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<document id="4A85F4D03BF2574017E355EE9B0AF6F1" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FFE9FFB2FF6871B4DA78FD02" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Tetrabothrus borneensis CAMERON 1943" docType="treatment" docVersion="11" lastPageNumber="102" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="101" updateTime="1699235685730" updateUser="ExternalLinkService">
<mods:mods id="92070FB276365C2A4168DFD7E9DDA048" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B79F43E72FDEAD4581D812D9044FC685">
<mods:title id="AC827A39D99B9663319EBCC643AE5A24">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="7A183A78D0A0B9BD1CC79232B5728512" type="personal">
<mods:role id="C95E252B202D882A04D1D0EEC10372B0">
<mods:roleTerm id="1DCE4F8CD3A22820C558A3A90382E48E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F0628E35B33590D28FB0571B2D01206">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="B2AD998CF57B2C05061341120D491069">text</mods:typeOfResource>
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<mods:title id="C676C498B9BEB31115AAD19A065B9D60">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="A370B698BEBA3E41807309DBE1944299">
<mods:date id="1D5E5BF819F32B1533169A0136B96FA1">2016</mods:date>
<mods:detail id="91D6373251A04A8086D8766674029074" type="pubDate">
<mods:number id="6847914CAEFF37E61FA1C7601709F95E">2016-12-20</mods:number>
</mods:detail>
<mods:detail id="A243994D50E6132FB863FCAB93F67068" type="volume">
<mods:number id="2AD018CA6B5AB2A50145E4B8919D4F63">66</mods:number>
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<mods:number id="771E20D26BE1A77E733A20A7EEDEEF2D">1</mods:number>
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<mods:url id="3EC300C23841D6F51A4DFD4FA5A6F761">https://www.contributions-to-entomology.org/article/view/1904</mods:url>
</mods:location>
<mods:classification id="95F6EAA271AEF59BC433615E7CA7F865">journal article</mods:classification>
<mods:identifier id="5E295021F203D917E67F8F854473CB36" type="CLB-Dataset">2419</mods:identifier>
<mods:identifier id="A4788E14CCF2A9A38F6EF6F9AE0E9944" type="DOI">10.21248/contrib.entomol.66.1.13-111</mods:identifier>
<mods:identifier id="B29F5203DA4D4CA5804D3181E77443E1" type="GBIF-Dataset">d2933fad-1cac-4a5a-a558-2633f8314541</mods:identifier>
<mods:identifier id="4767643E49D93BC97B2674428633E5EC" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="8DC037993E4E73D2C17A6C20D9C66A0D" type="Zenodo-Dep">6421171</mods:identifier>
</mods:mods>
<treatment id="766F7C36FFE9FFB2FF6871B4DA78FD02" ID-DOI="http://doi.org/10.5281/zenodo.5879529" ID-GBIF-Taxon="192602751" ID-Zenodo-Dep="5879529" LSID="urn:lsid:plazi:treatment:766F7C36FFE9FFB2FF6871B4DA78FD02" httpUri="http://treatment.plazi.org/id/766F7C36FFE9FFB2FF6871B4DA78FD02" lastPageId="89" lastPageNumber="102" pageId="88" pageNumber="101">
<subSubSection id="B6DC9EABFFE9FFB3FF6871B4DAFEFB2D" box="[227,646,1171,1198]" pageId="88" pageNumber="101" type="nomenclature">
<paragraph id="FE79CD20FFE9FFB3FF6871B4DAFEFB2D" blockId="88.[227,646,1171,1231]" box="[227,646,1171,1198]" pageId="88" pageNumber="101">
<heading id="A5317A4CFFE9FFB3FF6871B4DAFEFB2D" box="[227,646,1171,1198]" centered="true" fontSize="11" level="2" pageId="88" pageNumber="101" reason="3">
<taxonomicName id="39C6B6A3FFE9FFB3FF6871B4DAFEFB2D" ID-CoL="55L23" authority="CAMERON, 1943" authorityName="CAMERON" authorityYear="1943" box="[227,646,1171,1198]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="CCB21132FFE9FFB3FF6871B4D9A3FB2D" box="[227,475,1171,1198]" italics="true" pageId="88" pageNumber="101">Tetrabothrus borneensis</emphasis>
<bibRefCitation id="9A57B0D1FFE9FFB3FE6A71B4DAFEFB2D" author="CAMERON, M." box="[481,646,1171,1198]" pageId="88" pageNumber="101" pagination="139 - 143" refId="ref77169" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON, 1943</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE9FFB3FF68719ED9CEFB4C" box="[227,438,1209,1231]" pageId="88" pageNumber="101" type="description">
<paragraph id="FE79CD20FFE9FFB3FF68719ED9CEFB4C" blockId="88.[227,646,1171,1231]" box="[227,438,1209,1231]" pageId="88" pageNumber="101">
(
<figureCitation id="66FDD1A5FFE9FFB3FF6C719ED925FB4C" box="[231,349,1209,1231]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="88" pageNumber="101">Figs 160–162</figureCitation>
,
<figureCitation id="66FDD1A5FFE9FFB3FEEE719ED9C9FB4C" box="[357,433,1209,1231]" captionStart="Figs 442–458" captionStartId="31.[151,191,1971,1993]" captionTargetBox="[155,1398,240,1950]" captionTargetId="figure-119@31.[151,1398,232,1971]" captionText="Figs 442–458. Tetrabothrus collucatus (442–443), T. borneensis (444–449), T. punctiventris (450–451), T. sulawesicus (452–454), Zyras illecebrosus, paratype (455–456), and Z. quasar, holotype (457–458): median lobe of aedeagus in lateral and in ventral view (442–445, 450–453, 455–458); apico-ventral portion of median lobe in lateral view (446–448); male sternite VIII (449, 454). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421226" httpUri="https://zenodo.org/record/6421226/files/figure.png" pageId="88" pageNumber="101">444–449</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE9FFB3FF3671CCDA0AFA83" box="[189,626,1259,1280]" pageId="88" pageNumber="101" type="reference_group">
<paragraph id="FE79CD20FFE9FFB3FF3671CCDA0AFA83" blockId="88.[189,626,1259,1280]" box="[189,626,1259,1280]" pageId="88" pageNumber="101">
<treatmentCitationGroup id="DED6EA0EFFE9FFB3FF3671CCDA0AFA83" box="[189,626,1259,1280]" pageId="88" pageNumber="101">
<treatmentCitation id="7F67EB31FFE9FFB3FF3671CCDA16FA83" author="CAMERON, M." box="[189,622,1259,1280]" page="140" pageId="88" pageNumber="101" year="1943">
<taxonomicName id="39C6B6A3FFE9FFB3FF3671CCDA16FA83" ID-CoL="55L23" authority="CAMERON, 1943: 140" authorityName="CAMERON" authorityPageNumber="140" authorityYear="1943" box="[189,622,1259,1280]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="CCB21132FFE9FFB3FF3671CCD9E5FA83" box="[189,413,1259,1280]" italics="true" pageId="88" pageNumber="101">Tetrabothrus borneensis</emphasis>
<bibRefCitation id="9A57B0D1FFE9FFB3FE2971CCDA16FA83" author="CAMERON, M." box="[418,622,1259,1280]" pageId="88" pageNumber="101" pagination="139 - 143" refId="ref77169" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON, 1943: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE9FFB3FF36700EDAA9F942" pageId="88" pageNumber="101" type="materials_examined">
<paragraph id="FE79CD20FFE9FFB3FF36700EDA6EFA22" blockId="88.[189,794,1321,1441]" pageId="88" pageNumber="101">
<emphasis id="CCB21132FFE9FFB3FF36700ED9B3FAC2" bold="true" box="[189,459,1321,1345]" pageId="88" pageNumber="101">Type material examined</emphasis>
:
<materialsCitation id="4EAEC77DFFE9FFB3FE50700EDA6EFA22" ID-GBIF-Occurrence="3455118319" collectionCode="BMNH" collectorName="M. Cameron. Bequest." determinerName="R. G. Booth" location="Mt Poi" pageId="88" pageNumber="101" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="217D7382FFE9FFB3FE50700EDA46FAC2" box="[475,574,1321,1345]" pageId="88" pageNumber="101" type="holotype">Holotype</typeStatus>
<specimenCount id="E8C006A9FFE9FFB3FDC3700DDA25FAC2" box="[584,605,1322,1345]" pageId="88" pageNumber="101" type="female">♀</specimenCount>
: “
<location id="FB199BFBFFE9FFB3FDFE700EDABBFAC2" LSID="urn:lsid:plazi:treatment:766F7C36FFE9FFB2FF6871B4DA78FD02:FB199BFBFFE9FFB3FDFE700EDABBFAC2" box="[629,707,1321,1345]" name="Mt Poi" pageId="88" pageNumber="101">Mt Poi</location>
, 5000' /
<emphasis id="CCB21132FFE9FFB3FF36706DD892FAE2" box="[189,234,1354,1377]" italics="true" pageId="88" pageNumber="101">TeT.</emphasis>
<taxonomicName id="39C6B6A3FFE9FFB3FF64706ED9E5FAE2" authority="Cam." authorityName="CAMERON" authorityYear="1943" box="[239,413,1353,1377]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="CCB21132FFE9FFB3FF64706ED925FAE2" box="[239,349,1353,1377]" italics="true" pageId="88" pageNumber="101">borneensis</emphasis>
Cam.
</taxonomicName>
Type / M. Cameron. Bequest. 1955- 147 / Holotype / Holotype
<taxonomicName id="39C6B6A3FFE9FFB3FE50704EDB6EFA02" authority="Cam." authorityName="CAMERON" authorityYear="1943" box="[475,790,1385,1409]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="CCB21132FFE9FFB3FE50704EDAADFA02" box="[475,725,1385,1409]" italics="true" pageId="88" pageNumber="101">Tetrabothrus borneensis</emphasis>
Cam.
</taxonomicName>
, det.
<determinerName id="0BA521E4FFE9FFB3FF6070AED91BFA22" box="[235,355,1417,1441]" pageId="88" pageNumber="101">R.G. Booth</determinerName>
2015” (
<collectionCode id="98D755E5FFE9FFB3FE3E70AEDA73FA22" box="[437,523,1417,1441]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="88" pageNumber="101">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="FE79CD20FFE9FFB3FF3670EEDAA9F942" blockId="88.[189,794,1481,1729]" pageId="88" pageNumber="101">
<emphasis id="CCB21132FFE9FFB3FF3670EEDA73FA62" bold="true" box="[189,523,1481,1505]" pageId="88" pageNumber="101">Additional material examined</emphasis>
:
<materialsCitation id="4EAEC77DFFE9FFB3FD9170EEDA4AF9A2" ID-GBIF-Occurrence="3455118432" collectingDate="1997-03" collectingMethod="flight interception trap" collectionCode="BMNH" collectorName="S Lokan &amp; Chung" country="Malaysia" location="Sandakan" municipality="Borneo" pageId="88" pageNumber="101" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
<emphasis id="CCB21132FFE9FFB3FD9170EEDA03FA62" bold="true" box="[538,635,1481,1505]" pageId="88" pageNumber="101">
<collectingCountry id="86D18DB0FFE9FFB3FD9170EEDA03FA62" box="[538,635,1481,1505]" name="Malaysia" pageId="88" pageNumber="101">Malaysia</collectingCountry>
</emphasis>
:
<specimenCount id="E8C006A9FFE9FFB3FD0170EEDAD7FA62" box="[650,687,1481,1505]" pageId="88" pageNumber="101" type="male">1 ♂</specimenCount>
[teneral],
<collectingMunicipality id="1E1D575AFFE9FFB3FF3670CDD975F982" box="[189,269,1514,1537]" pageId="88" pageNumber="101">Borneo</collectingMunicipality>
,
<collectingRegion id="3C0203C2FFE9FFB3FE9670CED927F982" box="[285,351,1513,1537]" country="Malaysia" name="Sabah" pageId="88" pageNumber="101">Sabah</collectingRegion>
,
<location id="FB199BFBFFE9FFB3FEE570CED9A0F982" LSID="urn:lsid:plazi:treatment:766F7C36FFE9FFB2FF6871B4DA78FD02:FB199BFBFFE9FFB3FEE570CED9A0F982" box="[366,472,1513,1537]" country="Malaysia" municipality="Borneo" name="Sandakan" pageId="88" pageNumber="101" stateProvince="Sabah">Sandakan</location>
,
<collectorName id="5333A8F6FFE9FFB3FE6C70CEDA3DF982" box="[487,581,1513,1537]" pageId="88" pageNumber="101">S Lokan</collectorName>
,
<collectingMethod id="2787B537FFE9FFB3FDDF70CED892F9A2" pageId="88" pageNumber="101">flight interception trap</collectingMethod>
,
<date id="8A78EBE0FFE9FFB3FF7F732ED932F9A2" box="[244,330,1545,1569]" pageId="88" pageNumber="101" value="1997-03">
<collectingDate id="9A3C1208FFE9FFB3FF7F732ED932F9A2" box="[244,330,1545,1569]" pageId="88" pageNumber="101" value="1997-03">III.1997</collectingDate>
</date>
, leg.
<collectorName id="5333A8F6FFE9FFB3FE0B732ED9B1F9A2" box="[384,457,1545,1569]" pageId="88" pageNumber="101">Chung</collectorName>
(
<collectionCode id="98D755E5FFE9FFB3FE5D732EDA53F9A2" box="[470,555,1545,1569]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="88" pageNumber="101">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4EAEC77DFFE9FFB3FDB6732EDAB4F9C2" ID-GBIF-Occurrence="3455118322" collectingDate="1996-10" collectionCode="BMNH" collectorName="Chung" country="Malaysia" location="Sepilok" municipality="Sandakan" pageId="88" pageNumber="101" specimenCount="9" stateProvince="Sabah">
<specimenCount id="E8C006A9FFE9FFB3FDB6732EDA02F9A2" box="[573,634,1545,1569]" pageId="88" pageNumber="101" type="generic">9 exs.</specimenCount>
,
<collectingRegion id="3C0203C2FFE9FFB3FD0F732EDABEF9A2" box="[644,710,1545,1569]" country="Malaysia" name="Sabah" pageId="88" pageNumber="101">Sabah</collectingRegion>
,
<collectingMunicipality id="1E1D575AFFE9FFB3FD5B732ED89EF9C2" pageId="88" pageNumber="101">Sandakan</collectingMunicipality>
,
<location id="FB199BFBFFE9FFB3FF78730ED93CF9C2" LSID="urn:lsid:plazi:treatment:766F7C36FFE9FFB2FF6871B4DA78FD02:FB199BFBFFE9FFB3FF78730ED93CF9C2" box="[243,324,1577,1601]" country="Malaysia" municipality="Sandakan" name="Sepilok" pageId="88" pageNumber="101" stateProvince="Sabah">Sepilok</location>
,
<date id="8A78EBE0FFE9FFB3FEDB730ED9E6F9C2" box="[336,414,1577,1601]" pageId="88" pageNumber="101" value="1996-10">
<collectingDate id="9A3C1208FFE9FFB3FEDB730ED9E6F9C2" box="[336,414,1577,1601]" pageId="88" pageNumber="101" value="1996-10">X.1996</collectingDate>
</date>
, leg.
<collectorName id="5333A8F6FFE9FFB3FE5C730EDA58F9C2" box="[471,544,1577,1601]" pageId="88" pageNumber="101">Chung</collectorName>
(
<collectionCode id="98D755E5FFE9FFB3FDBB730EDAFCF9C2" box="[560,644,1577,1601]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="88" pageNumber="101">BMNH</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="4EAEC77DFFE9FFB3FD52730EDA3CF9E2" ID-GBIF-Occurrence="3455118328" collectingDate="1997-03" collectionCode="BMNH" collectorName="Chung" country="Malaysia" location="Sepilok" municipality="Sandakan" pageId="88" pageNumber="101" specimenCount="2" stateProvince="Sabah">
<specimenCount id="E8C006A9FFE9FFB3FD52730EDB6EF9C2" box="[729,790,1577,1601]" pageId="88" pageNumber="101" type="generic">2 exs.</specimenCount>
, same data, but
<date id="8A78EBE0FFE9FFB3FEF1736ED9B7F9E2" box="[378,463,1609,1633]" pageId="88" pageNumber="101" value="1997-03">
<collectingDate id="9A3C1208FFE9FFB3FEF1736ED9B7F9E2" box="[378,463,1609,1633]" pageId="88" pageNumber="101" value="1997-03">III.1997</collectingDate>
</date>
(
<collectionCode id="98D755E5FFE9FFB3FE6D736EDA44F9E2" box="[486,572,1609,1633]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="88" pageNumber="101">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4EAEC77DFFE9FFB3FDDD736EDAB4F942" ID-GBIF-Occurrence="3455118444" collectingDate="2001-08" collectingMethod="flight interception trap" collectionCode="BMNH" collectorName="Brendell &amp; Mendel" country="Indonesia" county="Borneo" latitude="-0.05" location="Rekut" longLatPrecision="1308" longitude="113.98333" municipality="Busang" pageId="88" pageNumber="101" specimenCount="9" stateProvince="Kalimantan Tengah">
<emphasis id="CCB21132FFE9FFB3FDDD736EDABBF9E2" bold="true" box="[598,707,1609,1633]" pageId="88" pageNumber="101">
<collectingCountry id="86D18DB0FFE9FFB3FDDD736EDABBF9E2" box="[598,707,1609,1633]" name="Indonesia" pageId="88" pageNumber="101">Indonesia</collectingCountry>
</emphasis>
:
<specimenCount id="E8C006A9FFE9FFB3FD52736EDB6EF9E2" box="[729,790,1609,1633]" pageId="88" pageNumber="101" type="generic">9 exs.</specimenCount>
,
<collectingCounty id="1718B5ACFFE9FFB3FF36734DD975F902" box="[189,269,1642,1665]" pageId="88" pageNumber="101">Borneo</collectingCounty>
,
<collectingRegion id="3C0203C2FFE9FFB3FEAF734EDA7DF902" box="[292,517,1641,1665]" country="Indonesia" name="Kalimantan Tengah" pageId="88" pageNumber="101">Kalimantan Tengah</collectingRegion>
, confluence of
<collectingMunicipality id="1E1D575AFFE9FFB3FD47734DDB62F901" box="[716,794,1642,1666]" pageId="88" pageNumber="101">Busang</collectingMunicipality>
and
<location id="FB199BFBFFE9FFB3FF7F73AED94DF922" LSID="urn:lsid:plazi:treatment:766F7C36FFE9FFB2FF6871B4DA78FD02:FB199BFBFFE9FFB3FF7F73AED94DF922" box="[244,309,1673,1697]" country="Indonesia" county="Borneo" latitude="-0.05" longLatPrecision="1308" longitude="113.98333" municipality="Busang" name="Rekut" pageId="88" pageNumber="101" stateProvince="Kalimantan Tengah">Rekut</location>
,
<geoCoordinate id="9BF2ABE7FFE9FFB3FEC373AED9F3F922" box="[328,395,1673,1697]" degrees="0" direction="south" minutes="03" orientation="latitude" pageId="88" pageNumber="101" precision="925" value="-0.05">0°03'S</geoCoordinate>
,
<geoCoordinate id="9BF2ABE7FFE9FFB3FE1573AED985F922" box="[414,509,1673,1697]" degrees="113" direction="east" minutes="59" orientation="longitude" pageId="88" pageNumber="101" precision="925" value="113.98333">113°59'E</geoCoordinate>
,
<collectingMethod id="2787B537FFE9FFB3FD9B73AEDB6EF922" box="[528,790,1673,1697]" pageId="88" pageNumber="101">flight interception trap</collectingMethod>
,
<date id="8A78EBE0FFE9FFB3FF36738ED95EF942" box="[189,294,1705,1729]" pageId="88" pageNumber="101" value="2001-08">
<collectingDate id="9A3C1208FFE9FFB3FF36738ED95EF942" box="[189,294,1705,1729]" pageId="88" pageNumber="101" value="2001-08">VIII.2001</collectingDate>
</date>
, leg.
<collectorName id="5333A8F6FFE9FFB3FED0738ED9CDF942" box="[347,437,1705,1729]" pageId="88" pageNumber="101">Brendell</collectorName>
&amp;
<collectorName id="5333A8F6FFE9FFB3FE5F738EDA5DF942" box="[468,549,1705,1729]" pageId="88" pageNumber="101">Mendel</collectorName>
(
<collectionCode id="98D755E5FFE9FFB3FDB9738EDAFEF942" box="[562,646,1705,1729]" pageId="88" pageNumber="101">BMNH</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE9FFB3FF3673CEDD15FEA2" pageId="88" pageNumber="101" type="discussion">
<paragraph id="FE79CD20FFE9FFB3FF3673CEDD15FEA2" blockId="88.[189,795,1769,2049]" lastBlockId="88.[831,1437,233,290]" pageId="88" pageNumber="101">
<emphasis id="CCB21132FFE9FFB3FF3673CED954F882" bold="true" box="[189,300,1769,1793]" pageId="88" pageNumber="101">Comment</emphasis>
: The original description is based on a unique type specimen from “Mt. Poi [= Mount Pueh;
<geoCoordinate id="9BF2ABE7FFE9FFB3FF36720ED97EF8C2" box="[189,262,1833,1857]" degrees="1" direction="north" minutes="48" orientation="latitude" pageId="88" pageNumber="101" precision="925" value="1.8">1°48'N</geoCoordinate>
,
<geoCoordinate id="9BF2ABE7FFE9FFB3FE9D720ED90FF8C2" box="[278,375,1833,1857]" degrees="109" direction="east" minutes="41" orientation="longitude" pageId="88" pageNumber="101" precision="925" value="109.683334">109°41'E</geoCoordinate>
], altitude
<quantity id="393E60C5FFE9FFB3FE62720EDA3AF8C2" box="[489,578,1833,1857]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="88" pageNumber="101" unit="ft" value="5000.0">5,000 ft.</quantity>
” (
<bibRefCitation id="9A57B0D1FFE9FFB3FDD4720EDB76F8C2" author="CAMERON, M." box="[607,782,1833,1857]" pageId="88" pageNumber="101" pagination="139 - 143" refId="ref77169" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON 1943</bibRefCitation>
). A dissection of the
<typeStatus id="217D7382FFE9FFB3FE1A726ED997F8E2" box="[401,495,1865,1889]" pageId="88" pageNumber="101" type="holotype">holotype</typeStatus>
revealed that it is a female. Remarkably, when distinguishing
<taxonomicName id="39C6B6A3FFE9FFB3FDA6724EDB61F802" authorityName="PACE" authorityYear="2014" box="[557,793,1897,1921]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CCB21132FFE9FFB3FDA6724EDB61F802" box="[557,793,1897,1921]" italics="true" pageId="88" pageNumber="101">Tetrabothrus femoralis</emphasis>
</taxonomicName>
from
<taxonomicName id="39C6B6A3FFE9FFB3FF7272ADDA77F822" authority=", PACE (2014)" authorityName="PACE" authorityYear="2014" box="[249,527,1929,1953]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="CCB21132FFE9FFB3FF7272ADD9F9F822" box="[249,385,1929,1953]" italics="true" pageId="88" pageNumber="101">T. borneensis</emphasis>
,
<bibRefCitation id="9A57B0D1FFE9FFB3FE0672AEDA77F822" author="PACE, R." box="[397,527,1929,1953]" pageId="88" pageNumber="101" pagination="727 - 794" refId="ref78194" refString="PACE, R. 2014: Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae). - Linzer Biologische Beitrage 46 (1): 727 - 794." type="journal article" year="2014">PACE (2014)</bibRefCitation>
</taxonomicName>
states that the “aedeagus of this new species is more similar to that of
<taxonomicName id="39C6B6A3FFE9FFB3FD3E728DD910F862" authority="CAMERON, 1939" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="CCB21132FFE9FFB3FD3E728DDB62F842" box="[693,794,1961,1985]" italics="true" pageId="88" pageNumber="101">T. indicus</emphasis>
CAMERON, 1939
</taxonomicName>
from Bengal, than to that of
<taxonomicName id="39C6B6A3FFE9FFB3FD1A72EDD909F782" authority="CAMERON, 1943" authorityName="CAMERON" authorityYear="1943" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="CCB21132FFE9FFB3FD1A72EDDB61F862" box="[657,793,1993,2017]" italics="true" pageId="88" pageNumber="101">T. borneensis</emphasis>
<bibRefCitation id="9A57B0D1FFE9FFB3FF3672CED909F782" author="CAMERON, M." box="[189,369,2025,2049]" pageId="88" pageNumber="101" pagination="139 - 143" refId="ref77169" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON, 1943</bibRefCitation>
</taxonomicName>
from Borneo, of which I have seen specimens of the type series”, although there appears to be only a single (undissected) female type specimen.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE9FFB2FCB4746ED951FEE2" lastPageId="89" lastPageNumber="102" pageId="88" pageNumber="101" type="description">
<paragraph id="FE79CD20FFE9FFB3FCB4746EDC6DFDA2" blockId="88.[830,1438,329,1505]" pageId="88" pageNumber="101">
<emphasis id="CCB21132FFE9FFB3FCB4746EDBA0FEE2" bold="true" box="[831,984,329,353]" pageId="88" pageNumber="101">Redescription</emphasis>
: Species of variable size; body length
<quantity id="393E60C5FFE9FFB3FCB4744EDBB9FE02" box="[831,961,361,385]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.8" metricValueMin="4.3" pageId="88" pageNumber="101" unit="mm" value="5.05" valueMax="5.8" valueMin="4.3">4.3–5.8 mm</quantity>
; length of forebody
<quantity id="393E60C5FFE9FFB3FB10744EDD65FE02" box="[1179,1309,361,385]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="88" pageNumber="101" unit="mm" value="2.1" valueMax="2.4" valueMin="1.8">1.8–2.4 mm</quantity>
. Coloration variable: body reddish to blackish-brown; elytra usually at least indistinctly bicoloured with the anterior portion paler than the posterior portion; legs pale-reddish to brown; antennae blackish-brown with reddish antennomeres I–III.
</paragraph>
<paragraph id="FE79CD20FFE9FFB3FCB4770EDC1FFD62" blockId="88.[830,1438,329,1505]" pageId="88" pageNumber="101">
Head (
<figureCitation id="66FDD1A5FFE9FFB3FC02770EDB98FDC2" box="[905,992,553,577]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="88" pageNumber="101">Fig. 160</figureCitation>
) strongly transverse, distinctly narrowed behind eyes, extensively impunctate in the middle; punctation in lateral dorsal portions very fine and sparse. Eyes large and bulging, approximately three times as long as postocular region in dorsal view, or nearly so. Antenna (
<figureCitation id="66FDD1A5FFE9FFB3FCCC77EEDBE7FD62" box="[839,927,713,737]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="88" pageNumber="101">Fig. 161</figureCitation>
) 1.0–
<quantity id="393E60C5FFE9FFB3FC5077EEDC53FD62" box="[987,1067,713,737]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="88" pageNumber="101" unit="mm" value="1.3">1.3 mm</quantity>
long.
</paragraph>
<paragraph id="FE79CD20FFE9FFB3FCB477CDDC1BFCE2" blockId="88.[830,1438,329,1505]" pageId="88" pageNumber="101">
Pronotum (
<figureCitation id="66FDD1A5FFE9FFB3FC3677CEDC6DFC82" box="[957,1045,745,769]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="88" pageNumber="101">Fig. 160</figureCitation>
) of somewhat variable shape, 1.12– 1.17 times as broad as long and 1.05–1.17 times as broad as head, strongly convex in cross-section; punctation very fine and of variable density.
</paragraph>
<paragraph id="FE79CD20FFE9FFB3FCB4764EDCC4FB82" blockId="88.[830,1438,329,1505]" pageId="88" pageNumber="101">
Elytra (
<figureCitation id="66FDD1A5FFE9FFB3FC13764EDB97FC02" box="[920,1007,873,897]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="88" pageNumber="101">Fig. 160</figureCitation>
) approximately 0.9 times as long as pronotum; punctation very fine and rather dense; pubescence long and suberect. Hind wings fully developed. Metatarsomere approximately as long as the combined length of metatarsomeres II and III.
</paragraph>
<paragraph id="FE79CD20FFE9FFB3FCB4712EDC65FAA2" blockId="88.[830,1438,329,1505]" pageId="88" pageNumber="101">
Abdomen (
<figureCitation id="66FDD1A5FFE9FFB3FC48712EDC62FBA2" box="[963,1050,1033,1057]" captionStart="Figs 160–180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160–180. Tetrabothrus borneensis, holotype (160–162), T. punctiventris (163–166), T. sulawesicus (167–169), Zyras illecebrosus, paratype (170–172), Z. gibbus, holotype (173–177), and Z. quasar, holotype (178–180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII– VIII (177). Scale bars: 160–165, 167–175, 178–180: 1.0 mm; 166, 176–177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="88" pageNumber="101">Fig. 162</figureCitation>
) narrower than elytra and with moderately deep anterior impressions on tergites III–VI; tergite III with two, tergites IV–VI with four setiferous punctures at posterior margin; tergite VII without nonsetiferous punctation and with pronounced palisade fringe at posterior margin; tergite VIII with numerous setae at and near posterior margin; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII truncate.
</paragraph>
<paragraph id="FE79CD20FFE9FFB3FCB4700DDC98FA62" blockId="88.[830,1438,329,1505]" pageId="88" pageNumber="101">
<specimenCount id="E8C006A9FFE9FFB3FCB4700DDB2CFAC2" box="[831,852,1322,1345]" pageId="88" pageNumber="101" type="male">♂</specimenCount>
: posterior margin of sternite VIII nearly truncate in the middle (
<figureCitation id="66FDD1A5FFE9FFB3FC75706EDC2FFAE2" box="[1022,1111,1353,1377]" captionStart="Figs 442–458" captionStartId="31.[151,191,1971,1993]" captionTargetBox="[155,1398,240,1950]" captionTargetId="figure-119@31.[151,1398,232,1971]" captionText="Figs 442–458. Tetrabothrus collucatus (442–443), T. borneensis (444–449), T. punctiventris (450–451), T. sulawesicus (452–454), Zyras illecebrosus, paratype (455–456), and Z. quasar, holotype (457–458): median lobe of aedeagus in lateral and in ventral view (442–445, 450–453, 455–458); apico-ventral portion of median lobe in lateral view (446–448); male sternite VIII (449, 454). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421226" httpUri="https://zenodo.org/record/6421226/files/figure.png" pageId="88" pageNumber="101">Fig. 449</figureCitation>
); median lobe of aedeagus (
<figureCitation id="66FDD1A5FFE9FFB3FCCC704EDBAFFA02" box="[839,983,1385,1409]" captionStart="Figs 442–458" captionStartId="31.[151,191,1971,1993]" captionTargetBox="[155,1398,240,1950]" captionTargetId="figure-119@31.[151,1398,232,1971]" captionText="Figs 442–458. Tetrabothrus collucatus (442–443), T. borneensis (444–449), T. punctiventris (450–451), T. sulawesicus (452–454), Zyras illecebrosus, paratype (455–456), and Z. quasar, holotype (457–458): median lobe of aedeagus in lateral and in ventral view (442–445, 450–453, 455–458); apico-ventral portion of median lobe in lateral view (446–448); male sternite VIII (449, 454). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421226" httpUri="https://zenodo.org/record/6421226/files/figure.png" pageId="88" pageNumber="101">Figs 444–448</figureCitation>
) of variable size,
<quantity id="393E60C5FFE9FFB3FB36704EDD2DFA02" box="[1213,1365,1385,1409]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.5" metricValueMin="4.3" pageId="88" pageNumber="101" unit="mm" value="0.49" valueMax="0.55" valueMin="0.43">0.43–0.55 mm</quantity>
long; ventral process rather short, its shape not very distinctive and somewhat variable; paramere longer than median lobe, its apical lobe of moderate length.
</paragraph>
<paragraph id="FE79CD20FFE9FFB3FCB4732EDD63F822" blockId="88.[831,1438,1545,1954]" pageId="88" pageNumber="101">
<emphasis id="CCB21132FFE9FFB3FCB4732EDC39F9A2" bold="true" box="[831,1089,1545,1569]" pageId="88" pageNumber="101">Intraspecific variation</emphasis>
: This species is subject to pronounced intraspecific variation of size, coloration, shape of pronotum, size of the median lobe of the aedeagus, and the shape of the ventral process of the aedeagus. In the material from
<collectingRegion id="3C0203C2FFE9FFB3FB9773AEDC26F922" box="[1052,1118,1673,1697]" country="Malaysia" name="Sabah" pageId="88" pageNumber="101">Sabah</collectingRegion>
, the size of the median lobe of the aedeagus ranges from
<quantity id="393E60C5FFE9FFB3FBD1738EDD72F942" box="[1114,1290,1705,1729]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.5" metricValueMin="4.8" pageId="88" pageNumber="101" unit="mm" value="0.515" valueMax="0.55" valueMin="0.48">0.48 to 0.55 mm</quantity>
and the apex of the ventral process tends to be longer and more slender in lateral view, whereas in the males from
<collectingRegion id="3C0203C2FFE9FFB3FA9473CEDBF5F8A2" country="Indonesia" name="Kalimantan Tengah" pageId="88" pageNumber="101">Kalimantan Tengah</collectingRegion>
the length of the median lobe ranges from
<quantity id="393E60C5FFE9FFB3FAE4722EDBC3F8C2" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.699999999999999" metricValueMin="4.3" pageId="88" pageNumber="101" unit="mm" value="0.44999999999999996" valueMax="0.47" valueMin="0.43">0.43 to 0.47 mm</quantity>
and the apex of the ventral process tends to be shorter. The
<typeStatus id="217D7382FFE9FFB3FC62726EDC3FF8E2" box="[1001,1095,1865,1889]" pageId="88" pageNumber="101" type="holotype">holotype</typeStatus>
differs from the examined nontype material by paler coloration, sparse punctation of the pronotum, and slightly greater body size.
</paragraph>
<paragraph id="FE79CD20FFE9FFB2FCB472EED951FEE2" blockId="88.[831,1436,1993,2081]" lastBlockId="89.[151,756,233,353]" lastPageId="89" lastPageNumber="102" pageId="88" pageNumber="101">
<emphasis id="CCB21132FFE9FFB3FCB472EEDC6AF862" bold="true" box="[831,1042,1993,2017]" pageId="88" pageNumber="101">Comparative notes</emphasis>
:
<taxonomicName id="39C6B6A3FFE9FFB3FBAA72EEDD66F862" authorityName="CAMERON" authorityYear="1943" box="[1057,1310,1993,2017]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="101" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="CCB21132FFE9FFB3FBAA72EEDD66F862" box="[1057,1310,1993,2017]" italics="true" pageId="88" pageNumber="101">Tetrabothrus borneensis</emphasis>
</taxonomicName>
is not very distinctive regarding its external characters and characterized particularly by the shape of median lobe of the aedeagus. The aedeagus somewhat resembles that of
<taxonomicName id="39C6B6A3FFE8FFB2FF1C742DD889FEA2" authorityName="Assing" authorityYear="2009" box="[151,241,266,289]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="102" phylum="Arthropoda" rank="species" species="puetzi">
<emphasis id="CCB21132FFE8FFB2FF1C742DD889FEA2" box="[151,241,266,289]" italics="true" pageId="89" pageNumber="102">T. puetzi</emphasis>
</taxonomicName>
(
<collectingCountry id="86D18DB0FFE8FFB2FE8E742ED932FEA2" box="[261,330,265,289]" name="China" pageId="89" pageNumber="102">China</collectingCountry>
,
<collectingRegion id="3C0203C2FFE8FFB2FED1742ED9CAFEA2" box="[346,434,265,289]" country="China" name="Taiwan)" pageId="89" pageNumber="102">Taiwan)</collectingRegion>
, from which
<taxonomicName id="39C6B6A3FFE8FFB2FDDB742DDAA0FEA2" authorityName="CAMERON" authorityYear="1943" box="[592,728,265,289]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="102" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis id="CCB21132FFE8FFB2FDDB742DDAA0FEA2" box="[592,728,265,289]" italics="true" pageId="89" pageNumber="102">T. borneensis</emphasis>
</taxonomicName>
is readily distinguished by much paler legs with undilated femora alone.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE8FFB2FF1C74AEDA78FD02" pageId="89" pageNumber="102" type="distribution">
<paragraph id="FE79CD20FFE8FFB2FF1C74AEDA78FD02" blockId="89.[150,758,393,641]" pageId="89" pageNumber="102">
<emphasis id="CCB21132FFE8FFB2FF1C74AED98DFE22" bold="true" box="[151,501,393,417]" pageId="89" pageNumber="102">Distribution and natural history</emphasis>
: The examined material was collected in four localities in Borneo, one in
<collectingCountry id="86D18DB0FFE8FFB2FD11748EDA8BFE42" box="[666,755,425,449]" name="Malaysia" pageId="89" pageNumber="102">Sarawak</collectingCountry>
(
<typeStatus id="217D7382FFE8FFB2FF1474EED8B7FE62" box="[159,207,457,481]" pageId="89" pageNumber="102">type</typeStatus>
locality), two in
<collectingRegion id="3C0203C2FFE8FFB2FE0874EED9BBFE62" box="[387,451,457,481]" country="Malaysia" name="Sabah" pageId="89" pageNumber="102">Sabah</collectingRegion>
(
<collectingCountry id="86D18DB0FFE8FFB2FE5A74EEDA4DFE62" box="[465,565,457,481]" name="Malaysia" pageId="89" pageNumber="102">Malaysia</collectingCountry>
), and one in
<collectingRegion id="3C0203C2FFE8FFB2FD3474EED947FD82" country="Indonesia" name="Kalimantan Tengah" pageId="89" pageNumber="102">Kalimantan Tengah</collectingRegion>
(
<collectingCountry id="86D18DB0FFE8FFB2FEC574CED9C5FD82" box="[334,445,489,513]" name="Indonesia" pageId="89" pageNumber="102">Indonesia</collectingCountry>
). The species was doubtfully recorded from the Danum Valley (Sabah) based on
<specimenCount id="E8C006A9FFE8FFB2FD34772ED89FFDC2" pageId="89" pageNumber="102" type="female">three females</specimenCount>
by
<bibRefCitation id="9A57B0D1FFE8FFB2FE80770ED9E8FDC2" author="PACE, R." box="[267,400,553,577]" pageId="89" pageNumber="102" pagination="727 - 794" refId="ref78194" refString="PACE, R. 2014: Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae). - Linzer Biologische Beitrage 46 (1): 727 - 794." type="journal article" year="2014">PACE (2014)</bibRefCitation>
. The examined non-type material was collected with flight interception traps. One specimen collected in March is teneral.
</paragraph>
</subSubSection>
</treatment>
</document>
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