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<document id="E93E7EDB858F49C8F3024F06DB2D4EAB" ID-CLB-Dataset="302193" ID-DOI="10.11646/phytotaxa.659.3.1" ID-GBIF-Dataset="52524fe3-dfb8-4afc-829a-31089bb4e306" ID-ISSN="1179-3163" ID-Zenodo-Dep="13217894" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1722613780751" checkinUser="felipe" docAuthor="Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, Zhou, Zhuo, Thepkayson, Khamphanh, Luong, Thien Tam &amp; Zhang, Li-Bing" docDate="2024" docId="4B175324C955A514D1BAFF5CFD26A9C1" docLanguage="en" docName="phytotaxa.659.3.1.pdf" docOrigin="Phytotaxa 659 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.659.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Leptochilus daklakensis Liang Zhang, X. M. Zhou, T. T. Luong &amp; Li Bing Zhang 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="220" masterDocId="B72E2B5CC953A513D132FFC2FFE5AB4D" masterDocTitle="Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot" masterLastPageNumber="235" masterPageNumber="213" pageNumber="219" updateTime="1725399130512" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8E53B057EFFBE6902FD6DE344341F90F">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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<mods:namePart id="B4DD46C02DCEE98A3A56A0A2D80E455F">Zhang, Liang</mods:namePart>
<mods:nameIdentifier id="AC27E3CFB8A3E2CC94C3F7BFBEB56453" type="ORCID">0000-0003-3784-3135</mods:nameIdentifier>
<mods:affiliation id="BDCA6F66AEBFBC23E9E9433E1A74E6AC">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier id="ABC897FFA62C529C95E81385A13B538C" type="email">zhangliang@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:namePart id="A200392C15BE328D22E31AB3BAA7C837">Lu, Ngan Thi</mods:namePart>
<mods:nameIdentifier id="BC253253C564607BD009ABFDC9FD14B1" type="ORCID">0000-0001-9823-2018</mods:nameIdentifier>
<mods:affiliation id="6288FC7980AD7A9029FCDD9DF0E8B370">Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="9FA95BA7294DD72E02BAB8F6702A94BF" type="email">vnmngan@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EBBA53140F3941F3913254372DC03061">Zhou, Xin-Mao</mods:namePart>
<mods:nameIdentifier id="0B28A8933C378F3C65668B941D0C696A" type="ORCID">0000-0003-3555-7784</mods:nameIdentifier>
<mods:affiliation id="F37F2F21B1DE1979AE762DD7B11FDD19">School of Ecology and Environmental Science, Yunnan University, Kunming, Yunnan 650091, China</mods:affiliation>
<mods:nameIdentifier id="E6CF822B73F914D499247CE6AD7C20BA" type="email">xinmao.zhou@ynu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="3DEEBAB46A327AFA36C788599B665D63">Zhou, Zhuo</mods:namePart>
<mods:nameIdentifier id="849A3E5B61C5812F2968BFA2FFBED924" type="ORCID">0000-0001-9682-5717</mods:nameIdentifier>
<mods:affiliation id="C49B1C4C5E60605008D8F8ADF3C0BC1C">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier id="CF8B55B9A6E2860F664A91E2C4990464" type="email">zhouzhuo@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:namePart id="FA9AA52E72037C88256A8471AA852BB6">Thepkayson, Khamphanh</mods:namePart>
<mods:nameIdentifier id="854D3F6F97AF5E0652938050DFB95C3B" type="ORCID">0009-0001-1757-9907</mods:nameIdentifier>
<mods:affiliation id="C87D3D9DA8A5AEAEB5C334F4B0992A9B">Institute of Traditional Medicine, Vientiane, Lao PDR</mods:affiliation>
<mods:nameIdentifier id="4FBA19E2F4B489A2B2BDC9AC77634F06" type="email">khamphanh@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="DA9426332F4475F151EB9DF212B8D8EB">Luong, Thien Tam</mods:namePart>
<mods:nameIdentifier id="6CAB7FB406DB4149FF8938FC26B2B95C" type="ORCID">0000-0001-8354-6321</mods:nameIdentifier>
<mods:affiliation id="17EB96B3B9C236B07B303FFD1EC65080">Department of Ecology - Evolutionary Biology, Vietnam National University Ho Chi Minh City (VNUHCM) - University of Science, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam</mods:affiliation>
<mods:nameIdentifier id="33F64F7C57863A7FB523A8C31D538FA2" type="email">lttam1511@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3E734BF9F57AF1F438035DCCA57916C7">Zhang, Li-Bing</mods:namePart>
<mods:nameIdentifier id="456BEC4450C6AF1119C0841FC2721C07" type="ORCID">0000-0002-4905-040X</mods:nameIdentifier>
<mods:affiliation id="C263E55E7827ED638CF23D693B25079B">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:nameIdentifier id="9BDDB6B5687DC47F2B38122BBDBA13C5" type="email">Libing.Zhang@mobot.org</mods:nameIdentifier>
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<mods:title id="B28B1AB4BC32E234A30793EC283F3604">Phytotaxa</mods:title>
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<mods:part id="3883E18E5D452C58045C1278ABD88C4E">
<mods:date id="024AF19B17FF19000ADD6E876C15B5FA">2024</mods:date>
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<mods:number id="17B0E5826AF3039526DF2DD4588622AE">2024-07-11</mods:number>
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<mods:number id="20CD891DB90EC4514E5C00B6050DCEF4">659</mods:number>
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<mods:number id="EB3A1FD4124B7B0A98FAD629B653C687">3</mods:number>
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<mods:classification id="3CAA5983EBFA80382C5B88A7D4F9323A">journal article</mods:classification>
<mods:identifier id="0F94009AF39B05793E72E7B9E58F70CB" type="CLB-Dataset">302193</mods:identifier>
<mods:identifier id="288775A7B9CB6F798DA828066E843999" type="DOI">10.11646/phytotaxa.659.3.1</mods:identifier>
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<mods:identifier id="E97DCDDDDB22F8F664128C16F51C46EB" type="Zenodo-Dep">13217894</mods:identifier>
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<treatment id="4B175324C955A514D1BAFF5CFD26A9C1" ID-DOI="http://doi.org/10.5281/zenodo.13652225" ID-GBIF-Taxon="240245555" ID-Zenodo-Dep="13652225" LSID="urn:lsid:plazi:treatment:4B175324C955A514D1BAFF5CFD26A9C1" httpUri="http://treatment.plazi.org/id/4B175324C955A514D1BAFF5CFD26A9C1" lastPageId="7" lastPageNumber="220" pageId="6" pageNumber="219">
<subSubSection id="8BA4B1B9C955A515D1BAFF5CFB17ABF5" box="[136,1266,158,185]" pageId="6" pageNumber="219" type="nomenclature">
<paragraph id="C301E232C955A515D1BAFF5CFB17ABF5" blockId="6.[136,1266,158,185]" box="[136,1266,158,185]" pageId="6" pageNumber="219">
<heading id="9849555EC955A515D1BAFF5CFB17ABF5" box="[136,1266,158,185]" fontSize="11" level="2" pageId="6" pageNumber="219" reason="4">
2.
<taxonomicName id="04BE99B1C955A515D196FF5CFBD4ABF5" authority="Liang Zhang, X.M. Zhou, T.T. Luong &amp; Li Bing Zhang" authorityName="Liang Zhang, X. M. Zhou, T. T. Luong &amp; Li Bing Zhang" authorityYear="2024" box="[164,1073,158,185]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="6" pageNumber="219" phylum="Tracheophyta" rank="species" species="daklakensis" status="sp. nov.">
<emphasis id="F1CA3E20C955A515D196FF5CFE5CABF4" bold="true" box="[164,441,158,185]" italics="true" pageId="6" pageNumber="219">Leptochilus daklakensis</emphasis>
Liang Zhang, X.M. Zhou, T.T. Luong &amp; Li Bing Zhang
</taxonomicName>
,
<emphasis id="F1CA3E20C955A515D50EFF5DFB74ABF5" box="[1084,1169,159,184]" italics="true" pageId="6" pageNumber="219">
<taxonomicNameLabel id="EAF9835BC955A515D50EFF5DFB74ABF5" box="[1084,1169,159,184]" pageId="6" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5B85FEB7C955A515D592FF5CFB00ABF5" box="[1184,1253,158,184]" captionStart="FIGURE 4" captionStartId="6.[136,229,1723,1745]" captionTargetBox="[163,1424,473,1687]" captionTargetId="figure-139@6.[151,1436,461,1699]" captionTargetPageId="6" captionText="FIGURE 4. Leptochilus daklakensis Liang Zhang, X.M. Zhou &amp; Li Bing Zhang,—A, B, C. Habit.—D.Abaxial view of fertile lamina.— E. Rhizome scales. Photo credit: Xin-Mao Zhou." figureDoi="http://doi.org/10.5281/zenodo.13217902" httpUri="https://zenodo.org/record/13217902/files/figure.png" pageId="6" pageNumber="219">Fig. 4</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C955A515D1BAFF24FCB2AA05" pageId="6" pageNumber="219" type="materials_examined">
<paragraph id="C301E232C955A515D1BAFF24FCB2AA05" blockId="6.[136,1452,230,437]" pageId="6" pageNumber="219">
<emphasis id="F1CA3E20C955A515D1BAFF24FF26AA4D" bold="true" box="[136,195,230,256]" pageId="6" pageNumber="219">Type</emphasis>
:
<collectingCountry id="BBA9A2A2C955A515D1E0FF24FED0AA4D" box="[210,309,230,256]" name="Vietnam" pageId="6" pageNumber="219">Vietnam</collectingCountry>
. Đắk Lắk: Chu Yang Sin National Park, Sub-area 1351,
<quantity id="04464FD7C955A515D28BFF24FBB2AA4C" box="[953,1111,230,257]" metricMagnitude="3" metricUnit="m" metricValue="1.165" metricValueMax="1.25" metricValueMin="1.08" pageId="6" pageNumber="219" unit="m" value="1165.0" valueMax="1250.0" valueMin="1080.0">1080–1250 m</quantity>
,
<geoCoordinate id="A68A84F5C955A515D556FF24FB27AA4D" box="[1124,1218,230,256]" degrees="12.41" direction="north" orientation="latitude" pageId="6" pageNumber="219" precision="555" value="12.41">12.41°N</geoCoordinate>
/
<geoCoordinate id="A68A84F5C955A515D5F9FF24FAD3AA4D" box="[1227,1334,230,256]" degrees="108.37" direction="east" orientation="longitude" pageId="6" pageNumber="219" precision="555" value="108.37">108.37°E</geoCoordinate>
, on rocks, small branch of Krông Kmar River,
<date id="B700C4F2C955A515D325FEC8FD31AA68" box="[535,724,266,293]" pageId="6" pageNumber="219" value="2015-10-30">30 October 2015</date>
,
<emphasis id="F1CA3E20C955A515D3ECFEC9FA4EAA69" box="[734,1451,266,292]" italics="true" pageId="6" pageNumber="219">Li Bing Zhang, Xin Mao Zhou, Thien Tam Luong, Minh Tri Dang</emphasis>
&amp;
<emphasis id="F1CA3E20C955A515D197FEECFE71AA05" box="[165,404,302,328]" italics="true" pageId="6" pageNumber="219">Bich Ha Duong 8944</emphasis>
(
<typeStatus id="1C055C90C955A515D090FEECFDE3AA05" box="[418,518,302,328]" pageId="6" pageNumber="219" type="holotype">holotype</typeStatus>
CDBI!,
<typeStatus id="1C055C90C955A515D355FEECFD26AA05" box="[615,707,302,328]" pageId="6" pageNumber="219" type="isotype">isotypes</typeStatus>
PHH, MO!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C955A514D18FFE90FE6BAA21" lastPageId="7" lastPageNumber="220" pageId="6" pageNumber="219" type="diagnosis">
<paragraph id="C301E232C955A515D18FFE90FC9CAAF9" blockId="6.[136,1452,230,437]" pageId="6" pageNumber="219">
<emphasis id="F1CA3E20C955A515D18FFE90FED5AA21" bold="true" box="[189,304,338,364]" pageId="6" pageNumber="219">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C955A515D00DFE90FDAEAA21" box="[319,587,338,364]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="6" pageNumber="219" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C955A515D00DFE90FDAEAA21" box="[319,587,338,364]" italics="true" pageId="6" pageNumber="219">Leptochilus daklakensis</emphasis>
</taxonomicName>
is distinguished from other species by its dimorphic fronds, fertile fronds two times longer than sterile fronds, sterile fronds with broadly winged petioles, petioles of fertile fronds four times longer than fertile laminae, and apex of fertile laminae abruptly narrowed.
</paragraph>
<caption id="97C1B2BAC955A515D1BAF979FDBBADBB" ID-DOI="http://doi.org/10.5281/zenodo.13217902" ID-Zenodo-Dep="13217902" httpUri="https://zenodo.org/record/13217902/files/figure.png" pageId="6" pageNumber="219" startId="6.[136,229,1723,1745]" targetBox="[163,1424,473,1687]" targetPageId="6" targetType="figure">
<paragraph id="C301E232C955A515D1BAF979FDBBADBB" blockId="6.[136,1451,1723,1782]" pageId="6" pageNumber="219">
<emphasis id="F1CA3E20C955A515D1BAF979FF12AD9D" bold="true" box="[136,247,1723,1745]" pageId="6" pageNumber="219">FIGURE 4</emphasis>
.
<taxonomicName id="04BE99B1C955A515D031F97EFC68AD9F" authority="Liang Zhang, X. M. Zhou &amp; Li Bing Zhang" authorityName="Liang Zhang, X. M. Zhou &amp; Li Bing Zhang" box="[259,909,1724,1746]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="6" pageNumber="219" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C955A515D031F97EFE0CAD9C" box="[259,489,1724,1745]" italics="true" pageId="6" pageNumber="219">Leptochilus daklakensis</emphasis>
Liang Zhang, X.M. Zhou &amp; Li Bing Zhang
</taxonomicName>
,—A, B, C. Habit.—D. Abaxial view of fertile lamina.— E. Rhizome scales. Photo credit: Xin-Mao Zhou.
</paragraph>
</caption>
<paragraph id="C301E232C955A514D18FF8E6FE6BAA21" blockId="6.[136,1452,1828,2071]" lastBlockId="7.[136,1452,158,652]" lastPageId="7" lastPageNumber="220" pageId="6" pageNumber="219">
<emphasis id="F1CA3E20C955A515D18FF8E6FEEDAC73" bold="true" box="[189,264,1828,1854]" pageId="6" pageNumber="219">Plants</emphasis>
perennial, evergreen, lithophytic.
<emphasis id="F1CA3E20C955A515D34DF8E6FD0CAC73" bold="true" box="[639,745,1828,1854]" pageId="6" pageNumber="219">Rhizome</emphasis>
long creeping, ca.
<quantity id="04464FD7C955A515D286F8E6FBDCAC73" box="[948,1081,1828,1855]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.5" metricValueMin="2.6" pageId="6" pageNumber="219" unit="mm" value="3.05" valueMax="3.5" valueMin="2.6">2.6–3.5 mm</quantity>
in diam., dorsiventrally flattened, densely scaly, scales lanceolate, 1.2–3.5 ×
<quantity id="04464FD7C955A515D35CF88AFD1DAC2F" box="[622,760,1864,1890]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="6" pageNumber="219" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4–0.7 mm</quantity>
, dark brown, basifixed, concolorous, clathrate, margin entire or sparely denticulate, apex acuminate.
<emphasis id="F1CA3E20C955A515D365F8AEFD49ACCB" bold="true" box="[599,684,1900,1926]" pageId="6" pageNumber="219">Fronds</emphasis>
dimorphic, fertile fronds usually much longer than sterile fronds, or rarely the same length as sterile fronds.
<emphasis id="F1CA3E20C955A515D345F852FCFEACE7" bold="true" box="[631,795,1936,1962]" pageId="6" pageNumber="219">Sterile fronds</emphasis>
(6–)
<quantity id="04464FD7C955A515D260F852FC20ACE7" box="[850,965,1936,1962]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="6" pageNumber="219" unit="cm" value="20.0" valueMax="22.0" valueMin="18.0">18–22 cm</quantity>
tall, sessile or shortly petiolated;
<emphasis id="F1CA3E20C955A515D47EF852FA4EACE7" bold="true" box="[1356,1451,1936,1962]" pageId="6" pageNumber="219">laminae</emphasis>
elliptic-lanceolate, base gradually narrowed, apex acuminate or cuspidate, (6–)
<quantity id="04464FD7C955A515D2CCF876FB8AAC83" box="[1022,1135,1972,1998]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="6" pageNumber="219" unit="cm" value="20.0" valueMax="22.0" valueMin="18.0">18–22 cm</quantity>
long,
<quantity id="04464FD7C955A515D58AF876FAD0AC83" box="[1208,1333,1972,1998]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="6" pageNumber="219" unit="cm" value="3.05" valueMax="3.2" valueMin="2.9">2.9–3.2 cm</quantity>
wide near middle, chartaceous, glabrous;
<emphasis id="F1CA3E20C955A515D0D7F81AFDDAACBF" bold="true" box="[485,575,2008,2034]" pageId="6" pageNumber="219">petioles</emphasis>
short, stramineous, grooved adaxially,
<quantity id="04464FD7C955A515D2C5F81AFB9BACBF" box="[1015,1150,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="6" pageNumber="219" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
in diam.,
<quantity id="04464FD7C955A515D5DCF81AFA8EACBF" box="[1262,1387,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="6" pageNumber="219" unit="cm" value="0.5" valueMax="0.7" valueMin="0.3">0.3–0.7 cm</quantity>
long, petiole base scaly; scales similar to rhizome scales but smaller, 1.3–1.5 ×
<quantity id="04464FD7C955A515D2F4F83EFBB6A35B" box="[966,1107,2044,2071]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="219" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
.
<emphasis id="F1CA3E20C955A515D56DF83EFAE1A35B" bold="true" box="[1119,1284,2044,2070]" pageId="6" pageNumber="219">Fertile fronds</emphasis>
(16)
<quantity id="04464FD7C955A515D40AF83EFA4EA35B" box="[1336,1451,2044,2071]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="6" pageNumber="219" unit="cm" value="38.0" valueMax="41.0" valueMin="35.0">35–41 cm</quantity>
tall, petiole much longer than lamina;
<emphasis id="F1CA3E20C954A514D307FF5CFD71ABF5" bold="true" box="[565,660,158,184]" pageId="7" pageNumber="220">laminae</emphasis>
deltate-lanceolate, margins entire, base cuneate, apex long acuminate, 5.4–9.0 cm long,
<quantity id="04464FD7C954A514D078FF00FE23AB90" box="[330,454,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="4.1" metricValueMin="2.2" pageId="7" pageNumber="220" unit="cm" value="3.15" valueMax="4.1" valueMin="2.2">2.2–4.1 cm</quantity>
wide bellow middle part, chartaceous, glabrous;
<emphasis id="F1CA3E20C954A514D2DAFF00FBA7AB91" bold="true" box="[1000,1090,194,220]" pageId="7" pageNumber="220">petioles</emphasis>
stramineous, grooved adaxially,
<quantity id="04464FD7C954A514D1BAFF24FEEAAA4C" box="[136,271,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="7" pageNumber="220" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
in diam., (11–)
<quantity id="04464FD7C954A514D084FF24FDCDAA4C" box="[438,552,230,257]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="7" pageNumber="220" unit="cm" value="29.5" valueMax="32.0" valueMin="27.0">27–32 cm</quantity>
long.
<emphasis id="F1CA3E20C954A514D35CFF24FD4BAA4D" bold="true" box="[622,686,230,256]" pageId="7" pageNumber="220">Veins</emphasis>
anastomosing, distinct adaxially and abaxially, veinlets visible.
<emphasis id="F1CA3E20C954A514D448FF24FA49AA4D" bold="true" box="[1402,1452,230,256]" pageId="7" pageNumber="220">Sori</emphasis>
linear, reach lamina margins, ca.
<quantity id="04464FD7C954A514D337FEC8FDB8AA68" box="[517,605,266,293]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="220" unit="mm" value="0.8">0.8 mm</quantity>
from margins and
<quantity id="04464FD7C954A514D208FEC8FC4AAA68" box="[826,943,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="7" pageNumber="220" unit="mm" value="1.75" valueMax="2.5" valueMin="1.0">1–2.5 mm</quantity>
from midrib, 15–23 on each side of midrib,
<quantity id="04464FD7C954A514D1BAFEECFEE0AA04" box="[136,261,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.9" metricValueMin="1.2" pageId="7" pageNumber="220" unit="cm" value="2.05" valueMax="2.9" valueMin="1.2">1.2–2.9 cm</quantity>
long,
<quantity id="04464FD7C954A514D07EFEECFE36AA04" box="[332,467,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="7" pageNumber="220" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.4–0.6 mm</quantity>
wide when mature, at angles of 42–47 degree with midribs, centers ca. (1–)1.5–3.0 mm apart from one another.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C954A514D18FFEB4FCB5AAF9" pageId="7" pageNumber="220" type="distribution">
<paragraph id="C301E232C954A514D18FFEB4FCB5AAF9" blockId="7.[136,1452,158,652]" pageId="7" pageNumber="220">
<emphasis id="F1CA3E20C954A514D18FFEB4FE09AADD" bold="true" box="[189,492,374,400]" pageId="7" pageNumber="220">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C954A514D0C6FEB4FCE7AADD" box="[500,770,374,400]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C954A514D0C6FEB4FCE7AADD" box="[500,770,374,400]" italics="true" pageId="7" pageNumber="220">Leptochilus daklakensis</emphasis>
</taxonomicName>
is currently known from south Đắk Lắk Province,
<collectingCountry id="BBA9A2A2C954A514D474FEB4FA4CAADD" box="[1350,1449,374,400]" name="Vietnam" pageId="7" pageNumber="220">Vietnam</collectingCountry>
. The species was observed only growing on rocks in rainforests.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C954A514D18FFE7CFD26A9C1" pageId="7" pageNumber="220" type="etymology">
<paragraph id="C301E232C954A514D18FFE7CFC30AAB1" blockId="7.[136,1452,158,652]" pageId="7" pageNumber="220">
<emphasis id="F1CA3E20C954A514D18FFE7CFEDBAA95" bold="true" box="[189,318,446,472]" pageId="7" pageNumber="220">Etymology</emphasis>
: The species epithet is from the Vietnamese name,
<emphasis id="F1CA3E20C954A514D24AFE7CFC26AA95" box="[888,963,446,472]" italics="true" pageId="7" pageNumber="220">daklak</emphasis>
, the name of the province in South
<collectingCountry id="BBA9A2A2C954A514D47FFE7CFA49AA95" box="[1357,1452,446,472]" name="Vietnam" pageId="7" pageNumber="220">Vietnam</collectingCountry>
where the
<typeStatus id="1C055C90C954A514D1CFFE21FEC8AAB0" box="[253,301,483,509]" pageId="7" pageNumber="220">type</typeStatus>
was collected, and the Latin suffix
<emphasis id="F1CA3E20C954A514D38EFE21FD18AAB1" box="[700,765,483,508]" italics="true" pageId="7" pageNumber="220">-ensis</emphasis>
, of origin or place.
</paragraph>
<paragraph id="C301E232C954A514D18FFDC4FD26A9C1" blockId="7.[136,1452,158,652]" pageId="7" pageNumber="220">
<emphasis id="F1CA3E20C954A514D18FFDC4FEE5A96D" bold="true" box="[189,256,518,544]" pageId="7" pageNumber="220">Notes</emphasis>
:
<taxonomicName id="04BE99B1C954A514D03DFDC4FDFFA96D" box="[271,538,518,544]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C954A514D03DFDC4FDFFA96D" box="[271,538,518,544]" italics="true" pageId="7" pageNumber="220">Leptochilus daklakensis</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="04BE99B1C954A514D3D8FDC5FCAAA96D" authorityName="Copeland" authorityYear="1914" box="[746,847,519,544]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C954A514D3D8FDC5FCAAA96D" box="[746,847,519,544]" italics="true" pageId="7" pageNumber="220">L. ovatus</emphasis>
</taxonomicName>
but differs the latter in having broadly winged petioles (vs. narrowly wringed petioles, or sometimes petioles wingless), apex of fertile laminae abruptly narrowed (vs. apex of fertile laminae rounded or gradually narrowed). Phylogenetically,
<taxonomicName id="04BE99B1C954A514D2B6FD8DFBC6A925" box="[900,1059,590,616]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="daklakensis">
<emphasis id="F1CA3E20C954A514D2B6FD8DFBC6A925" box="[900,1059,590,616]" italics="true" pageId="7" pageNumber="220">L. daklakensis</emphasis>
</taxonomicName>
was resolved as an isolated lineage quite distant from
<taxonomicName id="04BE99B1C954A514D064FDB1FE58A9C1" authorityName="Copeland" authorityYear="1914" box="[342,445,627,652]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C954A514D064FDB1FE58A9C1" box="[342,445,627,652]" italics="true" pageId="7" pageNumber="220">L. ovatus</emphasis>
</taxonomicName>
in the
<emphasis id="F1CA3E20C954A514D338FDB1FD9CA9C1" box="[522,633,626,652]" italics="true" pageId="7" pageNumber="220">
L.
<taxonomicName id="04BE99B1C954A514D315FDB0FD9CA9C1" authorityName="K.B.Presl" authorityYear="1851" box="[551,633,626,652]" class="Polypodiopsida" family="Polypodiaceae" genus="Colysis" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="220" phylum="Tracheophyta" rank="genus">Colysis</taxonomicName>
</emphasis>
clade.
</paragraph>
</subSubSection>
</treatment>
</document>
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