270 lines
36 KiB
XML
270 lines
36 KiB
XML
<document id="A2EBB6C23DC9BEE45219C2411AFC18F4" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FF9CFFC5FF367054DA7AFE42" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Orphnebius reductus Assing 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="59" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="58" updateTime="1699235685730" updateUser="ExternalLinkService">
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<mods:titleInfo id="FF8067E9D1971E572515B882234F0954">
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<mods:title id="1774978196F314270F1CFAF6850199E7">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="6C8486A67DD3E5D17BF813B7E0592964" type="personal">
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<mods:roleTerm id="EE9E89C6906B7F1942F57FC0EBC7A435">Author</mods:roleTerm>
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<mods:namePart id="EDED52E0DBB08E8B55098A238D99BDB4">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="AC469E53BE26D2ACFEEF0C5C15625C11">text</mods:typeOfResource>
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<mods:title id="1D0C1833D21BF9DE01586B60270DD448">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="B14B802F9EB3BD5CCD73A99D236F0A9B">
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<mods:date id="AF3D5B4FE9C82509EBE78D28B1A12890">2016</mods:date>
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<mods:number id="A63E38B4957D3DED11BC60FAD92C53D7">2016-12-20</mods:number>
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<mods:detail id="11675844942F40D44687928D025A2857" type="volume">
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<mods:number id="19EB7205C4547F896046843CDC80E211">66</mods:number>
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<mods:start id="C75006B296C16476668A68A8A3360557">13</mods:start>
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<mods:url id="05EBDA7CAF8C97B051FB78EAD45D557F">https://www.contributions-to-entomology.org/article/view/1904</mods:url>
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<mods:classification id="F73178A03E30EF39BE02FD2FDF3B9AEB">journal article</mods:classification>
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<mods:identifier id="309C81E601D42480DD769543FAE0097B" type="CLB-Dataset">2419</mods:identifier>
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<mods:identifier id="494D710B274041D12BA872CD0841DAD6" type="DOI">10.21248/contrib.entomol.66.1.13-111</mods:identifier>
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<mods:identifier id="7637CD01D48DE57B26D74D8A29AB7AF7" type="GBIF-Dataset">d2933fad-1cac-4a5a-a558-2633f8314541</mods:identifier>
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<treatment id="766F7C36FF9CFFC5FF367054DA7AFE42" ID-DOI="http://doi.org/10.5281/zenodo.5903426" ID-GBIF-Taxon="192602829" ID-Zenodo-Dep="5903426" LSID="urn:lsid:plazi:treatment:766F7C36FF9CFFC5FF367054DA7AFE42" httpUri="http://treatment.plazi.org/id/766F7C36FF9CFFC5FF367054DA7AFE42" lastPageId="46" lastPageNumber="59" pageId="45" pageNumber="58">
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<subSubSection id="B6DC9EABFF9CFFC6FF367054DA7BFA0D" box="[189,515,1395,1422]" pageId="45" pageNumber="58" type="nomenclature">
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<paragraph id="FE79CD20FF9CFFC6FF367054DA7BFA0D" blockId="45.[189,515,1395,1455]" box="[189,515,1395,1422]" pageId="45" pageNumber="58">
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<heading id="A5317A4CFF9CFFC6FF367054DA7BFA0D" box="[189,515,1395,1422]" centered="true" fontSize="11" level="2" pageId="45" pageNumber="58" reason="3">
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<taxonomicName id="39C6B6A3FF9CFFC6FF367054D9EBFA0D" ID-CoL="8Y7CK" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[189,403,1395,1422]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="58" phylum="Arthropoda" rank="species" species="reductus" status="sp. nov.">
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<emphasis id="CCB21132FF9CFFC6FF367054D9EBFA0D" box="[189,403,1395,1422]" italics="true" pageId="45" pageNumber="58">Orphnebius reductus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D781AC49FF9CFFC6FE117054DA7BFA0D" box="[410,515,1395,1422]" pageId="45" pageNumber="58" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B6DC9EABFF9CFFC6FF3670BED902FA2C" box="[189,378,1433,1455]" pageId="45" pageNumber="58" type="description">
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<paragraph id="FE79CD20FF9CFFC6FF3670BED902FA2C" blockId="45.[189,515,1395,1455]" box="[189,378,1433,1455]" pageId="45" pageNumber="58">
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(
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<figureCitation id="66FDD1A5FF9CFFC6FF4A70BED97AFA2C" box="[193,258,1433,1455]" captionStart="Figs 1–26" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 1–26. Antenna (1–21) and forebody (22–26) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 22–26: 1.0 mm; 1–21: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="45" pageNumber="58">Figs 18</figureCitation>
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,
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<figureCitation id="66FDD1A5FF9CFFC6FE8270BED959FA2C" box="[265,289,1433,1455]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="45" pageNumber="58">49</figureCitation>
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,
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<figureCitation id="66FDD1A5FF9CFFC6FEA270BED90CFA2C" box="[297,372,1433,1455]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="45" pageNumber="58">269–273</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B6DC9EABFF9CFFC6FF1C70EED96AF962" pageId="45" pageNumber="58" type="materials_examined">
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<paragraph id="FE79CD20FF9CFFC6FF1C70EED96AF962" blockId="45.[150,757,1481,1761]" pageId="45" pageNumber="58">
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<emphasis id="CCB21132FF9CFFC6FF1C70EED93AFA62" bold="true" box="[151,322,1481,1505]" pageId="45" pageNumber="58">Type material</emphasis>
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||
:
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<materialsCitation id="4EAEC77DFF9CFFC6FED570EEDA89F9C2" ID-GBIF-Occurrence="3455118423" collectingDate="1999-05-01" collectingDateMax="1999-05-16" collectingDateMin="1999-05-01" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos" determinerName="V. Assing" elevation="1200" location="Louangphrabang pr." longitude="102.23333" pageId="45" pageNumber="58" specimenCount="1" specimenCount-male="1" stateProvince="Louangphabang" typeStatus="holotype">
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||
<typeStatus id="217D7382FF9CFFC6FED570EED9BCFA62" box="[350,452,1481,1505]" pageId="45" pageNumber="58" type="holotype">Holotype</typeStatus>
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<specimenCount id="E8C006A9FF9CFFC6FE5170EDD997FA62" box="[474,495,1482,1505]" pageId="45" pageNumber="58" type="male">♂</specimenCount>
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: “
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||
<collectingCountry id="86D18DB0FF9CFFC6FD9E70EEDA33FA62" box="[533,587,1481,1505]" name="Laos" pageId="45" pageNumber="58">Laos</collectingCountry>
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,
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<date id="8A78EBE0FF9CFFC6FDED70EEDA97FA62" box="[614,751,1481,1505]" pageId="45" pageNumber="58" value="1999-05-01" valueMax="1999-05-16" valueMin="1999-05-01">
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<collectingDate id="9A3C1208FF9CFFC6FDED70EEDA97FA62" box="[614,751,1481,1505]" pageId="45" pageNumber="58" value="1999-05-01" valueMax="1999-05-16" valueMin="1999-05-01">1.–16.v.1999</collectingDate>
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</date>
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,
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<location id="FB199BFBFF9CFFC6FF1C70CED907F982" LSID="urn:lsid:plazi:treatment:766F7C36FF9CFFC5FF367054DA7AFE42:FB199BFBFF9CFFC6FF1C70CED907F982" box="[151,383,1513,1537]" country="Laos" longitude="102.23333" name="Louangphrabang pr." pageId="45" pageNumber="58" stateProvince="Louangphabang">Louangphrabang pr.</location>
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||
, 20°33–4'N 102°14'E,
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||
<location id="FB199BFBFF9CFFC6FD0E70CDD8BBF9A2" LSID="urn:lsid:plazi:treatment:766F7C36FF9CFFC5FF367054DA7AFE42:FB199BFBFF9CFFC6FD0E70CDD8BBF9A2" country="Laos" longitude="102.23333" name="Ban Song Cha" pageId="45" pageNumber="58" stateProvince="Louangphabang">Ban Song Cha</location>
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(
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<locationDeviation id="EFCBD402FF9CFFC6FF5D732ED949F9A2" box="[214,305,1545,1569]" pageId="45" pageNumber="58">
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W
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</locationDeviation>
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),
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<quantity id="393E60C5FF9CFFC6FEC3732ED9E3F9A2" box="[328,411,1545,1569]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="45" pageNumber="58" unit="m" value="1200.0">
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<elevation id="75EB2A13FF9CFFC6FEC3732ED9E3F9A2" box="[328,411,1545,1569]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="45" pageNumber="58" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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,
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||
<collectorName id="5333A8F6FF9CFFC6FE23732EDA62F9A2" box="[424,538,1545,1569]" pageId="45" pageNumber="58">Vít Kubáň</collectorName>
|
||
leg. / Holotypus ♂
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||
<emphasis id="CCB21132FF9CFFC6FF1C730ED90AF9C2" box="[151,370,1577,1601]" italics="true" pageId="45" pageNumber="58">Orphnebiusreductus</emphasis>
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||
<taxonomicNameLabel id="D781AC49FF9CFFC6FEFF730DD9D6F9C2" box="[372,430,1578,1601]" pageId="45" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
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||
, det.
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||
<determinerName id="0BA521E4FF9CFFC6FE6A730DDA3EF9C2" box="[481,582,1577,1601]" pageId="45" pageNumber="58">V. Assing</determinerName>
|
||
2015”(
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||
<collectionCode id="98D755E5FF9CFFC6FD1A730EDA90F9C2" box="[657,744,1577,1601]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="45" pageNumber="58">NHMB</collectionCode>
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||
)
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||
</materialsCitation>
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.
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||
<materialsCitation id="4EAEC77DFF9CFFC6FF1C736DD915F902" ID-GBIF-Occurrence="3455118464" collectingDate="1999-05-01" collectingDateMax="1999-05-16" collectingDateMin="1999-05-01" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos" determinerName="V. Assing" elevation="1200" location="Louangphrabang" longitude="102.23333" pageId="45" pageNumber="58" specimenCount="25" specimenCount-female="10" specimenCount-male="15" stateProvince="Louangphabang" typeStatus="paratype">
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||
<typeStatus id="217D7382FF9CFFC6FF1C736DD979F9E1" box="[151,257,1610,1634]" pageId="45" pageNumber="58" type="paratype">Paratypes</typeStatus>
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||
:
|
||
<specimenCount id="E8C006A9FF9CFFC6FE87736ED92BF9E2" box="[268,339,1609,1633]" pageId="45" pageNumber="58" type="male">15 ♂♂</specimenCount>
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,
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||
<specimenCount id="E8C006A9FF9CFFC6FED4736ED9DFF9E2" box="[351,423,1609,1633]" pageId="45" pageNumber="58" type="female">10 ♀♀</specimenCount>
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||
[3 teneral]: same data as holotype (
|
||
<collectionCode id="98D755E5FF9CFFC6FF58734ED95EF902" box="[211,294,1641,1665]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="45" pageNumber="58">NHMB</collectionCode>
|
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, cAss)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EAEC77DFF9CFFC6FEF3734ED885F922" ID-GBIF-Occurrence="3455118475" collectingDate="1999-04-24" collectingDateMax="1999-05-16" collectingDateMin="1999-04-24" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos" determinerName="V. Assing" elevation="1200" location="Louangphrabang" longitude="102.23333" pageId="45" pageNumber="58" specimenCount="1" specimenCount-female="1" stateProvince="Louangphabang" typeStatus="paratype">
|
||
<specimenCount id="E8C006A9FF9CFFC6FEF3734ED9E6F902" box="[376,414,1641,1665]" pageId="45" pageNumber="58" type="female">1 ♀</specimenCount>
|
||
: same data, but
|
||
<date id="8A78EBE0FF9CFFC6FDC7734EDA8CF902" box="[588,756,1641,1665]" pageId="45" pageNumber="58" value="1999-04-24" valueMax="1999-05-16" valueMin="1999-04-24">
|
||
<collectingDate id="9A3C1208FF9CFFC6FDC7734EDA8CF902" box="[588,756,1641,1665]" pageId="45" pageNumber="58" value="1999-04-24" valueMax="1999-05-16" valueMin="1999-04-24">24.iv.–16.v.1999</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="98D755E5FF9CFFC6FF1473AED88CF922" box="[159,244,1673,1697]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="45" pageNumber="58">NHMB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EAEC77DFF9CFFC6FE8D73AED976F962" ID-GBIF-Occurrence="3455118530" collectingDate="2003-05-28" collectingDateMax="2003-06-20" collectingDateMin="2003-05-28" collectorName="Vit Kuban" country="Laos" elevation="1500" location="Phongsaly prov." pageId="45" pageNumber="58" specimenCount="1" specimenCount-male="1" stateProvince="Phongsali" typeStatus="paratype">
|
||
<specimenCount id="E8C006A9FF9CFFC6FE8D73AED952F922" box="[262,298,1673,1697]" pageId="45" pageNumber="58" type="male">1♂</specimenCount>
|
||
: “
|
||
<collectingCountry id="86D18DB0FF9CFFC6FECB73AED912F922" box="[320,362,1673,1697]" pageId="45" pageNumber="58">Lao</collectingCountry>
|
||
,
|
||
<location id="FB199BFBFF9CFFC6FEF873AEDA67F922" LSID="urn:lsid:plazi:treatment:766F7C36FF9CFFC5FF367054DA7AFE42:FB199BFBFF9CFFC6FEF873AEDA67F922" box="[371,543,1673,1697]" country="Laos" name="Phongsaly prov." pageId="45" pageNumber="58" stateProvince="Phongsali">Phongsaly prov.</location>
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||
, 21°41–2'N 102°06– 8'E,
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||
<date id="8A78EBE0FF9CFFC6FF48738ED917F942" box="[195,367,1705,1729]" pageId="45" pageNumber="58" value="2003-05-28" valueMax="2003-06-20" valueMin="2003-05-28">
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||
<collectingDate id="9A3C1208FF9CFFC6FF48738ED917F942" box="[195,367,1705,1729]" pageId="45" pageNumber="58" value="2003-05-28" valueMax="2003-06-20" valueMin="2003-05-28">28.v.–20.vi.2003</collectingDate>
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||
</date>
|
||
,
|
||
<location id="FB199BFBFF9CFFC6FEF3738EDA6FF942" LSID="urn:lsid:plazi:treatment:766F7C36FF9CFFC5FF367054DA7AFE42:FB199BFBFF9CFFC6FEF3738EDA6FF942" box="[376,535,1705,1729]" country="Laos" name="Phongsaly env." pageId="45" pageNumber="58" stateProvince="Phongsali">Phongsaly env.</location>
|
||
, ~
|
||
<quantity id="393E60C5FF9CFFC6FDA6738EDAF8F942" box="[557,640,1705,1729]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="45" pageNumber="58" unit="m" value="1500.0">
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<elevation id="75EB2A13FF9CFFC6FDA6738EDAF8F942" box="[557,640,1705,1729]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="45" pageNumber="58" unit="m" value="1500.0">1500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="5333A8F6FF9CFFC6FD02738EDA8CF942" box="[649,756,1705,1729]" pageId="45" pageNumber="58">Vít Kubáň</collectorName>
|
||
leg.” (cAss)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9CFFC6FF1C722EDA92F8C2" pageId="45" pageNumber="58" type="etymology">
|
||
<paragraph id="FE79CD20FF9CFFC6FF1C722EDA92F8C2" blockId="45.[151,756,1801,1857]" pageId="45" pageNumber="58">
|
||
<emphasis id="CCB21132FF9CFFC6FF1C722ED976F8A2" bold="true" box="[151,270,1801,1825]" pageId="45" pageNumber="58">Etymology</emphasis>
|
||
: The specific epithet (Latin, adjective) alludes to the conspicuously reduced condylite of the paramere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9CFFC6FF1C724EDC18F7A2" pageId="45" pageNumber="58" type="description">
|
||
<paragraph id="FE79CD20FF9CFFC6FF1C724EDBEAFEA2" blockId="45.[150,756,1897,2049]" lastBlockId="45.[792,1399,233,1794]" pageId="45" pageNumber="58">
|
||
<emphasis id="CCB21132FF9CFFC6FF1C724ED961F802" bold="true" box="[151,281,1897,1921]" pageId="45" pageNumber="58">Description</emphasis>
|
||
: Body length
|
||
<quantity id="393E60C5FF9CFFC6FE22724EDA52F802" box="[425,554,1897,1921]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.6" metricValueMin="3.8" pageId="45" pageNumber="58" unit="mm" value="4.199999999999999" valueMax="4.6" valueMin="3.8">3.8–4.6 mm</quantity>
|
||
; length of forebody
|
||
<quantity id="393E60C5FF9CFFC6FF1D72AED960F822" box="[150,280,1929,1953]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="45" pageNumber="58" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 mm</quantity>
|
||
. Coloration: forebody blackish; abdomen pale-reddish; legs yellowish-red, with the profemora yellowish and the meso- and metafemora brown; antennae blackish, with the basal 1–3 antennomeres yellowish; maxillary palpi reddish-yellow with yellowish terminal palpomere.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9CFFC6FC91740EDB16FC82" blockId="45.[792,1399,233,1794]" pageId="45" pageNumber="58">
|
||
Head (
|
||
<figureCitation id="66FDD1A5FF9CFFC6FCE3740EDBCBFEC2" box="[872,947,297,321]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="45" pageNumber="58">Fig. 49</figureCitation>
|
||
) strongly transverse, 1.30–1.35 times as broad as long; punctation of lateral portions extremely fine, barely visible, and very sparse; median and posterior dorsal portions extensively impunctate; interstices without microsculpture. Eyes enormous, nearly reaching posterior margin, and strongly bulging. Antenna (
|
||
<figureCitation id="66FDD1A5FF9CFFC6FCAA74CEDB14FD82" box="[801,876,489,513]" captionStart="Figs 1–26" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 1–26. Antenna (1–21) and forebody (22–26) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 22–26: 1.0 mm; 1–21: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="45" pageNumber="58">Fig. 18</figureCitation>
|
||
) sexually dimorphic,
|
||
<quantity id="393E60C5FF9CFFC6FBEB74CEDCA7FD82" box="[1120,1247,489,513]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="45" pageNumber="58" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
|
||
long in male and
|
||
<quantity id="393E60C5FF9CFFC6FCDB772EDBB7FDA2" box="[848,975,521,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="45" pageNumber="58" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
|
||
long in female, distinctly incrassate, and distinctly asymmetric; antennomere IV moderately transverse; antennomeres V–X of gradually and distinctly increasing width, increasingly asymmetric, and increasingly transverse, more transverse in female than in male; X approximately twice as wide as long; XI elongate, longer (male) or shorter (female) than the combined length of VIII–X.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9CFFC6FC91762DDB2DFC62" blockId="45.[792,1399,233,1794]" pageId="45" pageNumber="58">
|
||
Pronotum (
|
||
<figureCitation id="66FDD1A5FF9CFFC6FC28762EDB95FCA2" box="[931,1005,777,801]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="45" pageNumber="58">Fig. 49</figureCitation>
|
||
) moderately transverse, approximately 1.25 times as broad as long and 1.05–1.10 times as broad as head, moderately strongly convex in crosssection; posterior angles obsolete; disc with a median pair of punctures, otherwise practically impunctate; lateral margins each with (probably) four long and erect black setae.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9CFFC6FC9176CEDCA8FB02" blockId="45.[792,1399,233,1794]" pageId="45" pageNumber="58">
|
||
Elytra (
|
||
<figureCitation id="66FDD1A5FF9CFFC6FCEC76CEDBC9FB82" box="[871,945,1001,1025]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="45" pageNumber="58">Fig. 49</figureCitation>
|
||
) approximately 0.7 times as long as pronotum; suture distinctly gaping posteriorly; punctation sparse and very fine; pubescence long and pale. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of II and III.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9CFFC6FC9171AEDB08FA62" blockId="45.[792,1399,233,1794]" pageId="45" pageNumber="58">
|
||
Abdomen: paratergites strongly elevated and sharply edged; tergites III–VI each with two lateral setiferous punctures on either side, tergite VI additionally with four setiferous punctures at posterior margin; tergite VII with non-setiferous punctation in posterior two-thirds and with eight setiferous punctures near posterior margin, posterior margin with palisade fringe; tergite VIII (
|
||
<figureCitation id="66FDD1A5FF9CFFC6FCAA704EDB01FA02" box="[801,889,1385,1409]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="45" pageNumber="58">Fig. 272</figureCitation>
|
||
) with two transverse rows of black setae posteriorly and with broadly truncate posterior margin; sternite VIII with broadly, very weakly convex posterior margin.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9CFFC6FC9170CDDCC2F942" blockId="45.[792,1399,233,1794]" pageId="45" pageNumber="58">
|
||
<specimenCount id="E8C006A9FF9CFFC6FC9170CDDB57F982" box="[794,815,1514,1537]" pageId="45" pageNumber="58" type="male">♂</specimenCount>
|
||
: hemi-tergites IX and tergite X with extremely dense and long pubescence; median lobe of aedeagus (
|
||
<figureCitation id="66FDD1A5FF9CFFC6FCAA730EDBCAF9C2" box="[801,946,1577,1601]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="45" pageNumber="58">Figs 269–270</figureCitation>
|
||
)
|
||
<quantity id="393E60C5FF9CFFC6FC4C730EDC18F9C2" box="[967,1120,1577,1601]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.199999999999999" metricValueMin="6.8" pageId="45" pageNumber="58" unit="mm" value="0.7" valueMax="0.72" valueMin="0.68">0.68–0.72 mm</quantity>
|
||
long and of distinctive shape; internal sac without strongly sclerotized structures; paramere (
|
||
<figureCitation id="66FDD1A5FF9CFFC6FC59734EDC52F902" box="[978,1066,1641,1665]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="45" pageNumber="58">Fig. 271</figureCitation>
|
||
) approximately
|
||
<quantity id="393E60C5FF9CFFC6FB50734EDD41F902" box="[1243,1337,1641,1665]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="45" pageNumber="58" unit="mm" value="0.55">0.55 mm</quantity>
|
||
long, with apically hooked paramerite of modified shape and with condylite of strongly reduced size.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9CFFC6FC9173EDDD49F882" blockId="45.[792,1399,233,1794]" pageId="45" pageNumber="58">
|
||
<specimenCount id="E8C006A9FF9CFFC6FC9173EDDB57F962" box="[794,815,1738,1761]" pageId="45" pageNumber="58" type="female">♀</specimenCount>
|
||
: abdominal segments IX–X of similar shape and chaetotaxy as in
|
||
<taxonomicName id="39C6B6A3FF9CFFC6FC1273CEDC7DF882" authorityName="Eppelsheim" authorityYear="1895" box="[921,1029,1769,1793]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="58" phylum="Arthropoda" rank="species" species="hauseri">
|
||
<emphasis id="CCB21132FF9CFFC6FC1273CEDC7DF882" box="[921,1029,1769,1793]" italics="true" pageId="45" pageNumber="58">O. hauseri</emphasis>
|
||
</taxonomicName>
|
||
; spermatheca as in
|
||
<figureCitation id="66FDD1A5FF9CFFC6FB5D73CEDD55F882" box="[1238,1325,1769,1793]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="45" pageNumber="58">Fig. 273</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9CFFC6FC91720EDC18F7A2" blockId="45.[793,1400,1833,2081]" pageId="45" pageNumber="58">
|
||
<emphasis id="CCB21132FF9CFFC6FC91720EDB95F8C2" bold="true" box="[794,1005,1833,1857]" pageId="45" pageNumber="58">Comparative notes</emphasis>
|
||
: Based on the modifications of the abdominal segments IX–X, the shape and chaetotaxy of tergite and sternite VIII, and on the morphology of the aedeagus,
|
||
<taxonomicName id="39C6B6A3FF9CFFC6FC0372AEDC78F822" authorityName="Assing" authorityYear="2016" box="[904,1024,1929,1953]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="58" phylum="Arthropoda" rank="species" species="reductus">
|
||
<emphasis id="CCB21132FF9CFFC6FC0372AEDC78F822" box="[904,1024,1929,1953]" italics="true" pageId="45" pageNumber="58">O. reductus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="39C6B6A3FF9CFFC6FB2372AEDD6CF822" authorityName="Eppelsheim" authorityYear="1895" box="[1192,1300,1929,1953]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="58" phylum="Arthropoda" rank="species" species="hauseri">
|
||
<emphasis id="CCB21132FF9CFFC6FB2372AEDD6CF822" box="[1192,1300,1929,1953]" italics="true" pageId="45" pageNumber="58">O. hauseri</emphasis>
|
||
</taxonomicName>
|
||
group. It is characterized especially by the enormous eyes, by the shape and pronounced sexual dimorphism of the antennae, the shape of the median lobe of the aedeagus, and the derived shape of the paramere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9FFFC5FF3675CEDA7AFE42" pageId="46" pageNumber="59" type="distribution">
|
||
<paragraph id="FE79CD20FF9FFFC5FF3675CEDA7AFE42" blockId="46.[187,794,233,449]" pageId="46" pageNumber="59">
|
||
<emphasis id="CCB21132FF9FFFC5FF3675CEDA60FE82" bold="true" box="[189,536,233,257]" pageId="46" pageNumber="59">Distribution and natural history</emphasis>
|
||
: This species is currently known from two localities in
|
||
<collectingRegion id="3C0203C2FF9FFFC5FD9C742EDAFDFEA2" box="[535,645,265,289]" country="Laos" name="Phongsali" pageId="46" pageNumber="59">Phongsaly</collectingRegion>
|
||
and Louang Phrabang provinces, North
|
||
<collectingCountry id="86D18DB0FF9FFFC5FE7F740DDA51FEC2" box="[500,553,298,321]" name="Laos" pageId="46" pageNumber="59">Laos</collectingCountry>
|
||
. The specimens were collected at altitudes of 1200 and approximately
|
||
<quantity id="393E60C5FF9FFFC5FD4E746EDB6EFEE2" box="[709,790,329,353]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="46" pageNumber="59" unit="m" value="1500.0">1500 m</quantity>
|
||
, together with
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FED1744ED9BFFE02" authorityName="Assing" authorityYear="2016" box="[346,455,361,385]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="lunatus">
|
||
<emphasis id="CCB21132FF9FFFC5FED1744ED9BFFE02" box="[346,455,361,385]" italics="true" pageId="46" pageNumber="59">O. lunatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FE5C744EDA15FE02" authorityName="Assing" authorityYear="2016" box="[471,621,361,385]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrapicalis">
|
||
<emphasis id="CCB21132FF9FFFC5FE5C744EDA15FE02" box="[471,621,361,385]" italics="true" pageId="46" pageNumber="59">O. nigrapicalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FDF5744EDB6CFE02" authorityName="Assing" authorityYear="2016" box="[638,788,361,385]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="grandicollis">
|
||
<emphasis id="CCB21132FF9FFFC5FDF5744EDB6CFE02" box="[638,788,361,385]" italics="true" pageId="46" pageNumber="59">O. grandicollis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FF3674AED94CFE22" authorityName="Assing" authorityYear="2016" box="[189,308,393,417]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="extensus">
|
||
<emphasis id="CCB21132FF9FFFC5FF3674AED94CFE22" box="[189,308,393,417]" italics="true" pageId="46" pageNumber="59">O. extensus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FEC974AED9CCFE22" authorityName="Assing" authorityYear="2016" box="[322,436,393,417]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="serratus">
|
||
<emphasis id="CCB21132FF9FFFC5FEC974AED9CCFE22" box="[322,436,393,417]" italics="true" pageId="46" pageNumber="59">O. serratus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FE4874AEDA4BFE22" authorityName="Assing" authorityYear="2016" box="[451,563,393,417]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="bicuspis">
|
||
<emphasis id="CCB21132FF9FFFC5FE4874AEDA4BFE22" box="[451,563,393,417]" italics="true" pageId="46" pageNumber="59">O. bicuspis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FDC974AEDAEDFE22" authorityName="Assing" authorityYear="2016" box="[578,661,393,417]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="vates">
|
||
<emphasis id="CCB21132FF9FFFC5FDC974AEDAEDFE22" box="[578,661,393,417]" italics="true" pageId="46" pageNumber="59">O. vates</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FD2874AED8A4FE42" authorityName="Assing" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="baccillatus">
|
||
<emphasis id="CCB21132FF9FFFC5FD2874AED8A4FE42" italics="true" pageId="46" pageNumber="59">O. baccillatus</emphasis>
|
||
</taxonomicName>
|
||
, and two unnamed species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |