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<document id="0BBC45AACAAE14648C561D1AAD24145E" ID-DOI="10.11646/zootaxa.5403.4.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10556468" ID-ZooBank="D338D548-35CB-4A25-89F7-7EB5D7CC0348" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706018333181" checkinUser="plazi" docAuthor="Watung, Jackson F., Darmawan, Darmawan, Suwito, Awit, Narakusumo, Raden Pramesa, Nugroho, Hari, Encilia, Encilia, Qodri, Agmal, Peggie, Djunijanti, Ubaidillah, Rosichon &amp; Sutrisno, Hari" docDate="2024" docId="32239640FFF2F170FF78FA98FD80FB24" docLanguage="en" docName="zootaxa.5403.4.5.pdf" docOrigin="Zootaxa 5403 (4)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5403.4.5/52740" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyphodes ahsanae Watung &amp; Darmawan &amp; Suwito &amp; Narakusumo &amp; Nugroho &amp; Encilia &amp; Qodri &amp; Peggie &amp; Ubaidillah &amp; Sutrisno 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="476" masterDocId="CE1AEE38FFF4F177FFEFFFC3FFD1FF85" masterDocTitle="Description of two new species in the genus Glyphodes Guenée from Indonesia (Lepidoptera: Crambidae, Spilomelinae)" masterLastPageNumber="478" masterPageNumber="469" pageNumber="475" updateTime="1706019061947" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0E8AF693688D001976B235700ADABBED">Description of two new species in the genus Glyphodes Guenée from Indonesia (Lepidoptera: Crambidae, Spilomelinae)</mods:title>
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<mods:namePart id="FD15DF943C00F7A48F8B050CBA64E366">Watung, Jackson F.</mods:namePart>
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<mods:namePart id="6C78FC2DADD8682A37404CF5E9341AA8">Encilia, Encilia</mods:namePart>
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<mods:namePart id="6BFDD1DEE463509B0E0B90D48168AA9A">Qodri, Agmal</mods:namePart>
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<mods:namePart id="D64A7BCF76EC0A456FE31785FB2F18EA">Peggie, Djunijanti</mods:namePart>
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<mods:namePart id="5022676D57CAC8163E3330B3499F141A">Ubaidillah, Rosichon</mods:namePart>
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<heading id="E17D903AFFF2F171FF78FA98FDB1FAF3" bold="true" box="[151,608,1371,1398]" fontSize="11" level="1" pageId="6" pageNumber="475" reason="1">
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<taxonomicName id="7D8A5CD5FFF2F171FF78FA98FE27FAF0" authority="Sutrisno" authorityName="Watung &amp; Darmawan &amp; Suwito &amp; Narakusumo &amp; Nugroho &amp; Encilia &amp; Qodri &amp; Peggie &amp; Ubaidillah &amp; Sutrisno" authorityYear="2024" box="[151,502,1371,1398]" class="Insecta" family="Crambidae" genus="Glyphodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="475" phylum="Arthropoda" rank="species" species="ahsanae" status="sp. nov.">
<emphasis id="88FEFB44FFF2F171FF78FA98FE52FAF3" bold="true" box="[151,387,1371,1398]" italics="true" pageId="6" pageNumber="475">Glyphodes ahsanae</emphasis>
Sutrisno
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, 
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<subSubSection id="F29074DDFFF2F171FF78FABCFE7AFA1F" box="[151,427,1407,1434]" pageId="6" pageNumber="475" type="description">
<paragraph id="BA352756FFF2F171FF78FABCFE7AFA1F" blockId="6.[151,608,1371,1434]" box="[151,427,1407,1434]" pageId="6" pageNumber="475">
(
<figureCitation id="22B13BD3FFF2F171FF70FABCFEC4FA1F" box="[159,277,1407,1434]" captionStart="FIGURES 1" captionStartId="3.[151,264,1761,1785]" captionTargetBox="[154,1433,486,1735]" captionTargetId="figure-159@3.[154,1433,486,1735]" captionTargetPageId="3" captionText="FIGURES 1A–F. (A) Glyphodes stolalis ♂ (MZB. Lepi. 715), (B) G. stolalis ♀ (MZB. Lepi. 716), (C) G. nurfitriae sp. nov. ♂ (holotype; MZB. Lepi. 687.01) (D) G. nurfitriae sp. nov. ♀ (paratype; MZB. Lepi. 678.01) (E) G. ahsanae sp. nov. ♂ (holotype; MZB. Lepi. 682.01), (F) G. ahsanae sp. nov. ♀ (paratype; MZB. Lepi. 713.01). Abbreviations, amb= antemedial band, ap= apex, bb= basal band, mb= medial band, pmb= postmedial band. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556511" httpUri="https://zenodo.org/record/10556511/files/figure.png" pageId="6" pageNumber="475">Figs 1E–F</figureCitation>
, 
<figureCitation id="22B13BD3FFF2F171FECFFA43FE8FFA1F" box="[288,350,1408,1434]" captionStart="FIGURES 2" captionStartId="4.[151,264,1891,1915]" captionTargetBox="[302,1285,181,1866]" captionTargetId="figure-24@4.[302,1285,181,1866]" captionTargetPageId="4" captionText="FIGURES 2A–F. Male genitalia of Glyphodes spp.: G. stolalis (MZB. Lepi.715): (A) whole apparatus, (B) Phallus. G. nurfitriae sp. nov. (holotype; MZB. Lepi. 687): (C) whole apparatus, (D) Phallus. G. ahsanae sp. nov. (holotype; MZB. Lepi. 682): (E) whole apparatus, (F) Phallus. Abbreviations, blc= bundle lamellar coremata, ct= cornuti, fb= fibula, sc= saccus, tg= tegumen, tr= transtilla, uc= uncus, vc= vinculum, vlv= valva. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556515" httpUri="https://zenodo.org/record/10556515/files/figure.png" pageId="6" pageNumber="475">2E–F</figureCitation>
, 
<figureCitation id="22B13BD3FFF2F171FE86FA43FE72FA1F" box="[361,419,1408,1434]" captionStart="FIGURES 3" captionStartId="5.[151,264,1873,1897]" captionTargetBox="[328,1260,181,1849]" captionTargetId="figure-18@5.[328,1260,181,1849]" captionTargetPageId="5" captionText="FIGURES 3A–3I. Female genitalia of Glyphodes spp.: G. stolalis (MZB. Lepi. 716): whole apparatus (A), ovipositor lobe (B), corpus bursae (C). G. nurfitriae sp. nov. (paratype; MZB. Lepi. 678): whole apparatus (D), ovipositor lobe (E), corpus bursae (F). G. ahsanae sp. nov. (paratype; MZB. Lepi. 713): whole apparatus (G), ovipositor lobe (H), corpus bursae (I). Abbreviations, an= antrum, cb= corpus bursae, dcb= ductus corpus bursae, lpv= lamella postvaginalis, ol= ovipositor lobe. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556517" httpUri="https://zenodo.org/record/10556517/files/figure.png" pageId="6" pageNumber="475">3G–I</figureCitation>
)
</paragraph>
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<subSubSection id="F29074DDFFF2F171FF78FA04FBD3F93F" pageId="6" pageNumber="475" type="diagnosis">
<paragraph id="BA352756FFF2F171FF78FA04FBD3F93F" blockId="6.[151,1437,1479,2010]" pageId="6" pageNumber="475">
<emphasis id="88FEFB44FFF2F171FF78FA04FEC3FA64" bold="true" box="[151,274,1479,1505]" pageId="6" pageNumber="475">Diagnosis.</emphasis>
<taxonomicName id="7D8A5CD5FFF2F171FEF4FA04FE28FA64" authority="Watung &amp; Darmawan &amp; Suwito &amp; Narakusumo &amp; Nugroho &amp; Encilia &amp; Qodri &amp; Peggie &amp; Ubaidillah &amp; Sutrisno, 2024" authorityName="Watung &amp; Darmawan &amp; Suwito &amp; Narakusumo &amp; Nugroho &amp; Encilia &amp; Qodri &amp; Peggie &amp; Ubaidillah &amp; Sutrisno" authorityYear="2024" box="[283,505,1479,1505]" class="Insecta" family="Crambidae" genus="Glyphodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="475" phylum="Arthropoda" rank="species" species="ahsanae" status="sp. nov.">
<emphasis id="88FEFB44FFF2F171FEF4FA04FE28FA64" box="[283,505,1479,1505]" italics="true" pageId="6" pageNumber="475">Glyphodes ahsanae</emphasis>
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is easily distinguished from the most similar species, 
<taxonomicName id="7D8A5CD5FFF2F171FB3AFA0BFA9AFA64" authorityName="Guenee" authorityYear="1854" box="[1237,1355,1479,1506]" class="Insecta" family="Crambidae" genus="Glyphodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="475" phylum="Arthropoda" rank="species" species="stolalis">
<emphasis id="88FEFB44FFF2F171FB3AFA0BFA9AFA64" box="[1237,1355,1479,1506]" italics="true" pageId="6" pageNumber="475">G. stolalis</emphasis>
</taxonomicName>
by the white, cashew nut-shaped marking in between the dark brown medial and postmedial bands, the oblong white spot at the posterior medial band, and the white medial spot at the margin of the hindwing (
<figureCitation id="22B13BD3FFF2F171FB4EF9CCFABDF9AF" box="[1185,1388,1551,1578]" captionStart="FIGURES 1" captionStartId="3.[151,264,1761,1785]" captionTargetBox="[154,1433,486,1735]" captionTargetId="figure-159@3.[154,1433,486,1735]" captionTargetPageId="3" captionText="FIGURES 1A–F. (A) Glyphodes stolalis ♂ (MZB. Lepi. 715), (B) G. stolalis ♀ (MZB. Lepi. 716), (C) G. nurfitriae sp. nov. ♂ (holotype; MZB. Lepi. 687.01) (D) G. nurfitriae sp. nov. ♀ (paratype; MZB. Lepi. 678.01) (E) G. ahsanae sp. nov. ♂ (holotype; MZB. Lepi. 682.01), (F) G. ahsanae sp. nov. ♀ (paratype; MZB. Lepi. 713.01). Abbreviations, amb= antemedial band, ap= apex, bb= basal band, mb= medial band, pmb= postmedial band. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556511" httpUri="https://zenodo.org/record/10556511/files/figure.png" pageId="6" pageNumber="475">Figs 1A–B, 1E–F</figureCitation>
; see arrows). The simple uncus, apically pointed, with dense setae, the phallus with a thorn cornutus and corpus bursae with a pair of signa, medially bearing a spur, are also diagnostic characters of this species (
<figureCitation id="22B13BD3FFF2F171FB45F994FAF3F9F7" box="[1194,1314,1623,1650]" captionStart="FIGURES 2" captionStartId="4.[151,264,1891,1915]" captionTargetBox="[302,1285,181,1866]" captionTargetId="figure-24@4.[302,1285,181,1866]" captionTargetPageId="4" captionText="FIGURES 2A–F. Male genitalia of Glyphodes spp.: G. stolalis (MZB. Lepi.715): (A) whole apparatus, (B) Phallus. G. nurfitriae sp. nov. (holotype; MZB. Lepi. 687): (C) whole apparatus, (D) Phallus. G. ahsanae sp. nov. (holotype; MZB. Lepi. 682): (E) whole apparatus, (F) Phallus. Abbreviations, blc= bundle lamellar coremata, ct= cornuti, fb= fibula, sc= saccus, tg= tegumen, tr= transtilla, uc= uncus, vc= vinculum, vlv= valva. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556515" httpUri="https://zenodo.org/record/10556515/files/figure.png" pageId="6" pageNumber="475">Figs 2E–F</figureCitation>
, 
<figureCitation id="22B13BD3FFF2F171FAC0F99BFABDF9F7" box="[1327,1388,1624,1650]" captionStart="FIGURES 3" captionStartId="5.[151,264,1873,1897]" captionTargetBox="[328,1260,181,1849]" captionTargetId="figure-18@5.[328,1260,181,1849]" captionTargetPageId="5" captionText="FIGURES 3A–3I. Female genitalia of Glyphodes spp.: G. stolalis (MZB. Lepi. 716): whole apparatus (A), ovipositor lobe (B), corpus bursae (C). G. nurfitriae sp. nov. (paratype; MZB. Lepi. 678): whole apparatus (D), ovipositor lobe (E), corpus bursae (F). G. ahsanae sp. nov. (paratype; MZB. Lepi. 713): whole apparatus (G), ovipositor lobe (H), corpus bursae (I). Abbreviations, an= antrum, cb= corpus bursae, dcb= ductus corpus bursae, lpv= lamella postvaginalis, ol= ovipositor lobe. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556517" httpUri="https://zenodo.org/record/10556517/files/figure.png" pageId="6" pageNumber="475">3G–I</figureCitation>
; see arrows). In the male genitalia of the 
<taxonomicName id="7D8A5CD5FFF2F171FDC3F9BFFD4EF910" authorityName="Guenee" authorityYear="1854" box="[556,671,1659,1686]" class="Insecta" family="Crambidae" genus="Glyphodes" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="475" phylum="Arthropoda" rank="species" species="stolalis">
<emphasis id="88FEFB44FFF2F171FDC3F9BFFD4EF910" box="[556,671,1659,1686]" italics="true" pageId="6" pageNumber="475">G. stolalis</emphasis>
</taxonomicName>
, the uncus is blunt apically, the phallus bears a single thorn cornutus without accessories, and the corpus bursae lack a signum (
<figureCitation id="22B13BD3FFF2F171FCCBF95CFC72F93F" box="[804,931,1695,1722]" captionStart="FIGURES 2" captionStartId="4.[151,264,1891,1915]" captionTargetBox="[302,1285,181,1866]" captionTargetId="figure-24@4.[302,1285,181,1866]" captionTargetPageId="4" captionText="FIGURES 2A–F. Male genitalia of Glyphodes spp.: G. stolalis (MZB. Lepi.715): (A) whole apparatus, (B) Phallus. G. nurfitriae sp. nov. (holotype; MZB. Lepi. 687): (C) whole apparatus, (D) Phallus. G. ahsanae sp. nov. (holotype; MZB. Lepi. 682): (E) whole apparatus, (F) Phallus. Abbreviations, blc= bundle lamellar coremata, ct= cornuti, fb= fibula, sc= saccus, tg= tegumen, tr= transtilla, uc= uncus, vc= vinculum, vlv= valva. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556515" httpUri="https://zenodo.org/record/10556515/files/figure.png" pageId="6" pageNumber="475">Figs 2A–B</figureCitation>
, 
<figureCitation id="22B13BD3FFF2F171FC41F963FC27F93F" box="[942,1014,1696,1722]" captionStart="FIGURES 3" captionStartId="5.[151,264,1873,1897]" captionTargetBox="[328,1260,181,1849]" captionTargetId="figure-18@5.[328,1260,181,1849]" captionTargetPageId="5" captionText="FIGURES 3A–3I. Female genitalia of Glyphodes spp.: G. stolalis (MZB. Lepi. 716): whole apparatus (A), ovipositor lobe (B), corpus bursae (C). G. nurfitriae sp. nov. (paratype; MZB. Lepi. 678): whole apparatus (D), ovipositor lobe (E), corpus bursae (F). G. ahsanae sp. nov. (paratype; MZB. Lepi. 713): whole apparatus (G), ovipositor lobe (H), corpus bursae (I). Abbreviations, an= antrum, cb= corpus bursae, dcb= ductus corpus bursae, lpv= lamella postvaginalis, ol= ovipositor lobe. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556517" httpUri="https://zenodo.org/record/10556517/files/figure.png" pageId="6" pageNumber="475">3A–C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F29074DDFFF2F170FF28F900FC40FD2C" lastPageId="7" lastPageNumber="476" pageId="6" pageNumber="475" type="description">
<paragraph id="BA352756FFF2F170FF28F900FC4BFE98" blockId="6.[151,1437,1479,2010]" lastBlockId="7.[151,1437,150,1185]" lastPageId="7" lastPageNumber="476" pageId="6" pageNumber="475">
<emphasis id="88FEFB44FFF2F171FF28F900FDC6F958" bold="true" box="[199,535,1731,1758]" pageId="6" pageNumber="475">
Description. 
<emphasis id="88FEFB44FFF2F171FE8EF900FE4DF95B" bold="true" box="[353,412,1731,1758]" italics="true" pageId="6" pageNumber="475">Male</emphasis>
(
<figureCitation id="22B13BD3FFF2F171FE43F900FDD6F958" box="[428,519,1731,1757]" captionStart="FIGURES 1" captionStartId="3.[151,264,1761,1785]" captionTargetBox="[154,1433,486,1735]" captionTargetId="figure-159@3.[154,1433,486,1735]" captionTargetPageId="3" captionText="FIGURES 1A–F. (A) Glyphodes stolalis ♂ (MZB. Lepi. 715), (B) G. stolalis ♀ (MZB. Lepi. 716), (C) G. nurfitriae sp. nov. ♂ (holotype; MZB. Lepi. 687.01) (D) G. nurfitriae sp. nov. ♀ (paratype; MZB. Lepi. 678.01) (E) G. ahsanae sp. nov. ♂ (holotype; MZB. Lepi. 682.01), (F) G. ahsanae sp. nov. ♀ (paratype; MZB. Lepi. 713.01). Abbreviations, amb= antemedial band, ap= apex, bb= basal band, mb= medial band, pmb= postmedial band. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556511" httpUri="https://zenodo.org/record/10556511/files/figure.png" pageId="6" pageNumber="475">Fig. 1E</figureCitation>
):
</emphasis>
Forewing length 
<quantity id="7D728AB3FFF2F171FD0EF907FC8CF95B" box="[737,861,1732,1758]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="6" pageNumber="475" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
. Frons oblique, white; vertex shabby white. Labial palpus sub-ascending with first segment covered by yellowish-white scales from base to middle and black scales at tip, second segment broadly scaled in front, third segment porrect and embedded in white scales of second segment. Maxillary palpus prominent, triangularly dilated with white scales at tip. Proboscis well-developed, white, scaled at base. Antennae filiform, extend to approximately full forewing length, dorsal surface covered with longitudinal row of brown scales, ventral surface with long white cilia (almost twice diameter of flagellum). Thorax dorsally brown and ventrally white, patagia dark brown, tegulae white at middle. Legs white, epiphysis covered with black scales, inner spur of mid-tibia less than half length of outer spur. Forewings subtriangular, costa straight in basal 2/3 and curved in distal third, basal and antemedial bands obliquely (45°) running outward from costa to dorsum, wide medial band with complex structure medially running from mid-costa obliquely outward to 3/4 of dorsum, margin with white terminal cilia from apex to M 
<subScript id="260E2513FFF3F170FD8CFF32FDBAFF7A" attach="left" box="[611,619,241,255]" fontSize="6" pageId="7" pageNumber="476">3</subScript>
and in between CuA 
<subScript id="260E2513FFF3F170FCB5FF32FCB3FF7A" attach="left" box="[858,866,241,255]" fontSize="6" pageId="7" pageNumber="476">1</subScript>
and CuP. Hindwing with white and diffuse medial spot at postmedial band. Abdomen slender with anal black scale tuft.
</paragraph>
<paragraph id="BA352756FFF3F170FF28FEE5FD96FE54" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FEE5FDD9FEC5" bold="true" box="[199,520,294,321]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FEE5FEBFFEC4" bold="true" box="[199,366,294,321]" italics="true" pageId="7" pageNumber="476">Male genitalia</emphasis>
(
<figureCitation id="22B13BD3FFF3F170FE93FEE5FE2AFEC5" box="[380,507,294,320]" captionStart="FIGURES 2" captionStartId="4.[151,264,1891,1915]" captionTargetBox="[302,1285,181,1866]" captionTargetId="figure-24@4.[302,1285,181,1866]" captionTargetPageId="4" captionText="FIGURES 2A–F. Male genitalia of Glyphodes spp.: G. stolalis (MZB. Lepi.715): (A) whole apparatus, (B) Phallus. G. nurfitriae sp. nov. (holotype; MZB. Lepi. 687): (C) whole apparatus, (D) Phallus. G. ahsanae sp. nov. (holotype; MZB. Lepi. 682): (E) whole apparatus, (F) Phallus. Abbreviations, blc= bundle lamellar coremata, ct= cornuti, fb= fibula, sc= saccus, tg= tegumen, tr= transtilla, uc= uncus, vc= vinculum, vlv= valva. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556515" httpUri="https://zenodo.org/record/10556515/files/figure.png" pageId="7" pageNumber="476">Figs 2E–F</figureCitation>
):
</emphasis>
Tegumen subtriangular; uncus simple, narrow from base to medial part, gradually widened towards subapex, apically pointed, with dense setae; valva simple, semi-ovate, medial costa convex; fibula curved outwardly, pointed apically; juxta prominent, tongue-shaped, weakly sclerotized; transtilla wide, weakly sclerotized; saccus simple, triangular. Coremata base simple, with a bundle of long setae. Phallus weakly sclerotized ventrally, with a single thorn cornutus.
</paragraph>
<paragraph id="BA352756FFF3F170FF28FE19FD2AFD9C" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FE19FE42FE71" bold="true" box="[199,403,474,501]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FE19FECFFE70" bold="true" box="[199,286,474,501]" italics="true" pageId="7" pageNumber="476">Female</emphasis>
(
<figureCitation id="22B13BD3FFF3F170FEC3FE19FE55FE71" box="[300,388,474,500]" captionStart="FIGURES 1" captionStartId="3.[151,264,1761,1785]" captionTargetBox="[154,1433,486,1735]" captionTargetId="figure-159@3.[154,1433,486,1735]" captionTargetPageId="3" captionText="FIGURES 1A–F. (A) Glyphodes stolalis ♂ (MZB. Lepi. 715), (B) G. stolalis ♀ (MZB. Lepi. 716), (C) G. nurfitriae sp. nov. ♂ (holotype; MZB. Lepi. 687.01) (D) G. nurfitriae sp. nov. ♀ (paratype; MZB. Lepi. 678.01) (E) G. ahsanae sp. nov. ♂ (holotype; MZB. Lepi. 682.01), (F) G. ahsanae sp. nov. ♀ (paratype; MZB. Lepi. 713.01). Abbreviations, amb= antemedial band, ap= apex, bb= basal band, mb= medial band, pmb= postmedial band. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556511" httpUri="https://zenodo.org/record/10556511/files/figure.png" pageId="7" pageNumber="476">Fig. 1F</figureCitation>
):
</emphasis>
Similar to male, except abdomen without anal scale tuft and ventral surface of antennae with short grey cilia, less than 1/4 of flagellomere diameter.
</paragraph>
<paragraph id="BA352756FFF3F170FF28FDE1FC40FD2C" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FDE1FDCAFDB9" bold="true" box="[199,539,546,573]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FDE1FE59FDB8" bold="true" box="[199,392,546,573]" italics="true" pageId="7" pageNumber="476">Female genitalia</emphasis>
(
<figureCitation id="22B13BD3FFF3F170FE7BFDE1FDDCFDB9" box="[404,525,546,572]" captionStart="FIGURES 3" captionStartId="5.[151,264,1873,1897]" captionTargetBox="[328,1260,181,1849]" captionTargetId="figure-18@5.[328,1260,181,1849]" captionTargetPageId="5" captionText="FIGURES 3A–3I. Female genitalia of Glyphodes spp.: G. stolalis (MZB. Lepi. 716): whole apparatus (A), ovipositor lobe (B), corpus bursae (C). G. nurfitriae sp. nov. (paratype; MZB. Lepi. 678): whole apparatus (D), ovipositor lobe (E), corpus bursae (F). G. ahsanae sp. nov. (paratype; MZB. Lepi. 713): whole apparatus (G), ovipositor lobe (H), corpus bursae (I). Abbreviations, an= antrum, cb= corpus bursae, dcb= ductus corpus bursae, lpv= lamella postvaginalis, ol= ovipositor lobe. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556517" httpUri="https://zenodo.org/record/10556517/files/figure.png" pageId="7" pageNumber="476">Figs 3G–I</figureCitation>
):
</emphasis>
Ovipositor lobe ovate with scattered faint setae; lamella postvaginalis moderately sclerotized; posterior apophyses short, anterior apophyses as long as posterior apophyses; entrance to ostium bursae narrow, antrum very short (less than 1/5 length of anterior apophyses); ductus bursae weakly sclerotized at basal 1/3; corpus bursae globular with a pair of signa, medially bearing a spur.
</paragraph>
</subSubSection>
<subSubSection id="F29074DDFFF3F170FF28FD70FA57FC68" pageId="7" pageNumber="476" type="materials_examined">
<paragraph id="BA352756FFF3F170FF28FD70FDB3FC90" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="476">
<materialsCitation id="0AE22D0BFFF3F170FF28FD70FDB3FC90" collectingDate="2010-06-22" collectionCode="R, MZB" country="Indonesia" county="North Kolaka" latitude="-3.6481388" location="Tinukari" longLatPrecision="1" longitude="121.12278" municipality="Wawo" pageId="7" pageNumber="476" specimenCount="2" specimenCount-male="2" stateProvince="Sulawesi Tenggara" typeStatus="holotype">
<emphasis id="88FEFB44FFF3F170FF28FD70FEE5FD48" bold="true" box="[199,308,691,717]" pageId="7" pageNumber="476">
<typeStatus id="653199F4FFF3F170FF28FD70FEE5FD48" box="[199,308,691,717]" pageId="7" pageNumber="476" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="AC8CECDFFFF3F170FEAEFD70FE87FD48" box="[321,342,691,717]" count="1" pageId="7" pageNumber="476" type="male">♂</specimenCount>
(
<figureCitation id="22B13BD3FFF3F170FE83FD71FE16FD49" box="[364,455,690,716]" captionStart="FIGURES 1" captionStartId="3.[151,264,1761,1785]" captionTargetBox="[154,1433,486,1735]" captionTargetId="figure-159@3.[154,1433,486,1735]" captionTargetPageId="3" captionText="FIGURES 1A–F. (A) Glyphodes stolalis ♂ (MZB. Lepi. 715), (B) G. stolalis ♀ (MZB. Lepi. 716), (C) G. nurfitriae sp. nov. ♂ (holotype; MZB. Lepi. 687.01) (D) G. nurfitriae sp. nov. ♀ (paratype; MZB. Lepi. 678.01) (E) G. ahsanae sp. nov. ♂ (holotype; MZB. Lepi. 682.01), (F) G. ahsanae sp. nov. ♀ (paratype; MZB. Lepi. 713.01). Abbreviations, amb= antemedial band, ap= apex, bb= basal band, mb= medial band, pmb= postmedial band. Arrows indicate diagnostic characters." figureDoi="http://doi.org/10.5281/zenodo.10556511" httpUri="https://zenodo.org/record/10556511/files/figure.png" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FE83FD71FE16FD49" bold="true" box="[364,455,690,716]" pageId="7" pageNumber="476">Fig. 1E</emphasis>
</figureCitation>
): “ 
<typeStatus id="653199F4FFF3F170FE1FFD70FD8EFD48" box="[496,607,691,717]" pageId="7" pageNumber="476" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="AC8CECDFFFF3F170FD80FD70FD56FD48" box="[623,647,691,717]" count="1" pageId="7" pageNumber="476" type="male">♂</specimenCount>
; 
<collectingCountry id="C29D67C6FFF3F170FD76FD70FCD8FD48" box="[665,777,691,717]" name="Indonesia" pageId="7" pageNumber="476">Indonesia</collectingCountry>
, 
<collectingRegion id="784EE9B4FFF3F170FCF5FD70FC2AFD48" box="[794,1019,691,717]" country="Indonesia" name="Sulawesi Tenggara" pageId="7" pageNumber="476">Southeast Sulawesi</collectingRegion>
, 
<collectingCounty id="53545FDAFFF3F170FBE2FD70FB7EFD48" box="[1037,1199,691,717]" pageId="7" pageNumber="476">North Kolaka</collectingCounty>
, 
<collectingMunicipality id="5A51BD2CFFF3F170FB50FD70FAD8FD48" box="[1215,1289,691,717]" pageId="7" pageNumber="476">Wawo</collectingMunicipality>
, 
<location id="BF55718DFFF3F170FAF6FD70FAAAFD48" LSID="urn:lsid:plazi:treatment:32239640FFF2F170FF78FA98FD80FB24:BF55718DFFF3F170FAF6FD70FAAAFD48" box="[1305,1403,691,717]" country="Indonesia" county="North Kolaka" latitude="-3.6481388" longLatPrecision="1" longitude="121.12278" municipality="Wawo" name="Tinukari" pageId="7" pageNumber="476" stateProvince="Sulawesi Tenggara">Tinukari</location>
, 
<geoCoordinate id="DFBE4191FFF3F170FA62FD70FEDFFD76" degrees="3" direction="south" minutes="38" orientation="latitude" pageId="7" pageNumber="476" precision="1" seconds="53.3" value="-3.6481388">S 3°38ʹ53.3ʺ</geoCoordinate>
<geoCoordinate id="DFBE4191FFF3F170FEFAFD14FE11FD76" box="[277,448,727,755]" degrees="121" direction="east" minutes="07" orientation="longitude" pageId="7" pageNumber="476" precision="1" seconds="22.0" value="121.12278">E 121°07ʹ22.0ʺ</geoCoordinate>
, 
<date id="CE340196FFF3F170FE21FD14FD98FD74" box="[462,585,727,753]" pageId="7" pageNumber="476" value="2010-06-22">
<collectingDate id="DE70F87EFFF3F170FE21FD14FD98FD74" box="[462,585,727,753]" pageId="7" pageNumber="476" value="2010-06-22">22.vi.2010</collectingDate>
</date>
; Ubaidillah 
<collectionCode id="DC9BBF93FFF3F170FD3DFD14FD36FD74" box="[722,743,727,753]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="476">R</collectionCode>
, Kimsey B, Nugroho H, Lupyaningdyah P, Darmawan, 
<collectionCode id="DC9BBF93FFF3F170FAB6FD14FA48FD74" box="[1369,1433,727,753]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="476" type="Museum">MZB</collectionCode>
. Lepi. 682 (
<collectionCode id="DC9BBF93FFF3F170FEFEFD38FE80FC90" box="[273,337,763,789]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="476" type="Museum">MZB</collectionCode>
) / 
<typeStatus id="653199F4FFF3F170FE80FD38FDD8FC90" box="[367,521,763,789]" pageId="7" pageNumber="476" type="holotype">HOLOTYPE</typeStatus>
”; 
<collectionCode id="DC9BBF93FFF3F170FDF1FD38FD8FFC90" box="[542,606,763,789]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="476" type="Museum">MZB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="BA352756FFF3F170FF28FCDCFA57FC68" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="476">
<materialsCitation id="0AE22D0BFFF3F170FF28FCDCFD55FCBC" box="[199,644,799,825]" collectingDate="2010-06-22" collectionCode="R, MZB" country="Indonesia" county="North Kolaka" latitude="-3.6481388" location="Tinukari" longLatPrecision="1" longitude="121.12278" municipality="Wawo" pageId="7" pageNumber="476" specimenCount="2" specimenCount-male="2" stateProvince="Sulawesi Tenggara" typeStatus="paratype">
<emphasis id="88FEFB44FFF3F170FF28FCDCFE99FCBC" bold="true" box="[199,328,799,825]" pageId="7" pageNumber="476">
<typeStatus id="653199F4FFF3F170FF28FCDCFE92FCBC" box="[199,323,799,825]" pageId="7" pageNumber="476" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="AC8CECDFFFF3F170FEA0FCDCFEACFCBC" box="[335,381,799,825]" count="2" pageId="7" pageNumber="476" type="male">2 ♂</specimenCount>
; same data as holotype
</materialsCitation>
; 
<materialsCitation id="0AE22D0BFFF3F170FD60FCDCFB19FCBC" box="[655,1224,798,825]" collectingDate="2010-06-25" collectionCode="MZB" country="Indonesia" county="North Kolaka" latitude="-3.6481388" location="Tinukari" longLatPrecision="1" longitude="121.12278" municipality="Wawo" pageId="7" pageNumber="476" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Tenggara" typeStatus="paratype">
<specimenCount id="AC8CECDFFFF3F170FD60FCDCFD6DFCBC" box="[655,700,799,825]" count="1" pageId="7" pageNumber="476" type="male">1 ♂</specimenCount>
; same data as previous, but 
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(
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; 
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; N. 
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ʺE, 
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; 
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S &amp; 
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(
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)
</materialsCitation>
; 
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<specimenCount id="AC8CECDFFFF3F170FEA5FCA4FEA4FC04" box="[330,373,871,897]" count="1" pageId="7" pageNumber="476" type="female">1 ♀</specimenCount>
; 
<collectingCountry id="C29D67C6FFF3F170FE6CFCA4FE22FC04" box="[387,499,871,897]" name="Indonesia" pageId="7" pageNumber="476">Indonesia</collectingCountry>
, 
<collectingRegion id="784EE9B4FFF3F170FE11FCA4FDA0FC04" box="[510,625,871,897]" country="Indonesia" name="Jawa Barat" pageId="7" pageNumber="476">West Java</collectingRegion>
, 
<collectingMunicipality id="5A51BD2CFFF3F170FD92FCA4FC37FC04" box="[637,998,871,897]" pageId="7" pageNumber="476">Gunung Halimun National Park</collectingMunicipality>
, 
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, 
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<geoCoordinate id="DFBE4191FFF3F170FB3EFCA4FA80FC06" box="[1233,1361,871,899]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="7" pageNumber="476" precision="925" value="106.53333">E 106° 32 ʹ</geoCoordinate>
, 
<date id="CE340196FFF3F170FAB1FCA4FF02FC20" pageId="7" pageNumber="476" value="2000-10-28">
<collectingDate id="DE70F87EFFF3F170FAB1FCA4FF02FC20" pageId="7" pageNumber="476" value="2000-10-28">28.X. 2000</collectingDate>
</date>
; leg. 
<collectorName id="177F4280FFF3F170FEFBFC48FE02FC20" box="[276,467,907,933]" pageId="7" pageNumber="476">Woro, E. Cholik</collectorName>
, 
<collectorName id="177F4280FFF3F170FE0EFC48FDFBFC20" box="[481,554,907,933]" pageId="7" pageNumber="476">Sarino</collectorName>
(
<collectionCode id="DC9BBF93FFF3F170FDD3FC48FDAFFC20" box="[572,638,907,933]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="476" type="Museum">MZB</collectionCode>
)
</materialsCitation>
; 
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<specimenCount id="AC8CECDFFFF3F170FD7AFC48FD14FC20" box="[661,709,907,933]" count="2" pageId="7" pageNumber="476" type="male">2 ♂</specimenCount>
; 
<collectingCountry id="C29D67C6FFF3F170FD3BFC48FC95FC20" box="[724,836,907,933]" name="Indonesia" pageId="7" pageNumber="476">Indonesia</collectingCountry>
, 
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, 
<collectorName id="177F4280FFF3F170FC3AFC48FB98FC20" box="[981,1097,907,933]" pageId="7" pageNumber="476">Sukabumi</collectorName>
, 
<collectorName id="177F4280FFF3F170FBB8FC48FB6BFC20" box="[1111,1210,907,933]" pageId="7" pageNumber="476">Cikaniki</collectorName>
, 
<collectionCode id="DC9BBF93FFF3F170FB27FC48FAF4FC20" box="[1224,1317,907,933]" pageId="7" pageNumber="476">TNGHS</collectionCode>
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<materialsCitation id="0AE22D0BFFF3F170FAC0FC48FBB1FC4C" collectingDate="2009-10-08" collectionCode="MZB" collectorName="Legs. Woro &amp; Yayuk &amp; E. Cholik &amp; Sarino" country="Indonesia" latitude="-6.730278" longLatPrecision="21" longitude="106.70638" pageId="7" pageNumber="476" specimenCount="1" stateProvince="Jawa Barat" typeStatus="paratype">
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; 
<date id="CE340196FFF3F170FE93FC6CFE24FC4C" box="[380,501,943,969]" pageId="7" pageNumber="476" value="2009-10-08">
<collectingDate id="DE70F87EFFF3F170FE93FC6CFE24FC4C" box="[380,501,943,969]" pageId="7" pageNumber="476" value="2009-10-08">08.X.2009</collectingDate>
</date>
. 
<collectorName id="177F4280FFF3F170FDEEFC6CFD55FC4C" box="[513,644,943,969]" pageId="7" pageNumber="476">Legs. Woro</collectorName>
AN, 
<collectorName id="177F4280FFF3F170FD2FFC6CFCD9FC4C" box="[704,776,943,969]" pageId="7" pageNumber="476">Yayuk</collectorName>
RS, 
<collectorName id="177F4280FFF3F170FCADFC6CFC60FC4C" box="[834,945,943,969]" pageId="7" pageNumber="476">E. Cholik</collectorName>
, 
<collectorName id="177F4280FFF3F170FC52FC6CFBD7FC4C" box="[957,1030,943,969]" pageId="7" pageNumber="476">Sarino</collectorName>
(
<collectionCode id="DC9BBF93FFF3F170FBF9FC6CFB89FC4C" box="[1046,1112,943,969]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="476" type="Museum">MZB</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0AE22D0BFFF3F170FB82FC6CFE04FC68" collectionCode="TNGHS" collectorName="Sukabumi &amp; Cijeruk" country="Indonesia" location="West Java" pageId="7" pageNumber="476" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="AC8CECDFFFF3F170FB82FC6CFB48FC4C" box="[1133,1177,943,969]" count="1" pageId="7" pageNumber="476" type="female">1 ♀</specimenCount>
; 
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, 
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; 
<collectorName id="177F4280FFF3F170FF78FC10FEDAFC68" box="[151,267,979,1005]" pageId="7" pageNumber="476">Sukabumi</collectorName>
, 
<collectorName id="177F4280FFF3F170FEF8FC10FEBFFC68" box="[279,366,979,1005]" pageId="7" pageNumber="476">Cijeruk</collectorName>
, 
<collectionCode id="DC9BBF93FFF3F170FE97FC10FE04FC68" box="[376,469,979,1005]" pageId="7" pageNumber="476">TNGHS</collectionCode>
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<geoCoordinate id="DFBE4191FFF3F170FE32FC10FDA0FC6A" box="[477,625,979,1007]" degrees="06" direction="south" minutes="45" orientation="latitude" pageId="7" pageNumber="476" precision="15" seconds="14" value="-6.753889">S 06° 45ʹ 14ʺ</geoCoordinate>
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; 
<collectorName id="177F4280FFF3F170FCC5FC10FC0DFC68" box="[810,988,979,1005]" pageId="7" pageNumber="476">Legs. Woro AN</collectorName>
, 
<collectorName id="177F4280FFF3F170FC09FC10FBFFFC68" box="[998,1070,979,1005]" pageId="7" pageNumber="476">Yayuk</collectorName>
RS, 
<collectorName id="177F4280FFF3F170FB8AFC10FB05FC68" box="[1125,1236,979,1005]" pageId="7" pageNumber="476">E. Cholik</collectorName>
, 
<collectorName id="177F4280FFF3F170FB30FC10FAF9FC68" box="[1247,1320,979,1005]" pageId="7" pageNumber="476">Sarino</collectorName>
(
<collectionCode id="DC9BBF93FFF3F170FAD8FC10FAA8FC68" box="[1335,1401,979,1005]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="476" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F29074DDFFF3F170FF28FC34FEA7FBDC" pageId="7" pageNumber="476" type="etymology">
<paragraph id="BA352756FFF3F170FF28FC34FEA7FBDC" blockId="7.[151,1437,150,1185]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FC34FE99FB94" bold="true" box="[199,328,1015,1041]" pageId="7" pageNumber="476">Etymology</emphasis>
. The specific name 
<taxonomicName id="7D8A5CD5FFF3F170FDDBFC34FD40FB94" authorityName="Watung &amp; Darmawan &amp; Suwito &amp; Narakusumo &amp; Nugroho &amp; Encilia &amp; Qodri &amp; Peggie &amp; Ubaidillah &amp; Sutrisno" authorityYear="2024" box="[564,657,1015,1041]" class="Insecta" family="Crambidae" genus="Glyphodes" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="476" phylum="Arthropoda" rank="species" species="ahsanae">
<emphasis id="88FEFB44FFF3F170FDDBFC34FD40FB94" box="[564,657,1015,1041]" italics="true" pageId="7" pageNumber="476">ahsanae</emphasis>
</taxonomicName>
is derived from the name of the 2 
<superScript id="4DFF8A1EFFF3F170FBCFFC35FBE1FB81" attach="left" box="[1056,1072,1014,1028]" fontSize="6" pageId="7" pageNumber="476">nd</superScript>
daughter, Annisa Ahsan, of the senior author, Hari Sutrisno. This name is dedicated to her support during the preparation of this manuscript.A noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="F29074DDFFF3F170FF28FBA1FB45FBF8" box="[199,1172,1122,1149]" pageId="7" pageNumber="476" type="distribution">
<paragraph id="BA352756FFF3F170FF28FBA1FB45FBF8" blockId="7.[151,1437,150,1185]" box="[199,1172,1122,1149]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FBA1FE88FBF9" bold="true" box="[199,345,1122,1148]" pageId="7" pageNumber="476">Distribution</emphasis>
. 
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, 
<collectingRegion id="784EE9B4FFF3F170FE0AFBA0FD6FFBF8" box="[485,702,1123,1149]" country="Indonesia" name="Sulawesi Tenggara" pageId="7" pageNumber="476">Southeast Sulawesi</collectingRegion>
and 
<collectingRegion id="784EE9B4FFF3F170FD1BFBA0FC73FBF8" box="[756,930,1123,1149]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="476">North Sulawesi</collectingRegion>
in 
<collectingCountry id="C29D67C6FFF3F170FC29FBA0FBE2FBF8" box="[966,1075,1123,1149]" name="Indonesia" pageId="7" pageNumber="476">Indonesia</collectingCountry>
(
<figureCitation id="22B13BD3FFF3F170FBADFBA0FB56FBF8" box="[1090,1159,1123,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1229,1253]" captionTargetBox="[357,1230,849,1204]" captionTargetId="figure-259@6.[357,1231,849,1206]" captionTargetPageId="6" captionText="FIGURE 4. Distribution of the two new species in Indonesia: Glyphodes nurfitriae sp. nov. (white dot) and G. ahsanae sp. nov. (blue dots)." figureDoi="http://doi.org/10.5281/zenodo.10556519" httpUri="https://zenodo.org/record/10556519/files/figure.png" pageId="7" pageNumber="476">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F29074DDFFF3F170FF28FB44FD80FB24" box="[199,593,1159,1185]" pageId="7" pageNumber="476" type="discussion">
<paragraph id="BA352756FFF3F170FF28FB44FD80FB24" blockId="7.[151,1437,150,1185]" box="[199,593,1159,1185]" pageId="7" pageNumber="476">
<emphasis id="88FEFB44FFF3F170FF28FB44FEE3FB24" bold="true" box="[199,306,1159,1185]" pageId="7" pageNumber="476">Remark:</emphasis>
The adults are nocturnal.
</paragraph>
</subSubSection>
</treatment>
</document>