treatments-xml/data/76/6F/7C/766F7C36FF9BFFC0FCEE7634DCCEFCE2.xml

<document id="3E364F019A980BA351FB7D3C3954713A" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FF9BFFC0FCEE7634DCCEFCE2" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Orphnebius bicuspis Assing 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="56" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="55" updateTime="1699235685730" updateUser="ExternalLinkService">
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<mods:title id="FCE406AB0158961FB2E333E97938DB66">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="3739F8C6EA0C3729651AB31DDE0FD731">Assing, Volker</mods:namePart>
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<mods:title id="0445941D2B8656C59A5EEF386B216614">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="25AA21D477B4F21CDA980FD7E3267930">2016</mods:date>
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<treatment id="766F7C36FF9BFFC0FCEE7634DCCEFCE2" ID-DOI="http://doi.org/10.5281/zenodo.5903422" ID-GBIF-Taxon="192602834" ID-Zenodo-Dep="5903422" LSID="urn:lsid:plazi:treatment:766F7C36FF9BFFC0FCEE7634DCCEFCE2" httpUri="http://treatment.plazi.org/id/766F7C36FF9BFFC0FCEE7634DCCEFCE2" lastPageId="43" lastPageNumber="56" pageId="42" pageNumber="55">
<subSubSection id="B6DC9EABFF9BFFC1FCEE7634DCDDFCAD" box="[869,1189,787,814]" pageId="42" pageNumber="55" type="nomenclature">
<paragraph id="FE79CD20FF9BFFC1FCEE7634DCDDFCAD" blockId="42.[869,1189,787,847]" box="[869,1189,787,814]" pageId="42" pageNumber="55">
<heading id="A5317A4CFF9BFFC1FCEE7634DCDDFCAD" box="[869,1189,787,814]" centered="true" fontSize="11" level="2" pageId="42" pageNumber="55" reason="3">
<taxonomicName id="39C6B6A3FF9BFFC1FCEE7634DC4DFCAD" ID-CoL="8Y799" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[869,1077,787,814]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="55" phylum="Arthropoda" rank="species" species="bicuspis" status="sp. nov.">
<emphasis id="CCB21132FF9BFFC1FCEE7634DC4DFCAD" box="[869,1077,787,814]" italics="true" pageId="42" pageNumber="55">Orphnebius bicuspis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FF9BFFC1FBB77634DCDDFCAD" box="[1084,1189,787,814]" pageId="42" pageNumber="55" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF9BFFC1FCEE761EDC20FCCC" box="[869,1112,825,847]" pageId="42" pageNumber="55" type="description">
<paragraph id="FE79CD20FF9BFFC1FCEE761EDC20FCCC" blockId="42.[869,1189,787,847]" box="[869,1112,825,847]" pageId="42" pageNumber="55">
(
<figureCitation id="66FDD1A5FF9BFFC1FCE1761EDBB9FCCC" box="[874,961,825,847]" captionStart="Figs 1–26" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 1–26. Antenna (1–21) and forebody (22–26) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 22–26: 1.0 mm; 1–21: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="42" pageNumber="55">Figs 9–10</figureCitation>
, 
<figureCitation id="66FDD1A5FF9BFFC1FC43761EDB98FCCC" box="[968,992,825,847]" captionStart="Figs 27–39" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 27–39. Forebody (27–34), abdomen (35–37), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 27–37: 1.0 mm; 38–49: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="42" pageNumber="55">34</figureCitation>
, 
<figureCitation id="66FDD1A5FF9BFFC1FC63761EDC78FCCC" box="[1000,1024,825,847]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="42" pageNumber="55">44</figureCitation>
, 
<figureCitation id="66FDD1A5FF9BFFC1FB8C761EDC2BFCCC" box="[1031,1107,825,847]" captionStart-0="Figs 236–257" captionStart-1="Figs 258–276" captionStartId-0="21.[151,191,1971,1993]" captionStartId-1="22.[189,229,1970,1992]" captionTargetBox-0="[151,1397,239,1961]" captionTargetBox-1="[192,1432,240,1938]" captionTargetId-0="figure-144@21.[151,1398,232,1971]" captionTargetId-1="figure-135@22.[189,1436,232,1954]" captionTargetPageId-0="21" captionText-0="Figs 236–257. Orphnebius serratus (236–243), O. integer (244–250), O. latitibialis (251–252), and O. bicuspis from Laos (253– 257): median lobe of aedeagus in lateral and in ventral view (236–237, 244–245, 253–254); apex of median lobe in ventral view (238, 246, 255); paramere (239, 247, 256); tergite VIII (240, 249, 252); spermatheca (241–243, 251, 257); apex of paramerite (248); postero-median portion of tergite VIII (250). Scale bars: 0.2 mm." captionText-1="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6421202" figureDoi-1="http://doi.org/10.5281/zenodo.6421204" httpUri-0="https://zenodo.org/record/6421202/files/figure.png" httpUri-1="https://zenodo.org/record/6421204/files/figure.png" pageId="42" pageNumber="55">253–260</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF9BFFC1FCB4764EDCC8FB42" pageId="42" pageNumber="55" type="materials_examined">
<paragraph id="FE79CD20FF9BFFC1FCB4764EDCFBFC62" blockId="42.[831,1437,873,1217]" pageId="42" pageNumber="55">
<emphasis id="CCB21132FF9BFFC1FCB4764EDBAEFC02" bold="true" box="[831,982,873,897]" pageId="42" pageNumber="55">Type material</emphasis>
: 
<materialsCitation id="4EAEC77DFF9BFFC1FC68764EDCFBFC62" ID-GBIF-Occurrence="3455118419" collectingDate="2002-05-11" collectingDateMax="2002-05-21" collectingDateMin="2002-05-11" collectionCode="NHMB" collectorName="Vit Kuban" determinerName="V. Assing" elevation="750" latitude="19.583334" location="Laos-N" longLatPrecision="1271" longitude="101.96667" pageId="42" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Louangphabang" typeStatus="holotype">
<typeStatus id="217D7382FF9BFFC1FC68764EDC3EFC02" box="[995,1094,873,897]" pageId="42" pageNumber="55" type="holotype">Holotype</typeStatus>
<specimenCount id="E8C006A9FF9BFFC1FBC6764DDC1AFC02" box="[1101,1122,874,897]" pageId="42" pageNumber="55" type="male">♂</specimenCount>
: “ 
<location id="FB199BFBFF9BFFC1FBFC764EDCBFFC02" LSID="urn:lsid:plazi:treatment:766F7C36FF9BFFC0FCEE7634DCCEFCE2:FB199BFBFF9BFFC1FBFC764EDCBFFC02" box="[1143,1223,873,897]" latitude="19.583334" longLatPrecision="1271" longitude="101.96667" name="Laos-N" pageId="42" pageNumber="55" stateProvince="Louangphabang">Laos-N</location>
(
<location id="FB199BFBFF9BFFC1FB5E764EDDE9FC02" LSID="urn:lsid:plazi:treatment:766F7C36FF9BFFC0FCEE7634DCCEFCE2:FB199BFBFF9BFFC1FB5E764EDDE9FC02" box="[1237,1425,873,897]" latitude="19.583334" longLatPrecision="1271" longitude="101.96667" name="Louangphrabang" pageId="42" pageNumber="55" stateProvince="Louangphabang">Louangphrabang</location>
), 
<date id="8A78EBE0FF9BFFC1FCCB76AEDBB4FC22" box="[832,972,905,929]" pageId="42" pageNumber="55" value="2002-05-11" valueMax="2002-05-21" valueMin="2002-05-11">
<collectingDate id="9A3C1208FF9BFFC1FCCB76AEDBB4FC22" box="[832,972,905,929]" pageId="42" pageNumber="55" value="2002-05-11" valueMax="2002-05-21" valueMin="2002-05-11">11–21.v.2002</collectingDate>
</date>
, 
<geoCoordinate id="9BF2ABE7FF9BFFC1FC5176AEDC48FC22" box="[986,1072,905,929]" degrees="19" direction="north" minutes="35" orientation="latitude" pageId="42" pageNumber="55" precision="925" value="19.583334">19°35'N</geoCoordinate>
, 
<geoCoordinate id="9BF2ABE7FF9BFFC1FBB576AEDCE6FC22" box="[1086,1182,905,929]" degrees="101" direction="east" minutes="58" orientation="longitude" pageId="42" pageNumber="55" precision="925" value="101.96667">101°58'E</geoCoordinate>
, 
<location id="FB199BFBFF9BFFC1FB2076AEDD4FFC22" LSID="urn:lsid:plazi:treatment:766F7C36FF9BFFC0FCEE7634DCCEFCE2:FB199BFBFF9BFFC1FB2076AEDD4FFC22" box="[1195,1335,905,929]" latitude="19.583334" longLatPrecision="1271" longitude="101.96667" name="Thong Khan" pageId="42" pageNumber="55" stateProvince="Louangphabang">Thong Khan</location>
, ~ 
<quantity id="393E60C5FF9BFFC1FAD976AEDDE0FC22" box="[1362,1432,905,929]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="42" pageNumber="55" unit="m" value="750.0">
<elevation id="75EB2A13FF9BFFC1FAD976AEDDE0FC22" box="[1362,1432,905,929]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="42" pageNumber="55" unit="m" value="750.0">750 m</elevation>
</quantity>
, 
<collectorName id="5333A8F6FF9BFFC1FCB4768EDBD5FC42" box="[831,941,937,961]" pageId="42" pageNumber="55">Vít Kubáň</collectorName>
leg. / Holotypus ♂ 
<taxonomicName id="39C6B6A3FF9BFFC1FB08768EDD2FFC42" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[1155,1367,937,961]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="55" phylum="Arthropoda" rank="species" species="bicuspis" status="sp. nov.">
<emphasis id="CCB21132FF9BFFC1FB08768EDD2FFC42" box="[1155,1367,937,961]" italics="true" pageId="42" pageNumber="55">Orphnebius bicuspis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FF9BFFC1FAD5768DDDEFFC42" box="[1374,1431,938,961]" pageId="42" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
, det. 
<determinerName id="0BA521E4FF9BFFC1FCE676EDDBB7FC62" box="[877,975,969,993]" pageId="42" pageNumber="55">V. Assing</determinerName>
2015” (
<collectionCode id="98D755E5FF9BFFC1FBAA76EEDC0FFC62" box="[1057,1143,969,993]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="42" pageNumber="55">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="FE79CD20FF9BFFC1FCB476CDDCC8FB42" blockId="42.[831,1437,873,1217]" pageId="42" pageNumber="55">
<materialsCitation id="4EAEC77DFF9BFFC1FCB476CDDB99FBA2" ID-GBIF-Occurrence="3455118517" collectingDate="2002-05-11" collectingDateMax="2002-05-21" collectingDateMin="2002-05-11" collectionCode="NHMB" collectorName="Vit Kuban" determinerName="V. Assing" elevation="750" latitude="19.583334" location="Louangphrabang" longLatPrecision="1271" longitude="101.96667" pageId="42" pageNumber="55" specimenCount="12" stateProvince="Louangphabang" typeStatus="paratype">
<typeStatus id="217D7382FF9BFFC1FCB476CDDBD1FB81" box="[831,937,1002,1026]" pageId="42" pageNumber="55" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="E8C006A9FF9BFFC1FC3176CEDC7BFB82" box="[954,1027,1001,1025]" pageId="42" pageNumber="55" type="generic">12 exs.</specimenCount>
[3 teneral]: same data as holotype (
<collectionCode id="98D755E5FF9BFFC1FCCC712EDBE2FBA2" box="[839,922,1033,1057]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="42" pageNumber="55">NHMB</collectionCode>
, cAss)
</materialsCitation>
; 
<materialsCitation id="4EAEC77DFF9BFFC1FC65712EDD75FBE2" ID-GBIF-Occurrence="3455118534" collectingDate="1999-05-01" collectingDateMax="1999-05-16" collectingDateMin="1999-05-01" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos" elevation="1200" location="Ban Song Cha" longitude="102.23333" pageId="42" pageNumber="55" specimenCount="2" specimenCount-female="2" stateProvince="Louangphrabang" typeStatus="paratype">
<specimenCount id="E8C006A9FF9BFFC1FC65712EDC50FBA2" box="[1006,1064,1033,1057]" pageId="42" pageNumber="55" type="female">2 ♀♀</specimenCount>
: “ 
<collectingCountry id="86D18DB0FF9BFFC1FBB5712EDC0BFBA2" box="[1086,1139,1033,1057]" name="Laos" pageId="42" pageNumber="55">Laos</collectingCountry>
, 
<date id="8A78EBE0FF9BFFC1FBF4712EDD7DFBA2" box="[1151,1285,1033,1057]" pageId="42" pageNumber="55" value="1999-05-01" valueMax="1999-05-16" valueMin="1999-05-01">
<collectingDate id="9A3C1208FF9BFFC1FBF4712EDD7DFBA2" box="[1151,1285,1033,1057]" pageId="42" pageNumber="55" value="1999-05-01" valueMax="1999-05-16" valueMin="1999-05-01">1.–16.v.1999</collectingDate>
</date>
, 
<collectingRegion id="3C0203C2FF9BFFC1FA9A712EDB0BFBC2" country="Laos" name="Louangphabang" pageId="42" pageNumber="55">Louangphrabang</collectingRegion>
pr., 20°33–4'N 102°14'E, 
<location id="FB199BFBFF9BFFC1FB02710DDD5DFBC2" LSID="urn:lsid:plazi:treatment:766F7C36FF9BFFC0FCEE7634DCCEFCE2:FB199BFBFF9BFFC1FB02710DDD5DFBC2" box="[1161,1317,1065,1089]" country="Laos" longitude="102.23333" name="Ban Song Cha" pageId="42" pageNumber="55" stateProvince="Louangphrabang">Ban Song Cha</location>
(
<locationDeviation id="EFCBD402FF9BFFC1FABD710EDDE8FBC2" box="[1334,1424,1065,1089]" pageId="42" pageNumber="55">
<quantity id="393E60C5FF9BFFC1FABD710EDD15FBC2" box="[1334,1389,1065,1089]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="55" unit="km" value="5.0">5 km</quantity>
W
</locationDeviation>
), 
<quantity id="393E60C5FF9BFFC1FCB4716EDBE9FBE2" box="[831,913,1097,1121]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="42" pageNumber="55" unit="m" value="1200.0">
<elevation id="75EB2A13FF9BFFC1FCB4716EDBE9FBE2" box="[831,913,1097,1121]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="42" pageNumber="55" unit="m" value="1200.0">1200 m</elevation>
</quantity>
, 
<collectorName id="5333A8F6FF9BFFC1FC2F716EDC62FBE2" box="[932,1050,1097,1121]" pageId="42" pageNumber="55">Vít Kubáň</collectorName>
leg.” (
<collectionCode id="98D755E5FF9BFFC1FBE0716EDCC6FBE2" box="[1131,1214,1097,1121]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="42" pageNumber="55">NHMB</collectionCode>
, cAss)
</materialsCitation>
; 
<materialsCitation id="4EAEC77DFF9BFFC1FAAB716EDCD4FB42" ID-GBIF-Occurrence="3455118500" collectingDate="2012-05-12" collectingDateMax="2012-05-25" collectingDateMin="2012-05-12" collectingMethod="flight interception trap" collectionCode="ZFMK" collectorName="L. Dembicky" country="India" elevation="700" latitude="28.615557" location="Etalin" longLatPrecision="20" longitude="95.88917" pageId="42" pageNumber="55" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="E8C006A9FF9BFFC1FAAB716EDD21FBE2" box="[1312,1369,1097,1121]" pageId="42" pageNumber="55" type="male">2 ♂♂</specimenCount>
: “NE 
<collectingCountry id="86D18DB0FF9BFFC1FCB4714EDB01FB02" box="[831,889,1129,1153]" name="India" pageId="42" pageNumber="55">India</collectingCountry>
, Arunachal Pr., 
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vicinity, 
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<elevation id="75EB2A13FF9BFFC1FB44714EDD6BFB02" box="[1231,1299,1129,1153]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="42" pageNumber="55" unit="m" value="700.0">700 m</elevation>
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, 
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, 
<geoCoordinate id="9BF2ABE7FF9BFFC1FCB471AEDBCBFB22" box="[831,947,1161,1185]" degrees="95" direction="east" minutes="53" orientation="longitude" pageId="42" pageNumber="55" precision="15" seconds="21" value="95.88917">95°53'21&quot;E</geoCoordinate>
, FIT (
<collectingMethod id="2787B537FF9BFFC1FC7071AEDC82FB22" box="[1019,1274,1161,1185]" pageId="42" pageNumber="55">flight interception trap</collectingMethod>
), 
<collectorName id="5333A8F6FF9BFFC1FA9B71ADDDE4FB22" box="[1296,1436,1161,1185]" pageId="42" pageNumber="55">L. Dembický</collectorName>
leg., 
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<collectingDate id="9A3C1208FF9BFFC1FCFB718EDC7AFB42" box="[880,1026,1193,1217]" pageId="42" pageNumber="55" value="2012-05-12" valueMax="2012-05-25" valueMin="2012-05-12">12.–25.v.2012</collectingDate>
</date>
” (
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, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF9BFFC1FCB471CEDD73FAA2" pageId="42" pageNumber="55" type="etymology">
<paragraph id="FE79CD20FF9BFFC1FCB471CEDD73FAA2" blockId="42.[831,1437,1257,1313]" pageId="42" pageNumber="55">
<emphasis id="CCB21132FF9BFFC1FCB471CEDBCEFA82" bold="true" box="[831,950,1257,1281]" pageId="42" pageNumber="55">Etymology</emphasis>
: The specific epithet (Latin, adjective) alludes to the bifid ventral process of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF9BFFC0FCB4706EDD73FE42" lastPageId="43" lastPageNumber="56" pageId="42" pageNumber="55" type="description">
<paragraph id="FE79CD20FF9BFFC1FCB4706EDD28F9A2" blockId="42.[831,1438,1353,2049]" pageId="42" pageNumber="55">
<emphasis id="CCB21132FF9BFFC1FCB4706EDBBCFAE2" bold="true" box="[831,964,1353,1377]" pageId="42" pageNumber="55">Description</emphasis>
: Body length 
<quantity id="393E60C5FF9BFFC1FBEC706EDC93FAE2" box="[1127,1259,1353,1377]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="4.1" metricValueMin="2.7" pageId="42" pageNumber="55" unit="mm" value="3.4" valueMax="4.1" valueMin="2.7">2.7–4.1 mm</quantity>
; length of forebody 
<quantity id="393E60C5FF9BFFC1FCF0704EDC78FA02" box="[891,1024,1385,1409]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="42" pageNumber="55" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
. Coloration: forebody black; abdomen red; legs with the femora blackish-brown, the tibae brown to dark-brown, and the tarsi reddish; antennae blackish with antennomeres II–III often, rarely also IV, slightly paler (brown to dark-brown); maxillary palpi pale-brown with the apical palpomere yellowish.
</paragraph>
<paragraph id="FE79CD20FF9BFFC1FCB4730EDBEEF782" blockId="42.[831,1438,1353,2049]" pageId="42" pageNumber="55">
Head (
<figureCitation id="66FDD1A5FF9BFFC1FC06730EDBA7F9C2" box="[909,991,1577,1601]" captionStart="Figs 27–39" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 27–39. Forebody (27–34), abdomen (35–37), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 27–37: 1.0 mm; 38–49: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="42" pageNumber="55">Figs 34</figureCitation>
, 
<figureCitation id="66FDD1A5FF9BFFC1FC64730EDC73F9C2" box="[1007,1035,1577,1601]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="42" pageNumber="55">44</figureCitation>
) of transversely rectangular shape, approximately 1.4 times as broad as long; posterior angles nearly obsolete; posterior margin truncate or weakly convex; dorsal surface nearly impunctate. Eyes large, more than twice as long as postocular region in dorsal view. Antenna (
<figureCitation id="66FDD1A5FF9BFFC1FBF173EEDC9FF962" box="[1146,1255,1737,1761]" captionStart="Figs 1–26" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 1–26. Antenna (1–21) and forebody (22–26) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 22–26: 1.0 mm; 1–21: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="42" pageNumber="55">Figs 9–10</figureCitation>
) approximately 1.0 mm long; antennomere I very short, approximately as long as the combined length of antennomeres II and III; antennomere IV distinctly transverse and flattened; antennomere V much broader than antennomere IV; antennomeres V–X distinctly asymmetric, moderately flattened, and strongly transverse, more than twice as broad as long, and gradually increasing in width; XI approximately as long as the combined length of VIII–X.
</paragraph>
<paragraph id="FE79CD20FF9AFFC0FF1C75CDD969FE62" blockId="43.[151,758,233,1249]" pageId="43" pageNumber="56">
Pronotum (
<figureCitation id="66FDD1A5FF9AFFC0FE9975CED91AFE82" box="[274,354,233,257]" captionStart="Figs 27–39" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 27–39. Forebody (27–34), abdomen (35–37), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 27–37: 1.0 mm; 38–49: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="43" pageNumber="56">Figs 34</figureCitation>
, 
<figureCitation id="66FDD1A5FF9AFFC0FEE775CED9FFFE82" box="[364,391,233,257]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="43" pageNumber="56">44</figureCitation>
) approximately 1.4 times as broad as long and as broad as head, moderately convex in crosssection; lateral and posterior margins together nearly forming a semi-circle, posterior angles completely obsolete; disc with a median pair of punctures, otherwise practically impunctate; lateral margins with one long and stout, and with two moderately long and rather fine erect black setae.
</paragraph>
<paragraph id="FE79CD20FF9AFFC0FF1C74CED984FD02" blockId="43.[151,758,233,1249]" pageId="43" pageNumber="56">
Elytra (
<figureCitation id="66FDD1A5FF9AFFC0FF6E74CED94CFD82" box="[229,308,489,513]" captionStart="Figs 27–39" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 27–39. Forebody (27–34), abdomen (35–37), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 27–37: 1.0 mm; 38–49: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="43" pageNumber="56">Figs 34</figureCitation>
, 
<figureCitation id="66FDD1A5FF9AFFC0FEB574CED921FD82" box="[318,345,489,513]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="43" pageNumber="56">44</figureCitation>
) 0.70–0.77 times as long as pronotum; punctation sparse and fine; pubescence long, fine, pale, and sub-erect or depressed. Hind wings fully developed. Tibiae unmodified. Metatarsomere I slightly shorter than the combined length of II and III.
</paragraph>
<paragraph id="FE79CD20FF9AFFC0FF1C77AED982FC02" blockId="43.[151,758,233,1249]" pageId="43" pageNumber="56">
Abdomen: tergites III–VI with a lateral setiferous puncture on either side, VI with four additional punctures at posterior margin; tergite VII with dense non-setiferous punctures in posterior two-thirds, posterior margin with palisade fringe; tergite VIII (
<figureCitation id="66FDD1A5FF9AFFC0FD8A762EDA20FCA2" box="[513,600,777,801]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="43" pageNumber="56">Fig. 260</figureCitation>
) with a lateral cluster of fine setae on either side, posterior margin truncate in the middle and without teeth; sternite VIII with broadly convex posterior margin.
</paragraph>
<paragraph id="FE79CD20FF9AFFC0FF1C76ADDA66FBE2" blockId="43.[151,758,233,1249]" pageId="43" pageNumber="56">
<specimenCount id="E8C006A9FF9AFFC0FF1C76ADD8D4FC22" box="[151,172,906,929]" pageId="43" pageNumber="56" type="male">♂</specimenCount>
: segments IX–X distinctly modified (of the 
<taxonomicName id="39C6B6A3FF9AFFC0FD0E76AEDA8BFC22" authorityName="Eppelsheim" authorityYear="1895" box="[645,755,905,929]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="CCB21132FF9AFFC0FD0E76AEDA8BFC22" box="[645,755,905,929]" italics="true" pageId="43" pageNumber="56">O. hauseri</emphasis>
</taxonomicName>
<typeStatus id="217D7382FF9AFFC0FF1C768ED8B2FC42" box="[151,202,937,961]" pageId="43" pageNumber="56">type</typeStatus>
), with dense and long pubescence; median lobe of aedeagus (
<figureCitation id="66FDD1A5FF9AFFC0FEB276EED9B4FC62" box="[313,460,969,993]" captionStart="Figs 236–257" captionStartId="21.[151,191,1971,1993]" captionTargetBox="[151,1397,239,1961]" captionTargetId="figure-144@21.[151,1398,232,1971]" captionTargetPageId="21" captionText="Figs 236–257. Orphnebius serratus (236–243), O. integer (244–250), O. latitibialis (251–252), and O. bicuspis from Laos (253– 257): median lobe of aedeagus in lateral and in ventral view (236–237, 244–245, 253–254); apex of median lobe in ventral view (238, 246, 255); paramere (239, 247, 256); tergite VIII (240, 249, 252); spermatheca (241–243, 251, 257); apex of paramerite (248); postero-median portion of tergite VIII (250). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421202" httpUri="https://zenodo.org/record/6421202/files/figure.png" pageId="43" pageNumber="56">Figs 253–255</figureCitation>
, 
<figureCitation id="66FDD1A5FF9AFFC0FE6976EEDA38FC62" box="[482,576,969,993]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="43" pageNumber="56">258–259</figureCitation>
) approximately 
<quantity id="393E60C5FF9AFFC0FF1C76CED892FB82" box="[151,234,1001,1025]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="43" pageNumber="56" unit="mm" value="0.7">0.7 mm</quantity>
long; ventral process short and bifid; paramere (
<figureCitation id="66FDD1A5FF9AFFC0FF2B712ED881FBA2" box="[160,249,1033,1057]" captionStart="Figs 236–257" captionStartId="21.[151,191,1971,1993]" captionTargetBox="[151,1397,239,1961]" captionTargetId="figure-144@21.[151,1398,232,1971]" captionTargetPageId="21" captionText="Figs 236–257. Orphnebius serratus (236–243), O. integer (244–250), O. latitibialis (251–252), and O. bicuspis from Laos (253– 257): median lobe of aedeagus in lateral and in ventral view (236–237, 244–245, 253–254); apex of median lobe in ventral view (238, 246, 255); paramere (239, 247, 256); tergite VIII (240, 249, 252); spermatheca (241–243, 251, 257); apex of paramerite (248); postero-median portion of tergite VIII (250). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421202" httpUri="https://zenodo.org/record/6421202/files/figure.png" pageId="43" pageNumber="56">Fig. 256</figureCitation>
) approximately 
<quantity id="393E60C5FF9AFFC0FE30712EDA76FBA2" box="[443,526,1033,1057]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="43" pageNumber="56" unit="mm" value="0.6">0.6 mm</quantity>
long, with slender condylite; paramerite of distinctive shape, external margin notched at apical two-fifths.
</paragraph>
<paragraph id="FE79CD20FF9AFFC0FF1C714DD9A5FB62" blockId="43.[151,758,233,1249]" pageId="43" pageNumber="56">
<specimenCount id="E8C006A9FF9AFFC0FF1C714DD8D4FB02" box="[151,172,1130,1153]" pageId="43" pageNumber="56" type="female">♀</specimenCount>
: segments IX–X distinctly modified (
<taxonomicName id="39C6B6A3FF9AFFC0FDCA714EDAD7FB02" authorityName="Eppelsheim" authorityYear="1895" box="[577,687,1129,1153]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="CCB21132FF9AFFC0FDCA714EDAD7FB02" box="[577,687,1129,1153]" italics="true" pageId="43" pageNumber="56">O. hauseri</emphasis>
</taxonomicName>
<typeStatus id="217D7382FF9AFFC0FD3E714EDA9FFB02" box="[693,743,1129,1153]" pageId="43" pageNumber="56">type</typeStatus>
), with dense and long pubescence; proximal portion of spermathecal capsule simply curved, near distal portion distinctly modified (
<figureCitation id="66FDD1A5FF9AFFC0FEFE71EED9AAFB62" box="[373,466,1225,1249]" captionStart="Figs 236–257" captionStartId="21.[151,191,1971,1993]" captionTargetBox="[151,1397,239,1961]" captionTargetId="figure-144@21.[151,1398,232,1971]" captionTargetPageId="21" captionText="Figs 236–257. Orphnebius serratus (236–243), O. integer (244–250), O. latitibialis (251–252), and O. bicuspis from Laos (253– 257): median lobe of aedeagus in lateral and in ventral view (236–237, 244–245, 253–254); apex of median lobe in ventral view (238, 246, 255); paramere (239, 247, 256); tergite VIII (240, 249, 252); spermatheca (241–243, 251, 257); apex of paramerite (248); postero-median portion of tergite VIII (250). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421202" httpUri="https://zenodo.org/record/6421202/files/figure.png" pageId="43" pageNumber="56">Fig. 257</figureCitation>
).
</paragraph>
<paragraph id="FE79CD20FF9AFFC0FF1C702EDA75FA62" blockId="43.[151,758,1289,1505]" pageId="43" pageNumber="56">
<emphasis id="CCB21132FF9AFFC0FF1C702ED9EBFAA2" bold="true" box="[151,403,1289,1313]" pageId="43" pageNumber="56">Intraspecific variation</emphasis>
: There are slight differences in the shape of the bifid apex of the ventral process of the aedeagus of the males from Arunachal Pradesh and from 
<collectingCountry id="86D18DB0FF9AFFC0FF58704DD971FA02" box="[211,265,1386,1409]" name="Laos" pageId="43" pageNumber="56">Laos</collectingCountry>
. However, the external characters (
<figureCitation id="66FDD1A5FF9AFFC0FD08704EDA97FA02" box="[643,751,1385,1409]" captionStart="Figs 1–26" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 1–26. Antenna (1–21) and forebody (22–26) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 22–26: 1.0 mm; 1–21: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="43" pageNumber="56">Figs 9–10</figureCitation>
, 
<figureCitation id="66FDD1A5FF9AFFC0FF1C70AED8CBFA22" box="[151,179,1417,1441]" captionStart="Figs 27–39" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 27–39. Forebody (27–34), abdomen (35–37), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 27–37: 1.0 mm; 38–49: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="43" pageNumber="56">34</figureCitation>
, 
<figureCitation id="66FDD1A5FF9AFFC0FF4B70AED8A4FA22" box="[192,220,1417,1441]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="43" pageNumber="56">44</figureCitation>
) and the shape of the paramere are identical, so that the observed differences are attributed to intrarather than interspecific variation.
</paragraph>
<paragraph id="FE79CD20FF9AFFC0FF1C732EDD73FE42" blockId="43.[151,757,1545,2081]" lastBlockId="43.[794,1399,233,449]" pageId="43" pageNumber="56">
<emphasis id="CCB21132FF9AFFC0FF1C732ED913F9A2" bold="true" box="[151,363,1545,1569]" pageId="43" pageNumber="56">Comparative notes</emphasis>
: Based on the shape of the head, the antennae, and the pronotum, the lateral clusters of setae on the abdominal tergite VIII, as well as on the bifid ventral process of the aedeagus and the morphology of the spermatheca, 
<taxonomicName id="39C6B6A3FF9AFFC0FED773AED9A8F922" authorityName="Assing" authorityYear="2016" box="[348,464,1673,1697]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="CCB21132FF9AFFC0FED773AED9A8F922" box="[348,464,1673,1697]" italics="true" pageId="43" pageNumber="56">O. bicuspis</emphasis>
</taxonomicName>
belongs to the 
<taxonomicName id="39C6B6A3FF9AFFC0FD0B73AEDA8CF922" authorityName="Assing" authorityYear="2016" box="[640,756,1673,1697]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="CCB21132FF9AFFC0FD0B73AEDA8CF922" box="[640,756,1673,1697]" italics="true" pageId="43" pageNumber="56">O. serratus</emphasis>
</taxonomicName>
subgroup. It is distinguished from 
<taxonomicName id="39C6B6A3FF9AFFC0FD9B738EDAFDF942" authorityName="Assing" authorityYear="2016" box="[528,645,1705,1729]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="CCB21132FF9AFFC0FD9B738EDAFDF942" box="[528,645,1705,1729]" italics="true" pageId="43" pageNumber="56">O. serratus</emphasis>
</taxonomicName>
and other representatives of this subgroup by the non-dentate posterior margin of tergite VIII, as well as by the different shapes of the median lobe and the parameres of the aedeagus, from 
<taxonomicName id="39C6B6A3FF9AFFC0FEFD720ED992F8C2" authorityName="Assing" authorityYear="2016" box="[374,490,1833,1857]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="CCB21132FF9AFFC0FEFD720ED992F8C2" box="[374,490,1833,1857]" italics="true" pageId="43" pageNumber="56">O. serratus</emphasis>
</taxonomicName>
additionally by smaller average body size, a narrower pronotum, and undilated protibiae. Regarding the bifid ventral process of the aedeagus, as well as the similar head shape, 
<taxonomicName id="39C6B6A3FF9AFFC0FD0872AEDA8CF822" authorityName="Assing" authorityYear="2016" box="[643,756,1929,1953]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="CCB21132FF9AFFC0FD0872AEDA8CF822" box="[643,756,1929,1953]" italics="true" pageId="43" pageNumber="56">O. bicuspis</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="39C6B6A3FF9AFFC0FEAB728EDA7FF842" authority="PACE, 2000" authorityName="PACE" authorityYear="2000" box="[288,519,1961,1985]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="CCB21132FF9AFFC0FEAB728ED9FFF842" box="[288,391,1961,1985]" italics="true" pageId="43" pageNumber="56">O. incisus</emphasis>
<bibRefCitation id="9A57B0D1FF9AFFC0FE06728EDA7FF842" author="PACE, R." box="[397,519,1961,1985]" pageId="43" pageNumber="56" pagination="39 - 86" refId="ref77836" refString="PACE, R. 2000: Aleocharinae della Thailandia (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 17 (1): 39 - 86." type="journal article" year="2000">PACE, 2000</bibRefCitation>
</taxonomicName>
, which was originally described from 
<collectingCountry id="86D18DB0FF9AFFC0FED872EED9CDF862" box="[339,437,1993,2017]" name="Thailand" pageId="43" pageNumber="56">Thailand</collectingCountry>
and subsequently reported also from Yunnan (
<bibRefCitation id="9A57B0D1FF9AFFC0FE0072CED986F782" author="PACE, R." box="[395,510,2025,2049]" pageId="43" pageNumber="56" pagination="39 - 86" refId="ref77836" refString="PACE, R. 2000: Aleocharinae della Thailandia (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 17 (1): 39 - 86." type="journal article" year="2000">PACE 2000</bibRefCitation>
, 
<bibRefCitation id="9A57B0D1FF9AFFC0FD9F72CEDA23F782" author="PACE, R." box="[532,603,2025,2049]" pageId="43" pageNumber="56" pagination="283 - 296" refId="ref78055" refString="PACE, R. 2012 b: New data, two new genera and new species of Aleocharinae from Indian and Chinese subregions (Coleoptera: Staphylinidae). - In: HARTMANN, M. &amp; WEIPERT, J. (eds), Biodiversitat und Naturausstattung im Himalaya, Erfurt 4: 283 - 296." type="journal article" year="2012">2012b</bibRefCitation>
). 
<taxonomicName id="39C6B6A3FF9AFFC0FDF372CED893F7A2" authorityName="Assing" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="CCB21132FF9AFFC0FDF372CED893F7A2" italics="true" pageId="43" pageNumber="56">Orphnebius bicuspis</emphasis>
</taxonomicName>
is distinguished from this species by slightly larger size (
<taxonomicName id="39C6B6A3FF9AFFC0FC2975CEDC71FE82" authorityName="PACE" authorityYear="2000" box="[930,1033,233,257]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="CCB21132FF9AFFC0FC2975CEDC71FE82" box="[930,1033,233,257]" italics="true" pageId="43" pageNumber="56">O. incisus</emphasis>
</taxonomicName>
: 
<quantity id="393E60C5FF9AFFC0FB9075CEDC0AFE82" box="[1051,1138,233,257]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="43" pageNumber="56" unit="mm" value="3.4">3.4 mm</quantity>
), the coloration of the antennae (
<taxonomicName id="39C6B6A3FF9AFFC0FC1B742EDB8FFEA2" authorityName="PACE" authorityYear="2000" box="[912,1015,265,289]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="CCB21132FF9AFFC0FC1B742EDB8FFEA2" box="[912,1015,265,289]" italics="true" pageId="43" pageNumber="56">O. incisus</emphasis>
</taxonomicName>
: antennomeres I–V brown, VI–XI black), the more transverse head, by the larger median lobe of the aedeagus (
<taxonomicName id="39C6B6A3FF9AFFC0FB84746EDC0EFEE2" authorityName="PACE" authorityYear="2000" box="[1039,1142,329,353]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="CCB21132FF9AFFC0FB84746EDC0EFEE2" box="[1039,1142,329,353]" italics="true" pageId="43" pageNumber="56">O. incisus</emphasis>
</taxonomicName>
: &lt;
<quantity id="393E60C5FF9AFFC0FB16746EDC8AFEE2" box="[1181,1266,329,353]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="43" pageNumber="56" unit="mm" value="0.5">0.5 mm</quantity>
, if the scale and the drawings are correct) and by the shorter and more broadly separated apices of the ventral process. For illustrations of 
<taxonomicName id="39C6B6A3FF9AFFC0FC61748EDC2AFE42" authorityName="PACE" authorityYear="2000" box="[1002,1106,425,449]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="CCB21132FF9AFFC0FC61748EDC2AFE42" box="[1002,1106,425,449]" italics="true" pageId="43" pageNumber="56">O. incisus</emphasis>
</taxonomicName>
see 
<bibRefCitation id="9A57B0D1FF9AFFC0FB0B748EDD70FE42" author="PACE, R." box="[1152,1288,425,449]" pageId="43" pageNumber="56" pagination="39 - 86" refId="ref77836" refString="PACE, R. 2000: Aleocharinae della Thailandia (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 17 (1): 39 - 86." type="journal article" year="2000">PACE (2000)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF9AFFC0FC9174CEDCCEFCE2" pageId="43" pageNumber="56" type="distribution">
<paragraph id="FE79CD20FF9AFFC0FC9174CEDCCEFCE2" blockId="43.[794,1400,489,866]" pageId="43" pageNumber="56">
<emphasis id="CCB21132FF9AFFC0FC9174CEDCF4FD82" bold="true" box="[794,1164,489,513]" pageId="43" pageNumber="56">Distribution and natural history</emphasis>
: 
<taxonomicName id="39C6B6A3FF9AFFC0FB1774CEDD0EFD82" authorityName="Assing" authorityYear="2016" box="[1180,1398,489,513]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="CCB21132FF9AFFC0FB1774CEDD0EFD82" box="[1180,1398,489,513]" italics="true" pageId="43" pageNumber="56">Orphnebius bicuspis</emphasis>
</taxonomicName>
is known from two localities in 
<collectingRegion id="3C0203C2FF9AFFC0FB36772EDB01FDC2" country="Laos" name="Louangphabang" pageId="43" pageNumber="56">Louangphrabang province</collectingRegion>
(
<collectingCountry id="86D18DB0FF9AFFC0FC05770EDBBDFDC2" box="[910,965,553,577]" name="Laos" pageId="43" pageNumber="56">Laos</collectingCountry>
) and one in Arunachal Pradesh; the 
<typeStatus id="217D7382FF9AFFC0FC91776DDB31FDE1" box="[794,841,586,610]" pageId="43" pageNumber="56">type</typeStatus>
locality is identical to that of 
<taxonomicName id="39C6B6A3FF9AFFC0FB48776EDD4FFDE2" authorityName="Assing" authorityYear="2016" box="[1219,1335,585,609]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="CCB21132FF9AFFC0FB48776EDD4FFDE2" box="[1219,1335,585,609]" italics="true" pageId="43" pageNumber="56">O. serratus</emphasis>
</taxonomicName>
. The specimens were collected at altitudes between 700 and 
<quantity id="393E60C5FF9AFFC0FC9177AEDB14FD22" box="[794,876,649,673]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="43" pageNumber="56" unit="m" value="1200.0">1200 m</quantity>
, in 
<collectingCountry id="86D18DB0FF9AFFC0FC1D77ADDBB1FD22" box="[918,969,650,673]" name="Laos" pageId="43" pageNumber="56">Laos</collectingCountry>
together with 
<taxonomicName id="39C6B6A3FF9AFFC0FBE577AEDC90FD22" authorityName="Assing" authorityYear="2016" box="[1134,1256,649,673]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="dilatatus">
<emphasis id="CCB21132FF9AFFC0FBE577AEDC90FD22" box="[1134,1256,649,673]" italics="true" pageId="43" pageNumber="56">O. dilatatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="39C6B6A3FF9AFFC0FB7C77AEDD08FD22" authorityName="Assing" authorityYear="2016" box="[1271,1392,649,673]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="extensus">
<emphasis id="CCB21132FF9AFFC0FB7C77AEDD08FD22" box="[1271,1392,649,673]" italics="true" pageId="43" pageNumber="56">O. extensus</emphasis>
</taxonomicName>
, 
<taxonomicName id="39C6B6A3FF9AFFC0FC91778EDBF6FD42" authorityName="Assing" authorityYear="2016" box="[794,910,681,705]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="CCB21132FF9AFFC0FC91778EDBF6FD42" box="[794,910,681,705]" italics="true" pageId="43" pageNumber="56">O. serratus</emphasis>
</taxonomicName>
, 
<taxonomicName id="39C6B6A3FF9AFFC0FC23778EDC4AFD42" authorityName="Assing" authorityYear="2016" box="[936,1074,681,705]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="latitibialis">
<emphasis id="CCB21132FF9AFFC0FC23778EDC4AFD42" box="[936,1074,681,705]" italics="true" pageId="43" pageNumber="56">O. latitibialis</emphasis>
</taxonomicName>
, 
<taxonomicName id="39C6B6A3FF9AFFC0FBC6778EDCC3FD42" authorityName="Assing" authorityYear="2016" box="[1101,1211,681,705]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="CCB21132FF9AFFC0FBC6778EDCC3FD42" box="[1101,1211,681,705]" italics="true" pageId="43" pageNumber="56">O. lunatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="39C6B6A3FF9AFFC0FB5D778EDD08FD42" authorityName="Assing" authorityYear="2016" box="[1238,1392,681,705]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="nigrapicalis">
<emphasis id="CCB21132FF9AFFC0FB5D778EDD08FD42" box="[1238,1392,681,705]" italics="true" pageId="43" pageNumber="56">O. nigrapicalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="39C6B6A3FF9AFFC0FC9177EEDBEBFD62" authorityName="Assing" authorityYear="2016" box="[794,915,713,737]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="reductus">
<emphasis id="CCB21132FF9AFFC0FC9177EEDBEBFD62" box="[794,915,713,737]" italics="true" pageId="43" pageNumber="56">O. reductus</emphasis>
</taxonomicName>
, 
<taxonomicName id="39C6B6A3FF9AFFC0FC2C77EEDC6FFD62" box="[935,1047,713,737]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="retunsu">
<emphasis id="CCB21132FF9AFFC0FC2C77EEDC6FFD62" box="[935,1047,713,737]" italics="true" pageId="43" pageNumber="56">O. retunsu</emphasis>
</taxonomicName>
s, 
<taxonomicName id="39C6B6A3FF9AFFC0FBBE77EEDCB8FD62" authorityName="Assing" authorityYear="2016" box="[1077,1216,713,737]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="cultellatus">
<emphasis id="CCB21132FF9AFFC0FBBE77EEDCB8FD62" box="[1077,1216,713,737]" italics="true" pageId="43" pageNumber="56">O. cultellatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="39C6B6A3FF9AFFC0FA8177EEDB42FC82" authorityName="Assing" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="CCB21132FF9AFFC0FA8177EEDB42FC82" italics="true" pageId="43" pageNumber="56">O. carinatus</emphasis>
</taxonomicName>
, and in 
<collectingRegion id="3C0203C2FF9AFFC0FC1F77CEDC1EFC82" box="[916,1126,745,769]" country="India" name="Arunachal Pradesh" pageId="43" pageNumber="56">Arunachal Pradesh</collectingRegion>
together with 
<taxonomicName id="39C6B6A3FF9AFFC0FA8377CEDD00FC82" authorityName="Assing" authorityYear="2016" box="[1288,1400,745,769]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="spinans">
<emphasis id="CCB21132FF9AFFC0FA8377CEDD00FC82" box="[1288,1400,745,769]" italics="true" pageId="43" pageNumber="56">O. spinans</emphasis>
</taxonomicName>
and 
<taxonomicName id="39C6B6A3FF9AFFC0FCDB762EDBC9FCA2" authorityName="Assing" authorityYear="2016" box="[848,945,777,801]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="56" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="CCB21132FF9AFFC0FCDB762EDBC9FCA2" box="[848,945,777,801]" italics="true" pageId="43" pageNumber="56">O. dispar</emphasis>
</taxonomicName>
. At least the material from 
<collectingCountry id="86D18DB0FF9AFFC0FA8C762EDD38FCA2" box="[1287,1344,777,801]" name="India" pageId="43" pageNumber="56">India</collectingCountry>
was collected with a flight interception trap. Three of the 
<typeStatus id="217D7382FF9AFFC0FC91766DDB31FCE1" box="[794,841,842,866]" pageId="43" pageNumber="56">type</typeStatus>
specimens from 
<collectingCountry id="86D18DB0FF9AFFC0FB88766DDC4EFCE2" box="[1027,1078,842,865]" name="Laos" pageId="43" pageNumber="56">Laos</collectingCountry>
are teneral.
</paragraph>
</subSubSection>
</treatment>
</document>