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<document id="E8CE6602002F4F4A419BAFD609079CF6" ID-DOI="10.26107/RBZ-2022-0023" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174433" ID-ZooBank="E4F49C16-DB5E-4593-AB8E-03D8859A59D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665290687137" checkinUser="felipe" docAuthor="Páll-Gergely, Barna &amp; Hunyadi, András" docDate="2022" docId="03B887D723032671ADAF30FA215AFC92" docLanguage="en" docName="RafflesBZool.70.417439.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Clostophis bactrianus Páll-Gergely &amp; Hunyadi 2022, new species" docType="treatment" docVersion="4" lastPageNumber="427" masterDocId="FF81FFAF2305267BAE6F347A2327FFA7" masterDocTitle="New and little-known species of Clostophis Benson, 1860 from Southeast Asia (Gastropoda: Eupulmonata: Hypselostomatidae)" masterLastPageNumber="439" masterPageNumber="417" pageNumber="423" updateTime="1699379781661" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="51C1B9E4429B985004690C5B73FB04F7">New and little-known species of Clostophis Benson, 1860 from Southeast Asia (Gastropoda: Eupulmonata: Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="0013747CECB47166447C1187C0D81DEF" type="personal">
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<mods:namePart id="8B99C7F7C84223DC9BCE3CE0A7608C24">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="342226229AEE1F2F2B6A2F2B4EABBE10">Hunyadi, András</mods:namePart>
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<mods:title id="53F13BD05220F427E5DF0B7AFDC8AA2B">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="3EDB1C5D9F92A9E41A0E3506F87244EB">
<mods:date id="5975EAD71AB6BB722270402BCB4505D6">2022</mods:date>
<mods:detail id="1C83251FBEEA98DC7E704E5B9BBB6738" type="pubDate">
<mods:number id="5F6841E22DB2E400D070788153B462EE">2022-09-09</mods:number>
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<mods:number id="6E270F8FFC153ACC02090302E10C896B">70</mods:number>
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<mods:identifier id="DEC54BE5D754F90F242D9933B4317A7D" type="DOI">10.26107/RBZ-2022-0023</mods:identifier>
<mods:identifier id="3FDCF900C1DFE89396CF4CB7D0F0D996" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="762BF4ED0DF968841E212F1F42967D3F" type="Zenodo-Dep">7174433</mods:identifier>
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<treatment id="03B887D723032671ADAF30FA215AFC92" ID-DOI="http://doi.org/10.5281/zenodo.7171119" ID-Zenodo-Dep="7171119" LSID="urn:lsid:plazi:treatment:03B887D723032671ADAF30FA215AFC92" httpUri="http://treatment.plazi.org/id/03B887D723032671ADAF30FA215AFC92" lastPageId="10" lastPageNumber="427" pageId="6" pageNumber="423">
<subSubSection id="C30B654A2303267DADAF30FA2661FB3F" box="[960,1350,1152,1176]" pageId="6" pageNumber="423" type="nomenclature">
<paragraph id="8BAE36C12303267DADAF30FA2661FB3F" blockId="6.[960,1350,1152,1208]" box="[960,1350,1152,1176]" pageId="6" pageNumber="423">
<heading id="D0E681AD2303267DADAF30FA2661FB3F" bold="true" box="[960,1350,1152,1176]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="423" reason="2">
<emphasis id="B965EAD32303267DADAF30FA2661FB3F" bold="true" box="[960,1350,1152,1176]" pageId="6" pageNumber="423">
<taxonomicName id="4C114D422303267DADAF30FA2796FB3F" ID-CoL="a846bde7-f20a-445e-9840-48a3c2fd5d29" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[960,1201,1152,1176]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="423" phylum="Mollusca" rank="species" species="bactrianus" status="sp. nov.">
<emphasis id="B965EAD32303267DADAF30FA2796FB3F" bold="true" box="[960,1201,1152,1176]" italics="true" pageId="6" pageNumber="423">Clostophis bactrianus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A82303267DAAAF30FA2661FB3F" box="[1216,1350,1152,1176]" pageId="6" pageNumber="423" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B654A2303267DAA3530DA278BFB1F" box="[1114,1196,1184,1208]" pageId="6" pageNumber="423" type="description">
<paragraph id="8BAE36C12303267DAA3530DA278BFB1F" blockId="6.[960,1350,1152,1208]" box="[1114,1196,1184,1208]" pageId="6" pageNumber="423">
(
<figureCitation id="132A2A442303267DAA0D30DA2785FB1F" box="[1122,1186,1184,1208]" captionStart="Fig" captionStartId="9.[128,163,2093,2115]" captionTargetBox="[128,1437,189,2077]" captionTargetId="figure-10@9.[128,1437,188,2079]" captionTargetPageId="9" captionText="Fig. 8. Clostophis bactrianus, new species (holotype, HNHM 105324)." figureDoi="http://doi.org/10.5281/zenodo.7174451" httpUri="https://zenodo.org/record/7174451/files/figure.png" pageId="6" pageNumber="423">Fig. 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B654A2303267DAD2C309A20C8FA3F" pageId="6" pageNumber="423" type="materials_examined">
<paragraph id="8BAE36C12303267DAD2C309A20C8FA3F" blockId="6.[835,1474,1248,1432]" pageId="6" pageNumber="423">
<emphasis id="B965EAD32303267DAD2C309A20DFFB5F" bold="true" box="[835,1016,1248,1272]" pageId="6" pageNumber="423">Type material.</emphasis>
<materialsCitation id="3B793C9C2303267DAA6C309A20DFFADF" collectingDate="2013-01-19" collectionCode="HNHM" collectorName="A. Hunyadi" county="Gua Bama" elevation="120" latitude="4.1942" location="Padang Tungku" longLatPrecision="1" longitude="101.9656" municipality="Kuala Lipis" pageId="6" pageNumber="423" specimenCode="HNHM 105324" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<typeStatus id="54AA88632303267DAA6C309A274BFB5F" box="[1027,1132,1248,1272]" pageId="6" pageNumber="423" type="holotype">Holotype</typeStatus>
(SH:
<quantity id="4CE99B242303267DAAD7309A2633FB5F" box="[1208,1300,1248,1272]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="423" unit="mm" value="1.1">1.1 mm</quantity>
, SW:
<quantity id="4CE99B242303267DAB0F309A269DFB5F" box="[1376,1466,1248,1272]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="423" unit="mm" value="1.6">1.6 mm</quantity>
) (
<specimenCode id="DBB79EBA2303267DAD25317A20D9FABF" box="[842,1022,1280,1304]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="423">HNHM 105324</specimenCode>
),
<collectingRegion id="49D5F8232303267DAA7A317A274FFABF" box="[1045,1128,1280,1304]" country="Malaysia" name="Pahang" pageId="6" pageNumber="423">Pahang</collectingRegion>
,
<collectingCounty id="62CF4E4D2303267DAA19317A27C8FABF" box="[1142,1263,1280,1304]" pageId="6" pageNumber="423">Gua Bama</collectingCounty>
,
<quantity id="4CE99B242303267DAA91317A2610FABF" box="[1278,1335,1280,1304]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="423" unit="km" value="9.0">9 km</quantity>
from
<collectingMunicipality id="6BCAACBB2303267DABEE317A205BFA9F" pageId="6" pageNumber="423">Kuala Lipis</collectingMunicipality>
toward
<location id="8ECE601A2303267DADB0315A27B3FA9F" LSID="urn:lsid:plazi:treatment:03B887D723032671ADAF30FA215AFC92:8ECE601A2303267DADB0315A27B3FA9F" box="[991,1172,1312,1336]" county="Gua Bama" latitude="4.1942" longLatPrecision="1" longitude="101.9656" municipality="Kuala Lipis" name="Padang Tungku" pageId="6" pageNumber="423" stateProvince="Pahang">Padang Tungku</location>
,
<elevation id="003CD1F22303267DAACD315A260BFA9F" box="[1186,1324,1312,1336]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="423" unit="m" value="120.0">
<quantity id="4CE99B242303267DAACD315A27CCFA9F" box="[1186,1259,1312,1336]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="423" unit="m" value="120.0">120 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE2550062303267DAB53315A2699FA9F" box="[1340,1470,1311,1336]" degrees="4" direction="north" minutes="11.652" orientation="latitude" pageId="6" pageNumber="423" precision="1" value="4.1942">4°11.652N</geoCoordinate>
,
<geoCoordinate id="EE2550062303267DAD2B313A20F0FAFF" box="[836,983,1343,1368]" degrees="101" direction="east" minutes="57.936" orientation="longitude" pageId="6" pageNumber="423" precision="1" value="101.9656">101°57.936E</geoCoordinate>
(locality code: 2013/28), coll.
<collectorName id="26E453172303267DAB56313A2699FAFF" box="[1337,1470,1344,1368]" pageId="6" pageNumber="423">A. Hunyadi</collectorName>
,
<date id="FFAF10012303267DAD2C311A20DFFADF" box="[835,1016,1376,1401]" pageId="6" pageNumber="423" value="2013-01-19">
<collectingDate id="EFEBE9E92303267DAD2C311A20DFFADF" box="[835,1016,1376,1401]" pageId="6" pageNumber="423" value="2013-01-19">19 January 2013</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="54AA88632303267DAA6D311B2749FADE" box="[1026,1134,1377,1401]" pageId="6" pageNumber="423" type="paratype">Paratypes</typeStatus>
: 24 shells (coll. HA), same data as for holotype.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A2303267DAD2C31BA20ECF9BF" pageId="6" pageNumber="423" type="diagnosis">
<paragraph id="8BAE36C12303267DAD2C31BA20ECF9BF" blockId="6.[835,1475,1472,1560]" pageId="6" pageNumber="423">
<emphasis id="B965EAD32303267DAD2C31BA209FFA7F" bold="true" box="[835,952,1472,1496]" pageId="6" pageNumber="423">Diagnosis.</emphasis>
Uniquely characterised within the genus by the two low, blunt teeth on the columellar and basal side of the aperture.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A23032671AD2C323A23FBFE12" lastPageId="10" lastPageNumber="427" pageId="6" pageNumber="423" type="description">
<paragraph id="8BAE36C123032671AD2C323A22FAFEF2" blockId="6.[834,1477,1600,2041]" lastBlockId="10.[136,776,189,341]" lastPageId="10" lastPageNumber="427" pageId="6" pageNumber="423">
<emphasis id="B965EAD32303267DAD2C323A20F5F9FF" bold="true" box="[835,978,1600,1624]" pageId="6" pageNumber="423">Description.</emphasis>
Shell conical, wider than high, apex blunt. Whorls rounded or slightly shouldered. Last whorl not detached from penultimate whorl, although aperture may be slightly protruding (parietal callus elevated from penultimate whorl). Protoconch consisting of 1.5 whorls, spirally striated except for the first ca. 0.25 whorl. Entire shell with 4.25 whorls. Umbilicus relatively narrow, less than one third of shell width. Entire teleoconch with strong, elevated, rather equidistant spiral striae (ca. 2022 on body whorl in apertural view), and some weak, irregular radial growth lines. Aperture almost heart-shaped due to large sinulus. Parietal tooth pronounced, relatively short, reaches peristome. Palatal tooth also strong, elevated, but blunt. Sinulus clearly separated from rest of aperture due to strong parietal and palatal teeth. Two low, blunt denticles situated next to each other on the peristome on the basal and columellar sides of the aperture; the lower one is situated slightly left (i.e., direction of the columella) from the basal most point, whereas the upper one sits on the columellar side.
</paragraph>
<caption id="DF6E66492302267CAEEE3C57275BF7E5" ID-DOI="http://doi.org/10.5281/zenodo.7174445" ID-Zenodo-Dep="7174445" box="[129,1148,2092,2115]" httpUri="https://zenodo.org/record/7174445/files/figure.png" pageId="7" pageNumber="424" startId="7.[129,164,2093,2115]" targetBox="[128,1435,187,2077]" targetPageId="7">
<paragraph id="8BAE36C12302267CAEEE3C57275BF7E5" blockId="7.[129,1148,2092,2115]" box="[129,1148,2092,2115]" pageId="7" pageNumber="424">
Fig. 6.
<taxonomicName id="4C114D422302267CAEA53C572196F7E4" authority="(van Benthem Jutting, 1961)" baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1961" box="[202,689,2093,2115]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="424" phylum="Mollusca" rank="species" species="neglectus">
<emphasis id="B965EAD32302267CAEA53C5722B4F7E5" box="[202,403,2093,2114]" italics="true" pageId="7" pageNumber="424">Clostophis neglectus</emphasis>
(van
<bibRefCitation id="EF804B302302267CAFA43C57218DF7E4" author="Benthem Jutting WSS van" box="[459,682,2093,2115]" pageId="7" pageNumber="424" pagination="34 - 48" refId="ref6425" refString="Benthem Jutting WSS van (1961) Additional new species and new localities of the family Vertiginidae and the genera Oophana and Opisthostoma from Malaya. Bulletin of the Raffles Museum, 26: 34 - 48." type="journal article" year="1961">Benthem Jutting, 1961</bibRefCitation>
)
</taxonomicName>
(syntype of
<taxonomicName id="4C114D422302267CAD5E3C572722F7E5" authority=": NHMUK" authorityName="NHMUK" box="[817,1029,2093,2114]" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="424" phylum="Mollusca" rank="species" species="laidlawi">
<emphasis id="B965EAD32302267CAD5E3C5720BBF7E5" box="[817,924,2093,2114]" italics="true" pageId="7" pageNumber="424">C. laidlawi</emphasis>
: NHMUK
</taxonomicName>
20191105).
</paragraph>
</caption>
<caption id="DF6E6649230D2673AEF93C5727DBF7E5" ID-DOI="http://doi.org/10.5281/zenodo.7174449" ID-Zenodo-Dep="7174449" box="[150,1276,2093,2115]" httpUri="https://zenodo.org/record/7174449/files/figure.png" pageId="8" pageNumber="425" startId="8.[150,185,2093,2115]" targetBox="[150,1459,187,2077]" targetPageId="8">
<paragraph id="8BAE36C1230D2673AEF93C5727DBF7E5" blockId="8.[150,1276,2093,2115]" box="[150,1276,2093,2115]" pageId="8" pageNumber="425">
Fig. 7.
<taxonomicName id="4C114D42230D2673AEB03C5721E1F7E4" authority="(van Benthem Jutting, 1961)" baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1961" box="[223,710,2093,2115]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="425" phylum="Mollusca" rank="species" species="neglectus">
<emphasis id="B965EAD3230D2673AEB03C57228EF7E5" box="[223,425,2093,2114]" italics="true" pageId="8" pageNumber="425">Clostophis neglectus</emphasis>
(van
<bibRefCitation id="EF804B30230D2673AF8E3C572198F7E4" author="Benthem Jutting WSS van" box="[481,703,2093,2115]" pageId="8" pageNumber="425" pagination="34 - 48" refId="ref6425" refString="Benthem Jutting WSS van (1961) Additional new species and new localities of the family Vertiginidae and the genera Oophana and Opisthostoma from Malaya. Bulletin of the Raffles Museum, 26: 34 - 48." type="journal article" year="1961">Benthem Jutting, 1961</bibRefCitation>
)
</taxonomicName>
from locality 2013/33. A, specimen 2; BH, specimen 1.
</paragraph>
</caption>
<caption id="DF6E6649230C2672AEEF3C57201EF7E5" ID-DOI="http://doi.org/10.5281/zenodo.7174451" ID-Zenodo-Dep="7174451" box="[128,825,2092,2115]" httpUri="https://zenodo.org/record/7174451/files/figure.png" pageId="9" pageNumber="426" startId="9.[128,163,2093,2115]" targetBox="[128,1437,189,2077]" targetPageId="9">
<paragraph id="8BAE36C1230C2672AEEF3C57201EF7E5" blockId="9.[128,825,2092,2115]" box="[128,825,2092,2115]" pageId="9" pageNumber="426">
Fig. 8.
<taxonomicName id="4C114D42230C2672AEA63C5722BAF7E5" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[201,413,2093,2114]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="426" phylum="Mollusca" rank="species" species="bactrianus" status="sp. nov.">
<emphasis id="B965EAD3230C2672AEA63C5722BAF7E5" box="[201,413,2093,2114]" italics="true" pageId="9" pageNumber="426">Clostophis bactrianus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A8230C2672AFC53C572138F7E4" box="[426,543,2093,2115]" pageId="9" pageNumber="426" rank="species">new species</taxonomicNameLabel>
(holotype, HNHM 105324).
</paragraph>
</caption>
<paragraph id="8BAE36C1230F2671AEE7350723FBFE12" blockId="10.[136,776,381,437]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AEE7350722BEFE32" bold="true" box="[136,409,381,405]" pageId="10" pageNumber="427">Measurements (in mm).</emphasis>
SH = 0.941.05, SW = 1.031.06 (n = 4).
</paragraph>
</subSubSection>
<subSubSection id="C30B654A230F2671AEE735A72122FD32" pageId="10" pageNumber="427" type="diagnosis">
<paragraph id="8BAE36C1230F2671AEE735A72122FD32" blockId="10.[135,775,477,661]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AEE735A722A4FE52" bold="true" box="[136,387,477,501]" pageId="10" pageNumber="427">Differential diagnosis.</emphasis>
<taxonomicName id="4C114D42230F2671AFE635A72154FE52" authority="Páll-Gergely &amp; Hunyadi, 2022" authorityName="Páll-Gergely &amp; Hunyadi" authorityYear="2022" box="[393,627,477,501]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="427" phylum="Mollusca" rank="species" species="bactrianus" status="sp. nov.">
<emphasis id="B965EAD3230F2671AFE635A72154FE52" box="[393,627,477,501]" italics="true" pageId="10" pageNumber="427">Clostophis bactrianus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25657A8230F2671ACEF35A42024FE52" box="[640,771,477,501]" pageId="10" pageNumber="427" rank="species">new species</taxonomicNameLabel>
, differs from
<taxonomicName id="4C114D42230F2671AF60358422B3FDB2" baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1961" box="[271,404,509,533]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="427" phylum="Mollusca" rank="species" species="neglectus">
<emphasis id="B965EAD3230F2671AF60358422B3FDB2" box="[271,404,509,533]" italics="true" pageId="10" pageNumber="427">C. neglectus</emphasis>
</taxonomicName>
by the smaller shell size, the more elevated parietal tooth, the more basally-situated palatal tooth (and as a result, a larger sinulus), and the two low denticles on the columellar and basal sides of the aperture.
<taxonomicName id="4C114D42230F2671ACFB362723F8FD32" baseAuthorityName="Collinge" baseAuthorityYear="1902" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="427" phylum="Mollusca" rank="species" species="laidlawi">
<emphasis id="B965EAD3230F2671ACFB362723F8FD32" italics="true" pageId="10" pageNumber="427">Clostophis laidlawi</emphasis>
</taxonomicName>
has five apertural barriers.
</paragraph>
</subSubSection>
<subSubSection id="C30B654A230F2671AEE736C7215AFC92" pageId="10" pageNumber="427" type="etymology">
<paragraph id="8BAE36C1230F2671AEE736C7215AFC92" blockId="10.[136,776,701,821]" pageId="10" pageNumber="427">
<emphasis id="B965EAD3230F2671AEE736C7222DFD72" bold="true" box="[136,266,701,725]" pageId="10" pageNumber="427">Etymology.</emphasis>
The specific epithet derives from the scientific name of the Bactrian camel (
<taxonomicName id="4C114D42230F2671AFAF36A723E5FCB2" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Mammalia" family="Camelidae" genus="Camelus" kingdom="Animalia" order="Artiodactyla" pageId="10" pageNumber="427" phylum="Chordata" rank="species" species="bactrianus">
<emphasis id="B965EAD3230F2671AFAF36A721B0FD52" box="[448,663,733,757]" italics="true" pageId="10" pageNumber="427">Camelus bactrianus</emphasis>
Linnaeus, 1758
</taxonomicName>
). The two blunt columellar teeth of this new species resemble the two humps on the camels back.
</paragraph>
</subSubSection>
</treatment>
</document>