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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.859.35465" ID-GBIF-Dataset="fb287528-7c30-429b-a88d-dc8611c6a48d" ID-PMC="PMC6616038" ID-Pensoft-Pub="1313-2970-859-63" ID-PubMed="31327923" ID-ZooBank="555B043451EC47888E45383068C0CAE8" ModsDocID="1313-2970-859-63" checkinTime="1565091139148" checkinUser="pensoft" docAuthor="Zhao, Qing-Hao, Xu, Wang &amp; Yin, Zi-Wei" docDate="2019" docId="4D798B20CC44559469A67EB28E3128E4" docLanguage="en" docName="ZooKeys 859: 63-68" docOrigin="ZooKeys 859" docSource="http://dx.doi.org/10.3897/zookeys.859.35465" docTitle="Linan qiniangmontis Zhao, Xu &amp; Yin, 2019, sp. nov." docType="treatment" docUuid="122313A9-AF98-429B-BBBE-D13402160D5C" docUuidSource="ZooBank" docVersion="6" lastPageNumber="67" masterDocId="B059FFB2FF86FF954D02DC1DC010FFCF" masterDocTitle="A new species of Linan Hlavacˇ (Coleoptera, Staphylinidae, Pselaphinae) from Shenzhen, China" masterLastPageNumber="68" masterPageNumber="63" pageNumber="64" updateTime="1668167469156" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Linan Hlavacˇ (Coleoptera, Staphylinidae, Pselaphinae) from Shenzhen, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Qing-Hao</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Wang</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Zi-Wei</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>859</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>63</mods:start>
<mods:end>68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.859.35465</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.859.35465</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-859-63</mods:identifier>
<mods:identifier type="ZooBank">555B043451EC47888E45383068C0CAE8</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="158498278" LSID="urn:lsid:zoobank.org:act:122313A9-AF98-429B-BBBE-D13402160D5C" httpUri="http://treatment.plazi.org/id/4D798B20CC44559469A67EB28E3128E4" lastPageId="4" lastPageNumber="67" pageId="1" pageNumber="64">
<subSubSection pageId="1" pageNumber="64" type="nomenclature">
<paragraph pageId="1" pageNumber="64">
<taxonomicName LSID="4D798B20CC44559469A67EB28E3128E4" class="Insecta" family="Staphylinidae" genus="Linan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Linan qiniangmontis" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="qiniangmontis">Linan qiniangmontis</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="64">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Linan qiniangmontis sp. nov. A male B female. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.859.35465.figure1" httpUri="https://binary.pensoft.net/fig/314064" pageId="1" pageNumber="64">Figs 1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="1" pageNumber="64">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="64" type="type material">
<paragraph pageId="1" pageNumber="64">Type material.</paragraph>
<paragraph pageId="1" pageNumber="64">
<emphasis bold="true" pageId="1" pageNumber="64">Holotype: CHINA</emphasis>
: ♂: 'China: Guangdong, Shenzhen City, Mt. Qiniang (七娘山),
<geoCoordinate degrees="23" direction="north" minutes="32" orientation="latitude" precision="1" seconds="28.73" value="23.541313">23°32'28.73&quot;N</geoCoordinate>
,
<geoCoordinate degrees="114" direction="east" minutes="35" orientation="longitude" precision="1" seconds="8.46" value="114.585686">114°35'8.46&quot;E</geoCoordinate>
, mixed leaf litter, sifted, 45 m, 23.III.2019, Tang, Shuai, Zhao, Zhou &amp; Xia leg.' (SNUC).
<emphasis bold="true" pageId="1" pageNumber="64">Paratypes: CHINA</emphasis>
: 1 ♂, 3 ♀♀, same label data as holotype (SNUC).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="64" type="diagnosis">
<paragraph pageId="1" pageNumber="64">Diagnosis.</paragraph>
<paragraph pageId="1" pageNumber="64">Body length slightly less than 2.5 mm. Male: antennal club almost simple, with antennomere IX slightly angulate at anteromesal corner; metaventral processes short and narrowing toward apex; protibiae with small denticle at apex; metacoxae with truncate, curved, ventral projection; aedeagus elongate, median lobe asymmetrically narrowed at apex. Female: identifiable only when in association with a male.</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="66" pageId="1" pageNumber="64" type="description">
<paragraph pageId="1" pageNumber="64">Description.</paragraph>
<paragraph lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64">
Male (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Linan qiniangmontis sp. nov. A male B female. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.859.35465.figure1" httpUri="https://binary.pensoft.net/fig/314064" pageId="1" pageNumber="64">Fig. 1A</figureCitation>
). Body length (combined length of head, pronotum, elytra, and abdomen) 2.32-2.33 mm. Head longer than wide, length from clypeal anterior margin to head base 0.52-0.54 mm, width across eyes 0.48-0.49 mm; eyes small, each composed of ca. 23 facets. Antennae elongate, 1.78-1.79 mm long, scape elongate, ca. 3.5 times as long as wide, antennomeres 2-8 each sub-moniliform, of similar width, antennal club (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="1" pageNumber="64">Fig. 2A</figureCitation>
) formed by antennomeres 9-11, antennomere 9 much longer than wide, broadening from base to apex, angulate at anteromesal corner (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="1" pageNumber="64">Fig. 2A</figureCitation>
, indicated by arrow), antennomere 10 slightly transverse, antennomere 11 truncate and broadest at base and narrowing apically, both antennomeres 10 and 11 simple. Pronotum (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="1" pageNumber="64">Fig. 2B</figureCitation>
) approximately as long as wide, with rounded lateral margins, length along midline 0.49-0.51 mm, maximum width 0.49-0.52 mm. Elytra
<pageBreakToken pageId="2" pageNumber="65" start="start">strongly</pageBreakToken>
transverse, length along suture 0.56-0.57 mm, maximum width 0.85-0.88 mm. Metaventral processes (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="2" pageNumber="65">Fig. 2C</figureCitation>
) short, narrowing apically. Protrochanters and profemora (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="2" pageNumber="65">Fig. 2D</figureCitation>
) simple, protibiae (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="2" pageNumber="65">Fig. 2E</figureCitation>
) with small but distinct denticle at apex; mesotrochanters, mesofemora, and mesotibiae (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="2" pageNumber="65">Fig. 2F</figureCitation>
) simple; metacoxae (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="2" pageNumber="65">Fig. 2G</figureCitation>
) with truncate curved projection on ventral margin; metatrochanters, metafemora, and metatibiae simple. Abdomen approximately as wide as elytra, length of dorsally visible part along midline 0.74-0.77 mm, maximum width 0.86-0.87 mm; tergite 1 (IV) more than twice as long as tergite 2 (V); sternite 7 (IX) (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="2" pageNumber="65">Fig. 2H</figureCitation>
) semi-membranous, elongate. Length of aedeagus (
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male diagnostic features of Linan qiniangmontis sp. nov. A antennal club B pronotum C metaventral process, lateral D protrochanter and profemur E apex of protibia F mesotrochanter and mesofemur G metacoxa, metatrochanter, and metafemur H sternite IX I-K aedeagus, dorsal (I), lateral (J), and ventral (K). Scale bars: 0.3 mm (A, B, D, F, G); 0.2 mm (C, I, J, K); 0.1 mm (H); 0.05 mm (E)." figureDoi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="2" pageNumber="65">
Fig. 2
<normalizedToken originalValue="IK">I-K</normalizedToken>
</figureCitation>
) 0.38-0.40 mm; median lobe asymmetrical dorso-ventrally, narrowing apically with pointed apex; elongate parameres slightly exceeding apex of median lobe, with rounded apices; endophallus with one broad, rounded triangular sclerite, and one much shorter, elongate sclerite forked at apex.
</paragraph>
<caption doi="10.3897/zookeys.859.35465.figure1" httpUri="https://binary.pensoft.net/fig/314064" pageId="2" pageNumber="65" start="Figure 1" startId="F1">
<paragraph pageId="2" pageNumber="65">
Figure 1. Dorsal habitus of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Linan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Linan qiniangmontis" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="qiniangmontis">
<emphasis italics="true" pageId="2" pageNumber="65">Linan qiniangmontis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="2" pageNumber="65">A</emphasis>
male
<emphasis bold="true" pageId="2" pageNumber="65">B</emphasis>
female. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.859.35465.figure2" httpUri="https://binary.pensoft.net/fig/314065" pageId="2" pageNumber="65" start="Figure 2" startId="F2">
<paragraph pageId="2" pageNumber="65">
Figure 2. Male diagnostic features of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Linan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Linan qiniangmontis" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="qiniangmontis">
<emphasis italics="true" pageId="2" pageNumber="65">Linan qiniangmontis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="2" pageNumber="65">A</emphasis>
antennal club
<emphasis bold="true" pageId="2" pageNumber="65">B</emphasis>
pronotum
<emphasis bold="true" pageId="2" pageNumber="65">C</emphasis>
metaventral process, lateral
<emphasis bold="true" pageId="2" pageNumber="65">D</emphasis>
protrochanter and profemur
<emphasis bold="true" pageId="2" pageNumber="65">E</emphasis>
apex of protibia
<emphasis bold="true" pageId="2" pageNumber="65">F</emphasis>
mesotrochanter and mesofemur
<emphasis bold="true" pageId="2" pageNumber="65">G</emphasis>
metacoxa, metatrochanter, and metafemur
<emphasis bold="true" pageId="2" pageNumber="65">H</emphasis>
sternite IX
<emphasis bold="true" pageId="2" pageNumber="65">
<normalizedToken originalValue="IK">I-K</normalizedToken>
</emphasis>
aedeagus, dorsal (
<emphasis bold="true" pageId="2" pageNumber="65">I</emphasis>
), lateral (
<emphasis bold="true" pageId="2" pageNumber="65">J</emphasis>
), and ventral (
<emphasis bold="true" pageId="2" pageNumber="65">K</emphasis>
). Scale bars: 0.3 mm (
<emphasis bold="true" pageId="2" pageNumber="65">A, B, D, F, G</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="2" pageNumber="65">C, I, J, K</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="2" pageNumber="65">H</emphasis>
); 0.05 mm (
<emphasis bold="true" pageId="2" pageNumber="65">E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="66">
<pageBreakToken pageId="3" pageNumber="66" start="start">Female</pageBreakToken>
. Similar to male in general morphology, with slightly shorter antennae and smaller eyes; antennae and legs simple; lacking metaventral processes. Eyes each composed of approximately 18 facets. Measurements (as of male): Body length 2.33-2.44 mm, length/width of head 0.53-0.55 /0.49-0.51 mm, length of antennae 1.63-1.70 mm, length/width of pronotum 0.50-0.51/0.51 mm, length/width of elytra 0.57/0.89-0.91 mm, length/width of abdomen 0.72
<normalizedToken originalValue="0.81/0.92">-0.81/0.92-</normalizedToken>
0.93 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="distribution">
<paragraph pageId="4" pageNumber="67">
<pageBreakToken pageId="4" pageNumber="67" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="67">China: Guangdong.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="etymology">
<paragraph pageId="4" pageNumber="67">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="67">The new species epithet refers to the type locality of the new species, Qiniang Mountain.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="67" type="comparisons">
<paragraph pageId="4" pageNumber="67">Comparative notes.</paragraph>
<paragraph pageId="4" pageNumber="67">
The new species is placed as a member of the
<taxonomicName lsidName="L. chinensis" pageId="4" pageNumber="67" rank="species" species="chinensis">
<emphasis italics="true" pageId="4" pageNumber="67">L. chinensis</emphasis>
</taxonomicName>
group by the almost unmodified antennal clubs in the male. It is most similar to
<taxonomicName lsidName="L. hujiayaoi" pageId="4" pageNumber="67" rank="species" species="hujiayaoi">
<emphasis italics="true" pageId="4" pageNumber="67">L. hujiayaoi</emphasis>
</taxonomicName>
Yin &amp; Li, 2013 and
<taxonomicName lsidName="L. mulunensis" pageId="4" pageNumber="67" rank="species" species="mulunensis">
<emphasis italics="true" pageId="4" pageNumber="67">L. mulunensis</emphasis>
</taxonomicName>
Zhang, Li &amp; Yin, 2018 (both from Guangxi) in sharing modified male metacoxae.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Linan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Linan qiniangmontis" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="qiniangmontis">
<emphasis italics="true" pageId="4" pageNumber="67">Linan qiniangmontis</emphasis>
</taxonomicName>
differs from both known species in the slightly angulate anteromesal corner of antennomere 9 (rounded in
<taxonomicName lsidName="L. hujiayaoi" pageId="4" pageNumber="67" rank="species" species="hujiayaoi">
<emphasis italics="true" pageId="4" pageNumber="67">L. hujiayaoi</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="L. mulunensis" pageId="4" pageNumber="67" rank="species" species="mulunensis">
<emphasis italics="true" pageId="4" pageNumber="67">L. mulunensis</emphasis>
</taxonomicName>
), a different form of the metaventral processes (processes stouter in
<taxonomicName lsidName="L. hujiayaoi" pageId="4" pageNumber="67" rank="species" species="hujiayaoi">
<emphasis italics="true" pageId="4" pageNumber="67">L. hujiayaoi</emphasis>
</taxonomicName>
and much more elongate in
<taxonomicName lsidName="L. mulunensis" pageId="4" pageNumber="67" rank="species" species="mulunensis">
<emphasis italics="true" pageId="4" pageNumber="67">L. mulunensis</emphasis>
</taxonomicName>
), the lack of additional projections above the metacoxae (present in
<taxonomicName lsidName="L. hujiayaoi" pageId="4" pageNumber="67" rank="species" species="hujiayaoi">
<emphasis italics="true" pageId="4" pageNumber="67">L. hujiayaoi</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="L. mulunensis" pageId="4" pageNumber="67" rank="species" species="mulunensis">
<emphasis italics="true" pageId="4" pageNumber="67">L. mulunensis</emphasis>
</taxonomicName>
), and a more elongate aedeagus with a different configuration of the endophallus.
</paragraph>
</subSubSection>
</treatment>
</document>