" pageNumber="102">Remarks.</paragraph>
<paragraph id="102D3CD22C9993AA190BF347185816E6" pageId="27" pageNumber="102">
<taxonomicName id="5F70A3C281CC745E43B3B0E50E753D33" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="27" pageNumber="102" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
was established by
<bibRefCitation id="AAB7152F06982809030CA8383898A125" author="Barr, WF" journalOrPublisher="Coleopterists Bulletin" pageId="74" pageNumber="149" pagination="121 - 127" title="A key to the genera and a classification of the North American Cleridae (Coleoptera)." volume="16" year="1962">Barr (1962)</bibRefCitation>
to separate
<taxonomicName id="B31D2A72B8AA0756E56691CE91CF7457" class="Insecta" family="Cleridae" genus="Tillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tillus collaris" order="Coleoptera" pageId="27" pageNumber="102" phylum="Arthropoda" rank="species" species="collaris">Tillus collaris</taxonomicName>
Spinola and
<taxonomicName id="1D6B304E5C91BA67DB6C6720C11B5427" lsidName="T. patagoniae" pageId="27" pageNumber="102" rank="species" species="patagoniae">T. patagoniae</taxonomicName>
Knull, two New World species, from
<taxonomicName id="179043C0D8D8EEF88C5030DBCBC2730D" class="Insecta" family="Cleridae" genus="Tillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tillus" order="Coleoptera" pageId="27" pageNumber="102" phylum="Arthropoda" rank="genus">Tillus</taxonomicName>
(Olivier), a widely distributed genus with a concentration of species in Africa and Oceania. Later on,
<bibRefCitation id="D4E795D7ADEC65960AD3C2B9C6E97862" author="Rifkind, J" journalOrPublisher="The Coleopterists Bulletin" pageId="77" pageNumber="152" pagination="279 - 284" title="A New Species of Cymatoderella Barr (Coleoptera: Cleridae) from Mexico and Central America, with a Key and Distributional Data for the Genus." volume="47" year="1993 a">Rifkind (1993a)</bibRefCitation>
described a third species,
<taxonomicName id="ED39D8C21BB50D77D4FE3BA9C6DE0E4B" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella morula" order="Coleoptera" pageId="27" pageNumber="102" phylum="Arthropoda" rank="species" species="morula">Cymatoderella morula</taxonomicName>
.
<taxonomicName id="28025D1A5C0B9057A284EBD07FA8C116" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella collaris" order="Coleoptera" pageId="27" pageNumber="102" phylum="Arthropoda" rank="species" species="collaris">Cymatoderella collaris</taxonomicName>
is widely distributed throughout North and Central America; the species ranges from the eastern and southern United States, extending southward to Mexico and the Central American countries of Guatemala, Honduras and El Salvador.
<taxonomicName id="D9E0A01B485396DEE33D4830379077F5" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella morula" order="Coleoptera" pageId="27" pageNumber="102" phylum="Arthropoda" rank="species" species="morula">Cymatoderella morula</taxonomicName>
and
<taxonomicName id="096208F211B62AA87260203D1CF12B5A" lsidName="C. patagoniae" pageId="27" pageNumber="102" rank="species" species="patagoniae">C. patagoniae</taxonomicName>
are species with a limited distributional range.
<taxonomicName id="C2E5566C063DA2F7E0EEE80193722C8E" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella morula" order="Coleoptera" pageId="27" pageNumber="102" phylum="Arthropoda" rank="species" species="morula">Cymatoderella morula</taxonomicName>
inhabits regions of southern Mexico, Guatemala, Honduras, and Nicaragua (
<bibRefCitation id="919CE348AD2FD0C83F0CA5219F020FD3" author="Rifkind, J" journalOrPublisher="The Coleopterists Bulletin" pageId="77" pageNumber="152" pagination="279 - 284" title="A New Species of Cymatoderella Barr (Coleoptera: Cleridae) from Mexico and Central America, with a Key and Distributional Data for the Genus." volume="47" year="1993 a">Rifkind 1993a</bibRefCitation>
), and
<taxonomicName id="DE26449A8A526F617F880457E7CC039B" lsidName="C. patagoniae" pageId="27" pageNumber="102" rank="species" species="patagoniae">C. patagoniae</taxonomicName>
is found in southern Arizona and Guerrero, Michoacan and Sonora, Mexico.
</paragraph>
</subSubSection>
<subSubSection id="6FD542F185107067E28926ED8128FFEF" lastPageId="28" lastPageNumber="103" pageId="27" pageNumber="102" type="key">
<paragraph id="D5F9335BB135DEFBFF8D99A6BF06C4F4" pageId="27" pageNumber="102">
Key to species of
<taxonomicName id="1B53834C5EC50F4898F20B4BF30F7E53" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="27" pageNumber="102" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
Barr
</paragraph>
<paragraph id="6F5BFCF3402B5989B11E4161B1F68FF4" pageId="28" pageNumber="103">
<table id="52FE59A96F54C5E8C628B70F48E1F21F" pageId="28" pageNumber="103">
<tr id="A64BAE198F6A2618A9BD1A7B0ABD4A7C" pageId="28" pageNumber="103">
<td id="12924E09B5D6836D8B32DEC7C6EC6A4C" colspan="1" pageId="28" pageNumber="103" rowspan="1">
<pageBreakToken id="5DF8C7DD8F42614A60F23A9E900F3BAB" pageId="28" pageNumber="103" start="start">3</pageBreakToken>
D
</td>
<td id="A055D9A9479C097A40A16D6E8C9B8818" colspan="1" pageId="28" pageNumber="103" rowspan="1">
<taxonomicName id="7D9907913E02C34D5F8FF8F9C61084AC" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella patagoniae" order="Coleoptera" pageId="28" pageNumber="103" phylum="Arthropoda" rank="species" species="patagoniae">Cymatoderella patagoniae</taxonomicName>
</td>
</tr>
<tr id="BA87AA054BE2E3382218693B40EB6D94" pageId="28" pageNumber="103">
<td id="F3F0BE926E4007DCA2EB99A7C92F6C16" colspan="1" pageId="28" pageNumber="103" rowspan="1">2</td>
</tr>
<tr id="40080867D9A387A545406D3E9CB5ECEF" pageId="28" pageNumber="103">
<td id="970A2DB69B298088B90FD850455E4ADE" colspan="1" pageId="28" pageNumber="103" rowspan="1">9D</td>
<td id="8EB116A68689D178A4589F00294CDD30" colspan="1" pageId="28" pageNumber="103" rowspan="1">
<taxonomicName id="EAC6BEFA17F51D0801CF06E5BAFFA77A" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella collaris" order="Coleoptera" pageId="28" pageNumber="103" phylum="Arthropoda" rank="species" species="collaris">Cymatoderella collaris</taxonomicName>
</td>
</tr>
<tr id="EC757FCFC532D1BD0A028FEDBA4954B6" pageId="28" pageNumber="103">
<td id="0596B2E4A8FAE7F2F088BADBD931DB86" colspan="1" pageId="28" pageNumber="103" rowspan="1">9E</td>
<td id="5C6E6B86D883C76492A32EE143872C24" colspan="1" pageId="28" pageNumber="103" rowspan="1">
<taxonomicName id="87A01C8E7A33AB7E0A9D768714C9255A" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella morula" order="Coleoptera" pageId="28" pageNumber="103" phylum="Arthropoda" rank="species" species="morula">Cymatoderella morula</taxonomicName>
</td>
</tr>
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</paragraph>
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