127 lines
17 KiB
XML
127 lines
17 KiB
XML
<document id="6B51E30EFBD8797B618CBE77981AAB83" ID-DOI="10.5281/zenodo.213150" ID-GBIF-Dataset="c619cb1b-fb4b-4450-a767-1a943b43b8b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="213150" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460460536108" checkinUser="plazi" docAuthor="Vršanský, Peter & Aristov, Danil" docDate="2012" docId="1B41B46C501B7669DE88FB19FEE3A3DA" docLanguage="en" docName="zt03247p031.pdf" docOrigin="Zootaxa 3247" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Mutovia intercalaria Vršanský & Aristov, 2012, sp. n." docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="E778CC14501E766FDE1FFFB3FFE6A30E" masterDocTitle="Enigmatic Late Permian cockroaches from Isady, Russia (Blattida: Mutoviidae fam. n.)" masterLastPageNumber="31" masterPageNumber="19" pageNumber="24" updateTime="1698303704797" updateUser="plazi">
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<mods:mods id="9FF57C6A146AC86F6EB1EBD6C370A5AE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="92D4FE95D782549C0A59CEDB342BE4E6">
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<mods:title id="A89EA85F45E409E0A9F4BEFBAD963AF3">Enigmatic Late Permian cockroaches from Isady, Russia (Blattida: Mutoviidae fam. n.)</mods:title>
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</mods:titleInfo>
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<mods:name id="5A8582B7C9679384FDFA121F4DF1B565" type="personal">
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<mods:roleTerm id="EAF39EAF8FCECBC49EACFFC6839E9FBF">Author</mods:roleTerm>
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<mods:namePart id="B10626B3C4119BF8CB4C5B3991EBE49B">Vršanský, Peter</mods:namePart>
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<mods:name id="FDC48C2EAECF7993BE0A1D65F16918B0" type="personal">
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<mods:roleTerm id="9CB090B6BAF5413A9B32ED5581149BDF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="16E9EC4D7B29DBB57D92118D16E45C4D">Aristov, Danil</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="DDD453BCD3529CE4DF6B3A726C260F6C">text</mods:typeOfResource>
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<mods:titleInfo id="9A5D4295100ED7C9667109D905E0671F">
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<mods:title id="AA9B0F4630591224A64DF7C5068975C8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5C7CFB18EA6577C8DF49BBAD21DFB842">
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<mods:date id="2E46B14251E79B3B74D5A1C6CA595911">2012</mods:date>
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<mods:detail id="01BBF611E31D765F92AD23DEAC752F18" type="volume">
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<mods:number id="1810445F59CD77554B54DAFE1EB9CCF6">3247</mods:number>
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<mods:start id="2F2F246CC4DD27D1011103F692573D62">19</mods:start>
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<mods:end id="F093A2555C3F00F2221C8EDD102D662B">31</mods:end>
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<mods:classification id="F4B1A0C9511EE581644C0C96C034492D">journal article</mods:classification>
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<mods:identifier id="AB4C3A9C6C3D6E395C095FC7B46B7E15" type="DOI">10.5281/zenodo.213150</mods:identifier>
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<mods:identifier id="AE5006898BD3AA5D70268CEEBA12117E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C237ACFD02B6C527F11E901DEAA86973" type="Zenodo-Dep">213150</mods:identifier>
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<treatment id="1B41B46C501B7669DE88FB19FEE3A3DA" ID-DOI="http://doi.org/10.5281/zenodo.5674732" ID-GBIF-Taxon="119512307" ID-Zenodo-Dep="5674732" LSID="urn:lsid:plazi:treatment:1B41B46C501B7669DE88FB19FEE3A3DA" httpUri="http://treatment.plazi.org/id/1B41B46C501B7669DE88FB19FEE3A3DA" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24">
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<subSubSection id="DBF256F1501B766ADE88FB19FEF4A7EB" pageId="5" pageNumber="24" type="nomenclature">
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<paragraph id="9357057A501B766ADE88FB19FE3CA7CA" blockId="5.[151,474,1194,1253]" box="[151,474,1194,1220]" pageId="5" pageNumber="24">
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<heading id="C81FB216501B766ADE88FB19FE3CA7CA" bold="true" box="[151,474,1194,1220]" fontSize="11" level="1" pageId="5" pageNumber="24" reason="1">
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<emphasis id="A19CD968501B766ADE88FB19FE3CA7CA" bold="true" box="[151,474,1194,1220]" pageId="5" pageNumber="24">
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<taxonomicName id="54E87EF9501B766ADE88FB19FE76A7CA" box="[151,400,1194,1220]" class="Insecta" family="Mutoviidae" genus="Mutovia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="intercalaria" status="sp. nov.">
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<emphasis id="A19CD968501B766ADE88FB19FE76A7CA" bold="true" box="[151,400,1194,1220]" italics="true" pageId="5" pageNumber="24">Mutovia intercalaria</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BAAF6413501B766ADF87FB19FE3CA7CA" box="[408,474,1194,1220]" pageId="5" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9357057A501B766ADE88FB7EFEF4A7EB" blockId="5.[151,474,1194,1253]" box="[151,274,1229,1253]" pageId="5" pageNumber="24">
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(
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<figureCitation id="0BD319FF501B766ADE80FB7EFEEFA7EB" box="[159,265,1229,1253]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="2.[151,244,1945,1966]" captionStartId-1="3.[151,244,1856,1877]" captionStartId-2="4.[151,244,1607,1628]" captionStartId-3="6.[151,244,1662,1683]" captionStartId-4="7.[151,244,1769,1790]" captionStartId-5="8.[151,244,1841,1862]" captionTargetBox-0="[170,1417,410,1895]" captionTargetBox-1="[165,1422,184,1841]" captionTargetBox-2="[161,1423,412,1574]" captionTargetBox-3="[162,1424,299,1641]" captionTargetBox-4="[151,1436,339,1745]" captionTargetBox-5="[151,1436,411,1817]" captionTargetId-0="figure@2.[151,1436,369,1917]" captionTargetId-1="figure@3.[151,1436,177,1854]" captionTargetId-2="figure@4.[151,1436,395,1586]" captionTargetId-3="figure@6.[162,1424,299,1641]" captionTargetId-4="figure@7.[151,1436,339,1745]" captionTargetId-5="figure@8.[151,1436,411,1817]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 2. Forewings of Mutovia intercalaria sp. n. A, PIN 3840 / 52 (reverse, 19 mm); B, PIN 3840 / 53 (reverse, 19 mm); C, holotype PIN 3840 / 57 (reverse, length 16 mm; pale area is due to damage); Scale = 1 mm." captionText-1="FIGURE 3. Forewings of Mutovia intercalaria sp. n. A, PIN 3840 / 56 (reverse, 16 mm); B, PIN 3840 / 358 (reverse, 16 mm). C Hind wing tentatively attributed into male of Mutovia intercalaria PIN 3840 / 529 = 533 (length 17.5 mm) Scales = 1 mm." captionText-2="FIGURE 4. Explanatory drawing of forewings of Mutovia intercalaria sp. n. A, PIN 3840 / 518; B, PIN 3840 / 516; C, PIN 3840 / 520; D, PIN 3840 / 531; E, PIN 3840 / 356; F, PIN 3840 / 500; G, PIN 3840 / 501; H, PIN 3840 / 511; I, PIN 3840 / 532. Scales = 1 mm." captionText-3="FIGURE 5. Forewings of Mutovia intercalaria sp. n. A, holotype PIN 3840 / 57 (length 16 mm); B, PIN 3840 / 52 (19 mm); C Hind wing tentatively attributed into male of Mutovia intercalaria PIN 3840 / 529 = 533 (length 17.5 mm); D, PIN 3840 / 8 b, detail of venation; E, PIN 3840 / 66; F, PIN 3840 / 55 showing characteristic folding of posterior margin. Arrows show deformities. Scales = 1 mm." captionText-4="FIGURE 6. Forewings of Mutovia intercalaria sp. n. A, PIN 3840 / 56; B, PIN 3840 / 532; C, PIN 3840 / 53; D, PIN 3840 / 518; E, PIN 3840 / 54; F, PIN 3840 / 516; G, PIN 3840 / 356; H, PIN 3840 / 511; I, PIN 3840 / 500; J, PIN 3840 / 358. Scales = 10 mm." captionText-5="FIGURE 7. Forewings of Mutovia intercalaria sp. n. A, PIN 3840 / 523; B, PIN 3840 / 519; C, PIN 3840 / 360; D, PIN 3840 / 525; E, PIN 3840 / 531; F, PIN 3840 / 435; G, PIN 3840 / 520; H, PIN 3840 / 521. Scales = 10 mm." httpUri-0="https://zenodo.org/record/213152/files/figure.png" httpUri-1="https://zenodo.org/record/213153/files/figure.png" httpUri-2="https://zenodo.org/record/213154/files/figure.png" httpUri-3="https://zenodo.org/record/213155/files/figure.png" httpUri-4="https://zenodo.org/record/213156/files/figure.png" httpUri-5="https://zenodo.org/record/213157/files/figure.png" pageId="5" pageNumber="24">Figs. 2–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DBF256F1501B766ADE88FAA6FB88A6EF" pageId="5" pageNumber="24" type="materials_examined">
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<paragraph id="9357057A501B766ADE88FAA6FB88A6EF" blockId="5.[151,1436,1301,2009]" pageId="5" pageNumber="24">
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<emphasis id="A19CD968501B766ADE88FAA6FEACA620" bold="true" box="[151,330,1301,1326]" pageId="5" pageNumber="24">
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<typeStatus id="4C53BBD8501B766ADE88FAA6FF34A623" box="[151,210,1301,1325]" pageId="5" pageNumber="24">Type</typeStatus>
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Material.
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</emphasis>
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<typeStatus id="4C53BBD8501B766ADF4BFAA6FE27A623" box="[340,449,1301,1325]" pageId="5" pageNumber="24" type="holotype">Holotype</typeStatus>
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: Kitchuga Member, Poldars Formation (uppermost Severodvinian stage, Upper Permian),
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<taxonomicName id="54E87EF9501B766ADECFFA8AFDFEA65F" authority="Subzone" authorityName="Subzone" box="[208,536,1337,1361]" class="Amphibia" family="Chroniosuchidae" genus="Chroniosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anthracosauria" pageId="5" pageNumber="24" phylum="Chordata" rank="species" species="levis">
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<emphasis id="A19CD968501B766ADECFFA8AFE57A65F" box="[208,433,1337,1361]" italics="true" pageId="5" pageNumber="24">Chroniosaurus levis</emphasis>
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Subzone
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</taxonomicName>
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of the
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<emphasis id="A19CD968501B766ADC79FA8AFCA9A65F" box="[614,847,1337,1361]" italics="true" pageId="5" pageNumber="24">Proelginia permiana</emphasis>
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Zone. Isady, Vologda territory, northwestern
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<collectingCountry id="EBFF45EA501B766ADB55FA8AFA7EA65F" box="[1354,1432,1337,1361]" name="Russia" pageId="5" pageNumber="24">Russia</collectingCountry>
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.
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<collectionCode id="F5F99DBF501B766ADE88FAEDFF22A67B" box="[151,196,1374,1397]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="5" pageNumber="24">PIN</collectionCode>
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3840/57± (left forewing, male?).
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<typeStatus id="4C53BBD8501B766ADC50FAEDFD26A678" box="[591,704,1374,1398]" pageId="5" pageNumber="24" type="paratype">Paratypes</typeStatus>
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: The same as for
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<typeStatus id="4C53BBD8501B766ADD8CFAEEFC1FA67B" box="[915,1017,1373,1397]" pageId="5" pageNumber="24" type="holotype">holotype</typeStatus>
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:
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<collectionCode id="F5F99DBF501B766ADA18FAEDFBD2A67B" box="[1031,1076,1374,1397]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="5" pageNumber="24">PIN</collectionCode>
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3840/54±L, 55±R, 56±L, 66L, 67R, 358±L, 362R, 355, 356, 433, 434, 435, 501, 513, 519, 520, 524, 526, 528, 529=533 (hind wing), 531±, 532±, 534, 535±, 537, 539, 362, 540±, 656, 798 (?males); 8R, 52±L, 53±L, 355R, 356R, 359+-L, 360L, 500, 511, 516, 517, 518+-, 522, 523, 525 527, 539 (?females);
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<collectionCode id="F5F99DBF501B766ADCAFFA79FD3BA6EF" box="[688,733,1482,1505]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="5" pageNumber="24">PIN</collectionCode>
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3840/538, 521 (sex undetermined).
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</paragraph>
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</subSubSection>
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<subSubSection id="DBF256F1501B7669DED8FA5EFEE3A3DA" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24" type="description">
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<paragraph id="9357057A501B766ADED8FA5EFCFCA59B" blockId="5.[151,1436,1301,2009]" pageId="5" pageNumber="24">
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<emphasis id="A19CD968501B766ADED8FA5EFEBFA508" bold="true" box="[199,345,1517,1542]" pageId="5" pageNumber="24">Description.</emphasis>
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Forewing of medium size (
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<quantity id="5410A89F501B766ADC95FA5EFCE5A50B" box="[650,771,1517,1541]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="5" pageNumber="24" unit="mm" value="17.0" valueMax="19.0" valueMin="15.0">15–19 mm</quantity>
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x 5.1–6.5 mm). Sc (8–12 veins at margin), R1 (0–3 veins at margin) and RS (4–7 veins at margin) usually separated. M with 2–8 branches at margin, CuA with 6–8 veins. Clavus very short (less than one-fourth of wing length), CuP sometimes branched; anal veins simple or branched (up to 8–10 at margin). Colouration characteristically with pale main veins and dark intercalaries and reticulations, and with distinct coloured reticulation along clavus (
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<figureCitation id="0BD319FF501B766ADCABF9CEFD1EA59B" box="[692,760,1661,1685]" captionStart="FIGURE 2" captionStartId="2.[151,244,1945,1966]" captionTargetBox="[170,1417,410,1895]" captionTargetId="figure@2.[151,1436,369,1917]" captionTargetPageId="2" captionText="FIGURE 2. Forewings of Mutovia intercalaria sp. n. A, PIN 3840 / 52 (reverse, 19 mm); B, PIN 3840 / 53 (reverse, 19 mm); C, holotype PIN 3840 / 57 (reverse, length 16 mm; pale area is due to damage); Scale = 1 mm." httpUri="https://zenodo.org/record/213152/files/figure.png" pageId="5" pageNumber="24">Fig. 2</figureCitation>
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C).
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</paragraph>
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<paragraph id="9357057A501B766ADED8F912FB5FA40F" blockId="5.[151,1436,1301,2009]" pageId="5" pageNumber="24">
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<collectingCountry id="EBFF45EA501B766ADED8F912FF19A5B7" box="[199,255,1697,1721]" name="India" pageId="5" pageNumber="24">Hind</collectingCountry>
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wing (
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<figureCitation id="0BD319FF501B766ADF53F912FE7FA5B7" box="[332,409,1697,1721]" captionStart="FIGURE 3" captionStartId="3.[151,244,1856,1877]" captionTargetBox="[165,1422,184,1841]" captionTargetId="figure@3.[151,1436,177,1854]" captionTargetPageId="3" captionText="FIGURE 3. Forewings of Mutovia intercalaria sp. n. A, PIN 3840 / 56 (reverse, 16 mm); B, PIN 3840 / 358 (reverse, 16 mm). C Hind wing tentatively attributed into male of Mutovia intercalaria PIN 3840 / 529 = 533 (length 17.5 mm) Scales = 1 mm." httpUri="https://zenodo.org/record/213153/files/figure.png" pageId="5" pageNumber="24">Figs. 3</figureCitation>
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C, 5C) with long, simple, ascending branchelets, R1 with at least 5 veins at margin; RS simple or branched terminally. M expanded, with at least 5 veins at margin, with changed polarity from convex into concave near center of wing. CuA with about 5 veins (one secondarily branched), CuP simple.
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</paragraph>
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<paragraph id="9357057A501B766ADED8F8BEFE81A4D7" blockId="5.[151,1436,1301,2009]" pageId="5" pageNumber="24">
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<emphasis id="A19CD968501B766ADED8F8BEFDE0A428" bold="true" box="[199,518,1805,1830]" pageId="5" pageNumber="24">Character of preservation.</emphasis>
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Of forty-nine similar wing fossils, 48 are forewings, with 30 of the wings probably corresponding to males, 17 to females, and 1 wing indeterminable. One wing is a male hindwing. Nine of the identified wings are left wings and six are right wings. The apical posterior margin is often folded, suggesting that the wing was fragile at that area (similar to
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<taxonomicName id="54E87EF9501B766ADC8DF8C9FC03A49F" authority="Grimaldi, 2003" authorityName="Grimaldi" authorityYear="2003" box="[658,997,1913,1937]" class="Insecta" family="Santanmantidae" genus="Santanmantis" kingdom="Animalia" order="Mantodea" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
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<emphasis id="A19CD968501B766ADC8DF8C9FCCCA49F" box="[658,810,1914,1937]" italics="true" pageId="5" pageNumber="24">Santanmantis</emphasis>
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<bibRefCitation id="F779788B501B766ADD2BF8CAFC03A49F" author="Grimaldi" box="[820,997,1913,1937]" pageId="5" pageNumber="24" refString="Grimaldi, D. (2003) A revision of Cretaceous mantises and their relationships, including new taxa (Insecta: Dictyoptera: Mantodea). American Museum Novitates, 3412, 1 - 47." type="journal article" year="2003">Grimaldi, 2003</bibRefCitation>
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</taxonomicName>
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). The lengths of completely preserved specimens are
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<quantity id="5410A89F501B766ADF22F82EFE6AA4BB" box="[317,396,1949,1973]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="24" unit="mm" value="13.0">13 mm</quantity>
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(2 wings),
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<quantity id="5410A89F501B766ADC11F82EFDBBA4BB" box="[526,605,1949,1973]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="24" unit="mm" value="14.0">14 mm</quantity>
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(2),
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<quantity id="5410A89F501B766ADC8BF82EFD05A4BB" box="[660,739,1949,1973]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="24" unit="mm" value="16.0">16 mm</quantity>
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(7) and
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<quantity id="5410A89F501B766ADD5CF82EFC74A4BB" box="[835,914,1949,1973]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="24" unit="mm" value="19.0">19 mm</quantity>
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(3). All other forewings are approximately 16 or
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<quantity id="5410A89F501B766ADEAAF872FEC1A4D7" box="[181,295,1985,2009]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="5" pageNumber="24" unit="mm" value="20.0" valueMax="21.0" valueMin="19.0">19–21mm</quantity>
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long.
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</paragraph>
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<paragraph id="9357057A50187669DED8FF24FEE3A3DA" blockId="6.[151,1436,151,212]" pageId="6" pageNumber="25">
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<emphasis id="A19CD96850187669DED8FF24FE52A3BE" bold="true" box="[199,436,151,176]" pageId="6" pageNumber="25">Derivation of name.</emphasis>
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The species name refers to the intercalary veins of the wings, which are well marked and colourful.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |