treatments-xml/data/03/A6/63/03A663458202FFC01BA8F9BBFCE95C3C.xml

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<mods:title id="CF82F56282370A7AC61B1892E8167148">Stenopodidean shrimps (Crustacea: Decapoda) from New Caledonian waters</mods:title>
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<mods:namePart id="65D3233DA109DDF4CDFFE5E4D4703847">Goy, Joseph W.</mods:namePart>
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<mods:date id="3033D87D1A671E0A2C59C9E0B7511E5E">2015</mods:date>
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<paragraph id="8BB0D2538202FFDF1BA8F9BBFD905EF9" blockId="12.[151,588,1656,1716]" box="[151,588,1656,1682]" pageId="12" pageNumber="313">
<heading id="D0F8653F8202FFDF1BA8F9BBFD905EF9" bold="true" box="[151,588,1656,1682]" fontSize="11" level="1" pageId="12" pageNumber="313" reason="1">
<emphasis id="B97B0E418202FFDF1BA8F9BBFD905EF9" bold="true" box="[151,588,1656,1682]" pageId="12" pageNumber="313">
<taxonomicName id="4C0FA9D08202FFDF1BA8F9BBFDDE5EF9" ID-CoL="8GJWZ" box="[151,514,1656,1682]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="neocaledonensis" status="sp. nov.">
<emphasis id="B97B0E418202FFDF1BA8F9BBFDDE5EF9" bold="true" box="[151,514,1656,1682]" italics="true" pageId="12" pageNumber="313">Spongiocaris neocaledonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248B33A8202FFDF1935F9BBFD905EF9" box="[522,588,1656,1682]" pageId="12" pageNumber="313" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB0D2538202FFDF1BA8F958FEF25ED8" blockId="12.[151,588,1656,1716]" box="[151,302,1691,1716]" pageId="12" pageNumber="313">
(
<figureCitation id="1334CED68202FFDF1BA0F958FEFA5ED8" box="[159,294,1691,1716]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="14.[151,250,1520,1542]" captionStartId-1="15.[151,250,1880,1902]" captionStartId-2="16.[151,250,1651,1673]" captionTargetBox-0="[248,1347,205,1487]" captionTargetBox-1="[232,1345,492,1851]" captionTargetBox-2="[248,1332,196,1611]" captionTargetId-0="figure@14.[237,1349,193,1498]" captionTargetId-1="figure@15.[222,1365,481,1859]" captionTargetId-2="figure@16.[237,1349,193,1629]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." captionText-1="FIGURE 13. Spongiocaris neocaledonensis sp. nov. Female allotype, MNHN Na- 12000. A, carapace and abdomen, lateral view; B, carapace, dorsal view; C, fifth, sixth pleomeres, telson and uropods; D, thoracic sternites; E, egg; F; dactylus, propodus and carpus third pereiopod. Scale bars = 1.0 mm." captionText-2="FIGURE 14. Spongiocaris neocaledonensis sp. nov. A – E, I, J, male holotype, MNHN Na- 12000; F, male paratype, NMNH Na- 12003; G, male paratype, MNHN Na- 12005; H, male paratype, MNHN Na- 12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F – H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm." httpUri-0="https://zenodo.org/record/240472/files/figure.png" httpUri-1="https://zenodo.org/record/240473/files/figure.png" httpUri-2="https://zenodo.org/record/240474/files/figure.png" pageId="12" pageNumber="313">Figs. 12–14</figureCitation>
)
</paragraph>
<paragraph id="8BB0D2538202FFDF1BA8F927FC7B5E92" blockId="12.[151,935,1763,1785]" box="[151,935,1763,1785]" pageId="12" pageNumber="313">
<treatmentCitationGroup id="AB1FF57D8202FFDF1BA8F927FC7B5E92" box="[151,935,1763,1785]" pageId="12" pageNumber="313">
Non 
<taxonomicName id="4C0FA9D08202FFDF1BF6F920FE6C5E92" ID-CoL="8GJX4" box="[201,432,1763,1785]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="B97B0E418202FFDF1BF6F920FE6C5E92" box="[201,432,1763,1785]" italics="true" pageId="12" pageNumber="313">Spongiocaris yaldwyni</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0AAEF4428202FFDF1AF0F920FDB35E92" author="Goy" box="[463,623,1763,1785]" page="220" pageId="12" pageNumber="313" year="2010">
<bibRefCitation id="EF9EAFA28202FFDF1AF0F920FDE15E92" author="Goy" box="[463,573,1763,1785]" pageId="12" pageNumber="313" refString="Goy, J. W. (2010) Infraorder Stenopodidea Claus 1872. In: Schram, F. R., von Vaupel Klein, J. C., Forest, J. &amp; Charmantier- Daures, M. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, Decapoda, Volume 9 Part A Eucarida: Euphausiacea, Amphionidacea, and Decapoda (partim). Vol. 9 A, 215 - 265. Brill, Leiden, Netherlands." type="journal article" year="2010">Goy, 2010</bibRefCitation>
: 220
</treatmentCitation>
, figs. 65.3A, B, 
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, 65.12J.
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material
</emphasis>
. 
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: 
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<collectingCountry id="F31892C38202FFDF1ACDF8E6FD775F55" box="[498,683,1829,1854]" name="New Caledonia" pageId="12" pageNumber="313">New Caledonia</collectingCountry>
, 
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Ridge
</emphasis>
. 
<collectionCode id="ED1E4A968202FFDF184BF8E5FC645F55" box="[884,952,1830,1854]" pageId="12" pageNumber="313">SMIB</collectionCode>
4, stn DW58, 
<geoCoordinate id="EE3BB4948202FFDF1F5EF8E6FB095F55" box="[1121,1237,1829,1854]" direction="south" orientation="latitude" pageId="12" pageNumber="313" precision="92" value="-22.996666">22°59.8’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948202FFDF1FDDF8E6FAB95F55" box="[1250,1381,1829,1854]" direction="east" orientation="longitude" pageId="12" pageNumber="313" precision="92" value="167.40334">167°24.2’E</geoCoordinate>
, 
<quantity id="4CF77FB68202FFDF1E4DF8E6FF735F0A" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="12" pageNumber="313" unit="m" value="560.0">560 m</quantity>
, 
<date id="FFB1F4938202FFDF1B82F889FEF35F09" box="[189,303,1866,1890]" pageId="12" pageNumber="313" value="1989-03-09">9.III.1989</date>
, male, 
<specimenCount id="9D0919DA8202FFDF1ABBF889FE3B5F09" box="[388,487,1865,1890]" pageId="12" pageNumber="313" type="female">1 female</specimenCount>
(MNHN-NA 12000). ALLOTYPE: 
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<collectingCountry id="F31892C38202FFDF18B4F88AFB985F09" box="[907,1092,1865,1890]" name="New Caledonia" pageId="12" pageNumber="313">New Caledonia</collectingCountry>
, 
<collectingCountry id="F31892C38202FFDF1F6DF88AFB735F09" box="[1106,1199,1865,1890]" name="Norfolk Island" pageId="12" pageNumber="313">Norfolk</collectingCountry>
Ridge
</emphasis>
. 
<collectionCode id="ED1E4A968202FFDF1E30F889FA8F5F09" box="[1295,1363,1866,1890]" pageId="12" pageNumber="313">SMIB</collectionCode>
4, stn DW58, 
<geoCoordinate id="EE3BB4948202FFDF1BD0F8AEFEBF5FED" box="[239,355,1901,1926]" direction="south" orientation="latitude" pageId="12" pageNumber="313" precision="92" value="-22.996666">22°59.8’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948202FFDF1A52F8AEFE2D5FED" box="[365,497,1901,1926]" direction="east" orientation="longitude" pageId="12" pageNumber="313" precision="92" value="167.40334">167°24.2’E</geoCoordinate>
, 
<quantity id="4CF77FB68202FFDF1AC3F8AEFD985FEE" box="[508,580,1901,1926]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="12" pageNumber="313" unit="m" value="560.0">560 m</quantity>
, 
<date id="FFB1F4938202FFDF196FF8ADFD1E5FED" box="[592,706,1902,1926]" pageId="12" pageNumber="313" value="1989-03-09">9.III.1989</date>
, female (MNHN-NA 12000). 
<typeStatus id="54B46CF18202FFDF1F2BF8ADFB6F5FED" box="[1044,1203,1902,1926]" pageId="12" pageNumber="313" type="paratype">PARATYPES</typeStatus>
: 
<collectionCode id="ED1E4A968202FFDF1F83F8ADFADC5FED" box="[1212,1280,1902,1926]" pageId="12" pageNumber="313">SMIB</collectionCode>
4, stn DW61, 
<geoCoordinate id="EE3BB4948202FFDF1BA8F852FED65FC1" box="[151,266,1937,1962]" direction="south" orientation="latitude" pageId="12" pageNumber="313" precision="92" value="-22.998333">22°59.9’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948202FFDF1A21F852FE7D5FC1" box="[286,417,1937,1962]" direction="east" orientation="longitude" pageId="12" pageNumber="313" precision="92" value="167.38">167°22.8’E</geoCoordinate>
, 
<quantity id="4CF77FB68202FFDF1A8AF852FDDA5FC2" box="[437,518,1937,1962]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="313" unit="m" value="550.0">550 m</quantity>
, 
<date id="FFB1F4938202FFDF1925F851FD465FC1" box="[538,666,1938,1962]" pageId="12" pageNumber="313" value="1989-03-10">10.III.1989</date>
, 
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(1 ov.) (MNHN-NA 12001).— 
<collectionCode id="ED1E4A968202FFDF1F9EF851FB345FC1" box="[1185,1256,1938,1962]" pageId="12" pageNumber="313">SMIB</collectionCode>
4, stn DW63, 
<geoCoordinate id="EE3BB4948202FFDF1BA8F876FED65FA5" box="[151,266,1973,1998]" direction="south" orientation="latitude" pageId="12" pageNumber="313" precision="92" value="-22.978333">22°58.7’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948202FFDF1A27F876FE475FA5" box="[280,411,1973,1998]" direction="east" orientation="longitude" pageId="12" pageNumber="313" precision="92" value="167.35167">167°21.1’E</geoCoordinate>
, 
<quantity id="4CF77FB68202FFDF1A96F876FE285FA6" box="[425,500,1973,1998]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="313" unit="m" value="520.0">520 m</quantity>
, 
<date id="FFB1F4938202FFDF193DF875FD5E5FA5" box="[514,642,1974,1998]" pageId="12" pageNumber="313" value="1989-03-10">10.III.1989</date>
, 
<specimenCount id="9D0919DA8202FFDF19AFF875FD2F5FA5" box="[656,755,1973,1998]" pageId="12" pageNumber="313" type="female">1 female</specimenCount>
ov. (MNHN-NA 12004).— 
<collectionCode id="ED1E4A968202FFDF1F0CF875FBAB5FA5" box="[1075,1143,1974,1998]" pageId="12" pageNumber="313">SMIB</collectionCode>
4, stn DW64, 
<geoCoordinate id="EE3BB4948202FFDF1E1BF876FA4B5FA5" box="[1316,1431,1973,1998]" direction="south" orientation="latitude" pageId="12" pageNumber="313" precision="92" value="-22.921667">22°55.3’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948202FFDF1BA8F81AFEC55F99" box="[151,281,2009,2034]" direction="east" orientation="longitude" pageId="12" pageNumber="313" precision="92" value="167.27333">167°16.4’E</geoCoordinate>
, 
<quantity id="4CF77FB68202FFDF1A19F81AFEAD5F9A" box="[294,369,2009,2034]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="12" pageNumber="313" unit="m" value="460.0">460 m</quantity>
, 
<date id="FFB1F4938202FFDF1A42F819FE225F99" box="[381,510,2010,2034]" pageId="12" pageNumber="313" value="1989-03-10">10.III.1989</date>
, 
<specimenCount id="9D0919DA8202FFDF1934F819FD865F99" box="[523,602,2009,2034]" pageId="12" pageNumber="313" type="male">1 male</specimenCount>
, (MNHN-NA 12005).— 
<collectingCountry id="F31892C38202FFDF184EF81AFBFB5F99" box="[881,1063,2009,2034]" name="New Caledonia" pageId="12" pageNumber="313">
<emphasis id="B97B0E418202FFDF184EF81AFBFB5F99" bold="true" box="[881,1063,2009,2034]" pageId="12" pageNumber="313">New Caledonia</emphasis>
</collectingCountry>
. BIOCAL, stn DW 46, 
<geoCoordinate id="EE3BB4948202FFDF1E06F81AFA4B5F99" box="[1337,1431,2009,2034]" direction="south" orientation="latitude" pageId="12" pageNumber="313" precision="925" value="-22.883333">22°53’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948203FFDE1BA8FF54FED858DB" box="[151,260,151,176]" direction="east" orientation="longitude" pageId="13" pageNumber="314" precision="925" value="167.28334">167°17’E</geoCoordinate>
, 
<quantity id="4CF77FB68203FFDE1A2BFF54FE4658C4" box="[276,410,151,176]" metricMagnitude="2" metricUnit="m" metricValue="5.9" metricValueMax="6.1" metricValueMin="5.7" pageId="13" pageNumber="314" unit="m" value="590.0" valueMax="610.0" valueMin="570.0">570–610 m</quantity>
, 
<date id="FFB1F4938203FFDE1A95FF54FDE358DB" box="[426,575,151,176]" pageId="13" pageNumber="314" value="1985-08-30">30.VIII.1985</date>
, 
<specimenCount id="9D0919DA8203FFDE1971FF5BFD7C58DB" box="[590,672,151,176]" pageId="13" pageNumber="314" type="male">1 male</specimenCount>
, (MNHN-NA 12007).—MUSORSTOM 4, stn CP215, 
<geoCoordinate id="EE3BB4948203FFDE1E1BFF54FA4458DB" box="[1316,1432,151,176]" direction="south" orientation="latitude" pageId="13" pageNumber="314" precision="92" value="-22.928333">22°55.7’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948203FFDE1BA8FF7FFED858BE" box="[151,260,188,213]" direction="east" orientation="longitude" pageId="13" pageNumber="314" precision="925" value="167.28334">167°17’E</geoCoordinate>
, 
<quantity id="4CF77FB68203FFDE1A2EFF7FFE4E58BF" box="[273,402,188,213]" metricMagnitude="2" metricUnit="m" metricValue="5.025" metricValueMax="5.2" metricValueMin="4.85" pageId="13" pageNumber="314" unit="m" value="502.5" valueMax="520.0" valueMin="485.0">485–520 m</quantity>
, 
<date id="FFB1F4938203FFDE1AA1FF7FFDFD58BE" box="[414,545,188,213]" pageId="13" pageNumber="314" value="1985-09-28">28.IX.1985</date>
, 
<specimenCount id="9D0919DA8203FFDE1912FF7FFD4558BE" box="[557,665,188,213]" pageId="13" pageNumber="314" type="female">3 females</specimenCount>
(2 ov.) in 
<emphasis id="B97B0E418203FFDE1831FF7FFBDE58BF" box="[782,1026,188,212]" italics="true" pageId="13" pageNumber="314">Regardella okinseana</emphasis>
, (
<collectionCode id="ED1E4A968203FFDE1F27FF7FFBB758BF" LSID="urn:lsid:biocol.org:col:34871" box="[1048,1131,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
256952).—BIOGEOCAL, stn CP290, 
<geoCoordinate id="EE3BB4948203FFDE1A26FF1CFE475893" box="[281,411,223,248]" direction="south" orientation="latitude" pageId="13" pageNumber="314" precision="9" value="-20.615166">20°36.91’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948203FFDE1A98FF1CFDE45893" box="[423,568,223,248]" direction="east" orientation="longitude" pageId="13" pageNumber="314" precision="9" value="167.05566">167°03.34’E</geoCoordinate>
, 970– 
<quantity id="4CF77FB68203FFDE1943FF1CFD19589C" box="[636,709,223,248]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="13" pageNumber="314" unit="m" value="760.0">760 m</quantity>
, 
<date id="FFB1F4938203FFDE19EEFF23FC935893" box="[721,847,224,248]" pageId="13" pageNumber="314" value="1987-04-27">27.IV.1987</date>
, 
<specimenCount id="9D0919DA8203FFDE1864FF23FC635893" box="[859,959,223,248]" pageId="13" pageNumber="314" type="female">1 female</specimenCount>
, (MNHN-NA12002). — 
<emphasis id="B97B0E418203FFDE1FE3FF1CFA485893" bold="true" box="[1244,1428,223,248]" pageId="13" pageNumber="314">Loyalty Islands</emphasis>
. W. Lifou, N. Baie du Santal, CALSUB, Dive 7, 
<geoCoordinate id="EE3BB4948203FFDE1988FEC7FCC95977" box="[695,789,260,284]" direction="south" orientation="latitude" pageId="13" pageNumber="314" precision="925" value="-22.8">22°48’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB4948203FFDE181DFEC7FC4C5976" box="[802,912,260,285]" direction="east" orientation="longitude" pageId="13" pageNumber="314" precision="925" value="167.08333">167°05’E</geoCoordinate>
, 966– 
<quantity id="4CF77FB68203FFDE18EAFEC7FBC25977" box="[981,1054,260,285]" metricMagnitude="2" metricUnit="m" metricValue="8.45" pageId="13" pageNumber="314" unit="m" value="845.0">845 m</quantity>
, 
<date id="FFB1F4938203FFDE1F15FEC7FB7D5976" box="[1066,1185,260,285]" pageId="13" pageNumber="314" value="1989-02-25">25.II.1989</date>
, 
<specimenCount id="9D0919DA8203FFDE1F92FEC7FB245976" box="[1197,1272,260,285]" pageId="13" pageNumber="314" type="male">1 male</specimenCount>
on gorgonian, (MNHN-NA 12003).
</paragraph>
</subSubSection>
<caption id="DF7082DB8203FFDE1BA8FAC5FB335D77" box="[151,1263,1286,1308]" httpUri="https://zenodo.org/record/240471/files/figure.png" pageId="13" pageNumber="314" targetBox="[151,1436,373,1264]" targetPageId="13">
<paragraph id="8BB0D2538203FFDE1BA8FAC5FB335D77" blockId="13.[151,1263,1286,1308]" box="[151,1263,1286,1308]" pageId="13" pageNumber="314">
<emphasis id="B97B0E418203FFDE1BA8FAC5FEC45D70" bold="true" box="[151,280,1286,1308]" pageId="13" pageNumber="314">FIGURE 11</emphasis>
. 
<taxonomicName id="4C0FA9D08203FFDE1A1BFAC5FD3C5D77" authority="Baba, 1979" authorityName="Baba" authorityYear="1979" box="[292,736,1286,1308]" class="Malacostraca" family="Spongicolidae" genus="Spongicoloides" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="novaezelandiae">
<emphasis id="B97B0E418203FFDE1A1BFAC5FDBE5D77" box="[292,610,1286,1308]" italics="true" pageId="13" pageNumber="314">Spongicoloides novaezelandiae</emphasis>
Baba, 1979
</taxonomicName>
. Female, MNHN Na-11997, photo by M. Segonzac.
</paragraph>
</caption>
<subSubSection id="C31581D88203FFDE1BF8FA8BFAEF5E85" pageId="13" pageNumber="314" type="diagnosis">
<paragraph id="8BB0D2538203FFDE1BF8FA8BFAEF5E85" blockId="13.[151,1437,1352,2025]" pageId="13" pageNumber="314">
<emphasis id="B97B0E418203FFDE1BF8FA8BFEE65D0A" bold="true" box="[199,314,1352,1377]" pageId="13" pageNumber="314">Diagnosis</emphasis>
. Small commensal spongicolid shrimp associated with sponges or gorgonians, with slightly depressed body bearing few spines; carapace with distinct cervical groove with 4–10 small spines; small branchiostegal spine present; small supraorbital, antennal, and hepatic spines sometimes present; rostrum moderately long reaching to middle of last antennular peduncle segment with 4–6 dorsal, 0–1 ventral, 0–2 lateral spines; pigment of cornea restricted to submedial ring in starburst pattern; unarmed epistome; endopod of maxillule unarmed; setiferous organ of third maxilliped covering distal half of propodal ventral margin; first pereiopod with well developed setiferous organ; males with spines on meri, propodi of third pereiopods, single forward curving ventral medial spine on fifth abdominal pleomere; females with entire third pereiopods glabrous, lacking ventral medial spine on fifth abdominal somite; third pereiopod ventral propodal margin distally with dense fringe of setal brushes; fourth and fifth pereiopods with carpi distinctly 3-segmented; first to fifth pereiopods lack epipods but bear cicatrices; posterior margin of fifth abdominal pleomere ending in 4 or 5 spines, that of sixth pleomere in 3–9 spines; few dorsal spines on sixth abdominal pleomere; uropodal endopodite with 2 terminal dorsal hairs.
</paragraph>
</subSubSection>
<subSubSection id="C31581D88203FFC11BF8F93BFD715C00" lastPageId="18" lastPageNumber="319" pageId="13" pageNumber="314" type="description">
<paragraph id="8BB0D2538203FFDE1BF8F93BFD225F32" blockId="13.[151,1437,1352,2025]" pageId="13" pageNumber="314">
<emphasis id="B97B0E418203FFDE1BF8F93BFE8D5F7A" bold="true" box="[199,337,1784,1809]" pageId="13" pageNumber="314">Description</emphasis>
. (
<typeStatus id="54B46CF18203FFDE1A57F93BFE085F7A" box="[360,468,1784,1809]" pageId="13" pageNumber="314" type="holotype">Holotype</typeStatus>
, male MNHN-Na-12000) Rostrum (
<figureCitation id="1334CED68203FFDE184DF93BFC125F7A" box="[882,974,1784,1809]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="13" pageNumber="314">Figs. 12</figureCitation>
A, B) broad at base, slightly compressed anteriorly; moderately long, reaching past middle of last antennular peduncle segment; dorsal margin with 5 spines followed by blunt tubercle; no ventral or lateral spines.
</paragraph>
<paragraph id="8BB0D2538203FFDE1BF8F8A6FDFB5F82" blockId="13.[151,1437,1352,2025]" pageId="13" pageNumber="314">
Carapace (
<figureCitation id="1334CED68203FFDE1A00F8A6FE4F5F15" box="[319,403,1893,1918]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="13" pageNumber="314">Figs.12</figureCitation>
A, B) with distinct cervical groove bearing 10 small spines. Small spine at rostral base with 2 smaller spines behind located dorsomesially. Small antennal and branchiostegal spines, large spine at pterygosmial angle. Group of 3 small anterolateral spines present. Ventrolateral angle rounded, posterolateral angle of branchiostegite broadly rounded.
</paragraph>
<caption id="DF7082DB8200FFDD1BA8FA33FB485E28" httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="14" pageNumber="315" targetBox="[248,1347,205,1487]" targetPageId="14">
<paragraph id="8BB0D2538200FFDD1BA8FA33FB485E28" blockId="14.[151,1436,1520,1603]" pageId="14" pageNumber="315">
<emphasis id="B97B0E418200FFDD1BA8FA33FEC75E6E" bold="true" box="[151,283,1520,1542]" pageId="14" pageNumber="315">FIGURE 12</emphasis>
. 
<taxonomicName id="4C0FA9D08200FFDD1A16FA32FD825E6D" box="[297,606,1520,1542]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="neocaledonensis" status="sp. nov.">
<emphasis id="B97B0E418200FFDD1A16FA32FD825E6D" box="[297,606,1520,1542]" italics="true" pageId="14" pageNumber="315">Spongiocaris neocaledonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248B33A8200FFDD1958FA33FD655E6E" box="[615,697,1520,1541]" pageId="14" pageNumber="315" rank="species">
<emphasis id="B97B0E418200FFDD1958FA33FD655E6E" bold="true" box="[615,697,1520,1541]" pageId="14" pageNumber="315">sp. nov.</emphasis>
</taxonomicNameLabel>
Male holotype, MNHN Na-12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB0D2538200FFDD1BF8F9A9FB145E84" blockId="14.[151,1436,1642,1991]" pageId="14" pageNumber="315">
Abdominal pleomeres (
<figureCitation id="1334CED68200FFDD1AEBF9A9FDFA5EE8" box="[468,550,1642,1667]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="14" pageNumber="315">Fig. 12</figureCitation>
A) generally glabrous, lacking carinae. First to fifth pleomeres with rounded pleural margins; sixth pleomere’s pleuron ending in acute point. Posterior margin of fifth pleomere ending in 4 spines, that of sixth in 3 spines. Dorsally, sixth pleomere bears row of 3 medial spines, 2 large submedial and 2 smaller lateral spines. Only fifth pleomere with strong median ventral spine, directed anteriorly.
</paragraph>
<paragraph id="8BB0D2538200FFDD1BF8F938FBF35F5C" blockId="14.[151,1436,1642,1991]" pageId="14" pageNumber="315">
Eyes (
<figureCitation id="1334CED68200FFDD1A2DF939FEAF5F78" box="[274,371,1786,1811]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="14" pageNumber="315">Figs. 12</figureCitation>
A, B) well developed, of moderate size; cornea globular with black pigment restricted to submedial ring arranged in starburst pattern. No spinules on ophthalmic peduncle.
</paragraph>
<paragraph id="8BB0D2538200FFDD1BF8F881FCEC5FAC" blockId="14.[151,1436,1642,1991]" pageId="14" pageNumber="315">
Telson (
<figureCitation id="1334CED68200FFDD1A1DF881FEA95F30" box="[290,373,1858,1883]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="14" pageNumber="315">Fig. 12</figureCitation>
C) long, broad subtriangular, with median groove flanked by 2 longitudinal carinae. Telsonal base with pairs of small outer and medial spines. Dorsal surface of carinae with left side bearing 5 spines, right bearing 4. Few setae within median groove and outside carinae. Lateral margins of telson with 7 left, 8 right acute teeth. Posterior margin rounded, setose with 10 small teeth.
</paragraph>
<paragraph id="8BB0D2538201FFDC1BF8FF54FEE05954" blockId="15.[151,1437,151,429]" pageId="15" pageNumber="316">
Uropods (
<figureCitation id="1334CED68201FFDC1A02FF54FE4958DB" box="[317,405,151,176]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="15" pageNumber="316">Fig. 12</figureCitation>
C) well developed, about as long as telson. Basal segment strong with 2 small teeth posterolaterally. Exopodite broader than ovate endopodite; outer margin of left exopodite with 10 teeth, that of right with 11 teeth; dorsal surface of exopodite with 2 strong ridges. Margins of endopodite unarmed, dorsally with 1 strong ridge, terminating in 2 curved dorsal hairs. Unarmed margins of exopoditess, endopodites with long plumose setae.
</paragraph>
<paragraph id="8BB0D2538201FFDC1BF8FE8FFB9259C6" blockId="15.[151,1437,151,429]" pageId="15" pageNumber="316">
Epistome (
<figureCitation id="1334CED68201FFDC1A75FE8FFE78590F" box="[330,420,332,357]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="15" pageNumber="316">Fig. 12</figureCitation>
E) unarmed, rounded anteriorly, with rounded lateral extensions. Labrum normally developed, posterior margin acutely rounded. Paragnath slightly bilobed. Thoracic sternites (
<figureCitation id="1334CED68201FFDC1F9EFEACFB2C59E3" box="[1185,1264,367,392]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="15" pageNumber="316">Fig. 12</figureCitation>
D) narrow with 2 submedian spinules on segments 4, 6, 7, 8 and 1 submedian spinules on segment 5.
</paragraph>
<caption id="DF7082DB8201FFDC1BA8F89BFD315FC0" httpUri="https://zenodo.org/record/240473/files/figure.png" pageId="15" pageNumber="316" targetBox="[232,1345,492,1851]" targetPageId="15">
<paragraph id="8BB0D2538201FFDC1BA8F89BFD315FC0" blockId="15.[151,1436,1880,1963]" pageId="15" pageNumber="316">
<emphasis id="B97B0E418201FFDC1BA8F89BFEC75F06" bold="true" box="[151,283,1880,1902]" pageId="15" pageNumber="316">FIGURE 13</emphasis>
. 
<taxonomicName id="4C0FA9D08201FFDC1A15F89AFD835F05" box="[298,607,1880,1902]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="neocaledonensis" status="sp. nov.">
<emphasis id="B97B0E418201FFDC1A15F89AFD835F05" box="[298,607,1880,1902]" italics="true" pageId="15" pageNumber="316">Spongiocaris neocaledonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248B33A8201FFDC1957F89BFD675F06" box="[616,699,1880,1901]" pageId="15" pageNumber="316" rank="species">
<emphasis id="B97B0E418201FFDC1957F89BFD675F06" bold="true" box="[616,699,1880,1901]" pageId="15" pageNumber="316">sp. nov.</emphasis>
</taxonomicNameLabel>
Female allotype, MNHN Na-12000. A, carapace and abdomen, lateral view; B, carapace, dorsal view; C, fifth, sixth pleomeres, telson and uropods; D, thoracic sternites; E, egg; F; dactylus, propodus and carpus third pereiopod. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF7082DB821EFFC31BA8F9B0FE3C5E8F" httpUri="https://zenodo.org/record/240474/files/figure.png" pageId="16" pageNumber="317" targetBox="[248,1332,196,1611]" targetPageId="16">
<paragraph id="8BB0D253821EFFC31BA8F9B0FE3C5E8F" blockId="16.[151,1436,1651,1764]" pageId="16" pageNumber="317">
<emphasis id="B97B0E41821EFFC31BA8F9B0FEC65EE3" bold="true" box="[151,282,1651,1673]" pageId="16" pageNumber="317">FIGURE 14</emphasis>
. 
<taxonomicName id="4C0FA9D0821EFFC31A16F9B0FD815EE2" box="[297,605,1651,1673]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="neocaledonensis" status="sp. nov.">
<emphasis id="B97B0E41821EFFC31A16F9B0FD815EE2" box="[297,605,1651,1673]" italics="true" pageId="16" pageNumber="317">Spongiocaris neocaledonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248B33A821EFFC31959F9B0FD645EE3" box="[614,696,1651,1672]" pageId="16" pageNumber="317" rank="species">
<emphasis id="B97B0E41821EFFC31959F9B0FD645EE3" bold="true" box="[614,696,1651,1672]" pageId="16" pageNumber="317">sp. nov.</emphasis>
</taxonomicNameLabel>
A–E, I, J, male holotype, MNHN Na-12000; F, male paratype, NMNH Na-12003; G, male paratype, MNHN Na-12005; H, male paratype, MNHN Na-12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F–H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB0D253821EFFC31BF8F8D2FECC5FD1" blockId="16.[151,1436,1809,1978]" pageId="16" pageNumber="317">
Antennular peduncle (
<figureCitation id="1334CED6821EFFC31AFAF8D2FDC55F41" box="[453,537,1809,1834]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="16" pageNumber="317">Fig. 12</figureCitation>
B) moderately long, extending to middle of scaphocerite. Basal segment more than 2.0 times length of second segment, third same length as second. Outer margin with small rounded stylocerite, few setae; 2 small spines on distodorsal inner margin. Upper and lower flagella short, but well developed; upper flagellum unarmed, lower flagellum consisting of 22 segments with numerous aesthetascs along proximal to midlength.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8FF5BFDDA590F" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
Antenna (
<figureCitation id="1334CED6821FFFC21A08FF54FE4858DB" box="[311,404,151,176]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="17" pageNumber="318">Figs. 12</figureCitation>
A, B) with stout basicerite with 2 strong distodorsal spines on outer margin, medial carina with rounded distal extremity, sharp hooked spine ventrolaterally. Carpocerite unarmed, short not exceeding first antennular peduncle segment. Antennal flagellum long, slender, unarmed, extending approximately to tip of telson. Scaphocerite broad, slightly quadrangular, dorsally bearing 2 strong longitudinal carinae; outer margin slightly concave bearing 3 small spines proximally, 5 small teeth distally, larger distal most tooth; inner margin convex, fringed with long plumose setae.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8FEACFE11599E" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
Mandible (
<figureCitation id="1334CED6821FFFC21A7AFEACFE7F59E3" box="[325,419,367,392]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="17" pageNumber="318">Figs. 12</figureCitation>
F, G) robust, with short, fused molar and incisor processes. Molar surface with 3 small teeth; incisor bearing 2 stout teeth distally followed by 6 small teeth proximally. Palp well developed, 3-segmented. Proximal segment without setae; middle segment setose mesially, on distal outer margin; distal segment densely setose laterally and distally.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8FE3CFE835A0B" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
Maxillule (
<figureCitation id="1334CED6821FFFC21A7BFE3CFE4A5A73" box="[324,406,511,536]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="17" pageNumber="318">Fig. 12</figureCitation>
H) with slender, unarmed, undivided endopodite. Proximal endite broad, roundly truncated distally with numerous simple and compound spinose setae. Distal endite much narrower, rounded, with setae along its margins.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8FDAFFEAE5A9B" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
Maxilla (
<figureCitation id="1334CED6821FFFC21A0EFDAFFE595AEF" box="[305,389,620,645]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="17" pageNumber="318">
Fig. 
<date id="FFB1F493821FFFC21A56FDAFFE595AEF" box="[361,389,620,644]" pageId="17" pageNumber="318" value="2012-01">12</date>
</figureCitation>
I) with numerous plumose setae on both lobes of coxal and basal endites, proximal coxal endite broadest. Endopodite long, but robust, not exceeding anterior margin of scaphgnathite, with 16 plumose setae along margins. Scaphognathite long, anterior portion broader than posterior portion, with numerous plumose setae along margin.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8FD3FFBCC5B87" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
First maxilliped (
<figureCitation id="1334CED6821FFFC21AACFD3FFE3B5B7F" box="[403,487,764,789]" captionStart="FIGURE 14" captionStartId="16.[151,250,1651,1673]" captionTargetBox="[248,1332,196,1611]" captionTargetId="figure@16.[237,1349,193,1629]" captionTargetPageId="16" captionText="FIGURE 14. Spongiocaris neocaledonensis sp. nov. A – E, I, J, male holotype, MNHN Na- 12000; F, male paratype, NMNH Na- 12003; G, male paratype, MNHN Na- 12005; H, male paratype, MNHN Na- 12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F – H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240474/files/figure.png" pageId="17" pageNumber="318">Fig. 14</figureCitation>
A) with 2-segmented endopodite. Proximal segment 2.0 times longer than distal segment with 10 long plumose along outer margin, 3 simple setae on inner margin. Distal segment tapering with 10 long plumose setae on outer margin, stout terminal spine, long plumose seta on inner margin. Basipodite large, rounded anteriorly and posteriorly, slightly concave near middle, with outer margin bearing dense fringe of setae. Coxopodite bilobed with numerous seate. Exopodite well developed, flagellum arising from unsegmented peduncle with 5 proximal and 20 distodorsal long plumose setae. Moderately large epipod with slender proximal and distal lobes approximately equal in length, small arthrobranch also present.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8FC34FD6A5C83" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
Second maxilliped (
<figureCitation id="1334CED6821FFFC21A99FC34FE2A5C7B" box="[422,502,1015,1040]" captionStart="FIGURE 14" captionStartId="16.[151,250,1651,1673]" captionTargetBox="[248,1332,196,1611]" captionTargetId="figure@16.[237,1349,193,1629]" captionTargetPageId="16" captionText="FIGURE 14. Spongiocaris neocaledonensis sp. nov. A – E, I, J, male holotype, MNHN Na- 12000; F, male paratype, NMNH Na- 12003; G, male paratype, MNHN Na- 12005; H, male paratype, MNHN Na- 12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F – H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240474/files/figure.png" pageId="17" pageNumber="318">Fig. 14</figureCitation>
B) with 5-segmented endopodite. Dactylus suboval, slightly longer than broad, with dense fringe of setae along distodorsal margin. Propodus equal to dactylar length, densely setose on dorsal margin, ventral margin unarmed, notched proximally, rounded distally. Carpus triangular, narrowed proximally, equal to propodal length, densely setose on outer margin. Merus long, slightly concave medially, about 4.0 times carpal length with fringe of short setae on inner margin, 4 setae on outer margin. Ischium and basis not fused, both with few setae. Coxa with small epipod, arthrobranch and lateral podobranch. Exopodite long, slender, undivided with 3 proximal and 20 distodorsal long plumose setae.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8FB37FC5A5DAB" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
Third maxilliped (
<figureCitation id="1334CED6821FFFC21AADFB37FE3D5D67" box="[402,481,1268,1293]" captionStart="FIGURE 14" captionStartId="16.[151,250,1651,1673]" captionTargetBox="[248,1332,196,1611]" captionTargetId="figure@16.[237,1349,193,1629]" captionTargetPageId="16" captionText="FIGURE 14. Spongiocaris neocaledonensis sp. nov. A – E, I, J, male holotype, MNHN Na- 12000; F, male paratype, NMNH Na- 12003; G, male paratype, MNHN Na- 12005; H, male paratype, MNHN Na- 12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F – H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240474/files/figure.png" pageId="17" pageNumber="318">Fig. 14</figureCitation>
C) with well developed, 5-segmnted endopodite. Dactylus slender, tapering, 2.0 times longer than broad, dorsal margin with 3 simple setae, numerous long simple setae mesially and on inner margin. Propodus 2.0 times dactylar length, 4 outer setae, entire inner margin with dense fringe of setae on distal setiferous organ. Carpus about equal propodal length, merus 2.0 times carpal length, ischium equal meral length; all with few setae on outer margins, densely setose in double row on inner margins. Basis without exopodite, sparsely setose on inner margin. Coxa bearing small epipod, two small arthrobranchs.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8FA0FFD5B5ED7" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
First pereiopod (
<figureCitation id="1334CED6821FFFC21AB4FA0FFE3D5D8F" box="[395,481,1484,1509]" captionStart="FIGURE 14" captionStartId="16.[151,250,1651,1673]" captionTargetBox="[248,1332,196,1611]" captionTargetId="figure@16.[237,1349,193,1629]" captionTargetPageId="16" captionText="FIGURE 14. Spongiocaris neocaledonensis sp. nov. A – E, I, J, male holotype, MNHN Na- 12000; F, male paratype, NMNH Na- 12003; G, male paratype, MNHN Na- 12005; H, male paratype, MNHN Na- 12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F – H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240474/files/figure.png" pageId="17" pageNumber="318">Fig. 14</figureCitation>
D) small, slender, when stretched reaching past scaphocerite, segments with few simple setae on margins. Dactylus less than 0.5 propodal length. Fingers slightly compressed, with hooked highly chitinized tips, overlapping when closed. Cutting edges rather indistinct with propodus and dactylus bearing chitinous ridge along inner margins. Fingers and distodorsal extremity of palm with small tufts of long setae arranged like brushes. Distoventral carpus and proximoventral propodus with setiferous organ. Carpus longest segment, over 2.0 times propodal length. Merus almost equal carpal length, ischium equal to propodal length. Basis and coxa short, unarmed, no epipod present.
</paragraph>
<paragraph id="8BB0D253821FFFC21BF8F904FD175F1B" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="318">
Second pereiopod (
<figureCitation id="1334CED6821FFFC21A95F904FDDC5E8B" box="[426,512,1735,1760]" captionStart="FIGURE 14" captionStartId="16.[151,250,1651,1673]" captionTargetBox="[248,1332,196,1611]" captionTargetId="figure@16.[237,1349,193,1629]" captionTargetPageId="16" captionText="FIGURE 14. Spongiocaris neocaledonensis sp. nov. A – E, I, J, male holotype, MNHN Na- 12000; F, male paratype, NMNH Na- 12003; G, male paratype, MNHN Na- 12005; H, male paratype, MNHN Na- 12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F – H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240474/files/figure.png" pageId="17" pageNumber="318">Fig. 14</figureCitation>
E) similar to first, but longer, stronger, segments with few simple setae along margins. Fingers with hooked, heavily chintinized tips, overlapping when closed. Cutting edges entire, chitinized ridges along inner dactylar and prpodal margins. Fingers and distdorsal extremity of palm with small tuft of long setae. Carpus longest segment, about 1.5 times propodal length, merus 0.8 carpal length, ischium 0.3 meral length. Basis and coxa short, unarmed, no epipod present.
</paragraph>
<paragraph id="8BB0D253821FFFC11BF8F8BFFBC65954" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,151,536]" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318">
Third pereiopod (
<figureCitation id="1334CED6821FFFC21AADF8BFFE335FFF" box="[402,495,1916,1941]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="17" pageNumber="318">Figs. 12</figureCitation>
A) largest, strongest, almost as long as entire body length, with few simple setae along margins of segments. Fingers elongate with heavily chitinized hooked crossing tips, distally bearing small tufts of long setae. Dactylus with distal chitinous ridge along cutting edge, proximally with large sharp tooth, smaller rounded tooth. Propodal cutting edge with distal chitinous ridge, proximally fossa for receiving large dactylar tooth, crowned with large rounded tooth followed by serrate ridge. Propodus longest segment with outer margin distally bearing dense fringe of setal brushes. Carpus less than 0.3 propodal length, narrowing proximally. Merus about 0.8 propodal length with distomedial ridge, 5 blunt dorsal spines, 4 blunt ventral spines. Ischium 0.5 meral length, unarmed. Basis and coxa short, stout, unarmed, no epipod present.
</paragraph>
<paragraph id="8BB0D253821CFFC11BF8FE8FFA82599E" blockId="18.[151,1437,151,536]" pageId="18" pageNumber="319">
Fourth, fifth pereiopods (
<figureCitation id="1334CED6821CFFC11AD9FE8FFD99590F" box="[486,581,332,357]" captionStart="FIGURE 14" captionStartId="16.[151,250,1651,1673]" captionTargetBox="[248,1332,196,1611]" captionTargetId="figure@16.[237,1349,193,1629]" captionTargetPageId="16" captionText="FIGURE 14. Spongiocaris neocaledonensis sp. nov. A – E, I, J, male holotype, MNHN Na- 12000; F, male paratype, NMNH Na- 12003; G, male paratype, MNHN Na- 12005; H, male paratype, MNHN Na- 12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F – H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240474/files/figure.png" pageId="18" pageNumber="319">
Figs. 
<date id="FFB1F493821CFFC11918FE8FFD99590F" box="[551,581,332,356]" pageId="18" pageNumber="319" value="2014-01">14</date>
</figureCitation>
I, J) long, slender, fifth slightly longer than fourth, with simple setae along margins. Dactyli triunguiculate, with smaller proximal tooth on accessory spine. Unguis short, curved, clearly separated from dactylar corpus. Propodi about 0.3 carpal lengths with 14 movable spines on ventral margins. Carpi longest segments, divided into 3 subsegments, last 2 terminating in movable spine at distodorsal margin. Meri about as long as carpi, ischia 0.3 times meral length. Bases and coxae short, stout, unarmed, without epipods.
</paragraph>
<paragraph id="8BB0D253821CFFC11BF8FE3CFAB75A73" blockId="18.[151,1437,151,536]" box="[199,1387,511,536]" pageId="18" pageNumber="319">
First pleopod (
<figureCitation id="1334CED6821CFFC11A54FE3CFE665A73" box="[363,442,511,536]" captionStart="FIGURE 12" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[248,1347,205,1487]" captionTargetId="figure@14.[237,1349,193,1498]" captionTargetPageId="14" captionText="FIGURE 12. Spongiocaris neocaledonensis sp. nov. Male holotype, MNHN Na- 12000. A, lateral view; B, carapace, dorsal view; C, fifth to sixth pleomeres, telson and uropods, dorsal view; D, thoracic sternites; E, epistome and labrum; F, right mandible, dorsal view; G, right mandible, ventral view; H, maxillae; I, maxilla. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240472/files/figure.png" pageId="18" pageNumber="319">Fig. 12</figureCitation>
A) uniramous, second to fifth biramous, all lacking appendices, bearing no spinules.
</paragraph>
<caption id="DF7082DB821CFFC11BA8FDAFFCD85AE8" ID-Table-UUID="DF7082DB821CFFC11BA8FDAFFCD85AE8" box="[151,772,619,643]" httpUri="http://table.plazi.org/id/DF7082DB821CFFC11BA8FDAFFCD85AE8" pageId="18" pageNumber="319" targetBox="[151,1200,777,922]" targetIsTable="true" targetPageId="18">
<paragraph id="8BB0D253821CFFC11BA8FDAFFCD85AE8" blockId="18.[151,772,619,643]" box="[151,772,619,643]" pageId="18" pageNumber="319">
<emphasis id="B97B0E41821CFFC11BA8FDAFFEDF5AE8" bold="true" box="[151,259,620,643]" pageId="18" pageNumber="319">TABLE 1</emphasis>
. Branchial formula of 
<taxonomicName id="4C0FA9D0821CFFC11AC8FDA8FD1D5AE8" box="[503,705,619,643]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="neocaledonensis" status="sp. nov.">
<emphasis id="B97B0E41821CFFC11AC8FDA8FD1D5AE8" box="[503,705,619,643]" italics="true" pageId="18" pageNumber="319">S. neocaledonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A248B33A821CFFC119F7FDAFFCD85AE8" box="[712,772,620,643]" pageId="18" pageNumber="319" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB0D253821CFFC11A8EFD6FFC7F5AA9" blockId="18.[433,1206,684,736]" box="[433,931,684,706]" pageId="18" pageNumber="319">Maxillipeds Pereiopods</paragraph>
<paragraph id="8BB0D253821CFFC11A88FD08FD715C00" blockId="18.[433,1206,684,736]" lastBlockId="18.[151,1437,998,2031]" pageId="18" pageNumber="319">
I II III I II III IV V 
<emphasis id="B97B0E41821CFFC11BF8FC25FEAA5B94" bold="true" box="[199,374,998,1023]" pageId="18" pageNumber="319">Measurements</emphasis>
(mm). 
<typeStatus id="54B46CF1821CFFC11AF5FC25FDEB5B94" box="[458,567,998,1023]" pageId="18" pageNumber="319" type="holotype">Holotype</typeStatus>
: PCL: 8.3; RCL: 11.8; TL: 31.3; Allotype—PCL: 10.5; RCL: 12.6; TL: 34.0. 
<typeStatus id="54B46CF1821CFFC11BA8FBC8FED85C48" box="[151,260,1035,1059]" pageId="18" pageNumber="319" type="paratype">Paratypes</typeStatus>
&amp; additional material: PCL: 4.3–11.4; RCL: 6.0–13.4; TL: 14.4–38.2. Eggs in 4 ovigerous females were few in number (2–9) while the allotype bore 
<specimenCount id="9D0919DA821CFFC119FAFBEDFCF95C2C" box="[709,805,1070,1095]" pageId="18" pageNumber="319" type="egg">62 eggs</specimenCount>
. All of the eggs were in early stages of embryonic development averaging 2.04 × 
<quantity id="4CF77FB6821CFFC11ACCFB90FD8B5C01" box="[499,599,1107,1131]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="319" unit="mm" value="1.7">1.70 mm</quantity>
in size.
</paragraph>
</subSubSection>
<paragraph id="8BB0D253821CFFC11BA8FCCAFB775BF1" pageId="18" pageNumber="319">
<table id="F90F20F3821C002C1BA8FCCAFB6C5BF1" box="[151,1200,777,922]" gridcols="9" gridrows="5" pageId="18" pageNumber="319">
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<th id="76EEB96D821C002C1BA8FCCAFEF55B74" box="[151,297,777,799]" gridcol="0" gridrow="0" pageId="18" pageNumber="319">Pleurobranchs</th>
<th id="76EEB96D821C002C1A8EFCCAFE635B74" box="[433,447,777,799]" gridcol="1" gridrow="0" pageId="18" pageNumber="319">-</th>
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<th id="76EEB96D821C002C1814FCCAFCE65B74" box="[811,826,777,799]" gridcol="4" gridrow="0" pageId="18" pageNumber="319">1</th>
<th id="76EEB96D821C002C18B6FCCAFC445B74" box="[905,920,777,799]" gridcol="5" gridrow="0" pageId="18" pageNumber="319">1</th>
<th id="76EEB96D821C002C18D8FCCAFC285B74" box="[999,1012,777,799]" gridcol="6" gridrow="0" pageId="18" pageNumber="319">1</th>
<th id="76EEB96D821C002C1F7AFCCAFB885B74" box="[1093,1108,777,799]" gridcol="7" gridrow="0" pageId="18" pageNumber="319">1</th>
<th id="76EEB96D821C002C1F9CFCCAFB6C5B74" box="[1187,1200,777,799]" gridcol="8" gridrow="0" pageId="18" pageNumber="319">1</th>
</tr>
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<th id="76EEB96D821C002C1BA8FCEBFEF55B55" box="[151,297,808,830]" gridcol="0" gridrow="1" pageId="18" pageNumber="319">Arthrobranchs</th>
<td id="76EEB96D821C002C1A8EFCEBFE635B55" box="[433,447,808,830]" gridcol="1" gridrow="1" pageId="18" pageNumber="319">1</td>
<td id="76EEB96D821C002C1930FCEBFDC05B55" box="[527,540,808,830]" gridcol="2" gridrow="1" pageId="18" pageNumber="319">1</td>
<td id="76EEB96D821C002C1952FCEBFDA05B55" box="[621,636,808,830]" gridcol="3" gridrow="1" pageId="18" pageNumber="319">2</td>
<td id="76EEB96D821C002C1814FCEBFCE65B55" box="[811,826,808,830]" gridcol="4" gridrow="1" pageId="18" pageNumber="319">2</td>
<td id="76EEB96D821C002C18B6FCEBFC445B55" box="[905,920,808,830]" gridcol="5" gridrow="1" pageId="18" pageNumber="319">2</td>
<td id="76EEB96D821C002C18D8FCEBFC285B55" box="[999,1012,808,830]" gridcol="6" gridrow="1" pageId="18" pageNumber="319">2</td>
<td id="76EEB96D821C002C1F7AFCEBFB885B55" box="[1093,1108,808,830]" gridcol="7" gridrow="1" pageId="18" pageNumber="319">2</td>
<td id="76EEB96D821C002C1F9CFCEBFB6C5B55" box="[1187,1200,808,830]" gridcol="8" gridrow="1" pageId="18" pageNumber="319">-</td>
</tr>
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<th id="76EEB96D821C002C1BA8FC85FEF55B37" box="[151,297,838,860]" gridcol="0" gridrow="2" pageId="18" pageNumber="319">Podobranchs</th>
<td id="76EEB96D821C002C1A8EFC85FE635B37" box="[433,447,838,860]" gridcol="1" gridrow="2" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1930FC85FDC05B37" box="[527,540,838,860]" gridcol="2" gridrow="2" pageId="18" pageNumber="319">1</td>
<td id="76EEB96D821C002C1952FC85FDA05B37" box="[621,636,838,860]" gridcol="3" gridrow="2" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1814FC85FCE65B37" box="[811,826,838,860]" gridcol="4" gridrow="2" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C18B6FC85FC445B37" box="[905,920,838,860]" gridcol="5" gridrow="2" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C18D8FC85FC285B37" box="[999,1012,838,860]" gridcol="6" gridrow="2" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1F7AFC85FB885B37" box="[1093,1108,838,860]" gridcol="7" gridrow="2" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1F9CFC85FB6C5B37" box="[1187,1200,838,860]" gridcol="8" gridrow="2" pageId="18" pageNumber="319">-</td>
</tr>
<tr id="353FD011821C002C1BA8FCA6FB6C5B10" box="[151,1200,869,891]" gridrow="3" pageId="18" pageNumber="319">
<th id="76EEB96D821C002C1BA8FCA6FEF55B10" box="[151,297,869,891]" gridcol="0" gridrow="3" pageId="18" pageNumber="319">Epipods</th>
<td id="76EEB96D821C002C1A8EFCA6FE635B10" box="[433,447,869,891]" gridcol="1" gridrow="3" pageId="18" pageNumber="319">1</td>
<td id="76EEB96D821C002C1930FCA6FDC05B10" box="[527,540,869,891]" gridcol="2" gridrow="3" pageId="18" pageNumber="319">1</td>
<td id="76EEB96D821C002C1952FCA6FDA05B10" box="[621,636,869,891]" gridcol="3" gridrow="3" pageId="18" pageNumber="319">1</td>
<td id="76EEB96D821C002C1814FCA6FCE65B10" box="[811,826,869,891]" gridcol="4" gridrow="3" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C18B6FCA6FC445B10" box="[905,920,869,891]" gridcol="5" gridrow="3" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C18D8FCA6FC285B10" box="[999,1012,869,891]" gridcol="6" gridrow="3" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1F7AFCA6FB885B10" box="[1093,1108,869,891]" gridcol="7" gridrow="3" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1F9CFCA6FB6C5B10" box="[1187,1200,869,891]" gridcol="8" gridrow="3" pageId="18" pageNumber="319">-</td>
</tr>
<tr id="353FD011821C002C1BA8FC47FB6C5BF1" box="[151,1200,900,922]" gridrow="4" pageId="18" pageNumber="319">
<th id="76EEB96D821C002C1BA8FC47FEF55BF1" box="[151,297,900,922]" gridcol="0" gridrow="4" pageId="18" pageNumber="319">Exopods</th>
<td id="76EEB96D821C002C1A8EFC47FE635BF1" box="[433,447,900,922]" gridcol="1" gridrow="4" pageId="18" pageNumber="319">1</td>
<td id="76EEB96D821C002C1930FC47FDC05BF1" box="[527,540,900,922]" gridcol="2" gridrow="4" pageId="18" pageNumber="319">1</td>
<td id="76EEB96D821C002C1952FC47FDA05BF1" box="[621,636,900,922]" gridcol="3" gridrow="4" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1814FC47FCE65BF1" box="[811,826,900,922]" gridcol="4" gridrow="4" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C18B6FC47FC445BF1" box="[905,920,900,922]" gridcol="5" gridrow="4" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C18D8FC47FC285BF1" box="[999,1012,900,922]" gridcol="6" gridrow="4" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1F7AFC47FB885BF1" box="[1093,1108,900,922]" gridcol="7" gridrow="4" pageId="18" pageNumber="319">-</td>
<td id="76EEB96D821C002C1F9CFC47FB6C5BF1" box="[1187,1200,900,922]" gridcol="8" gridrow="4" pageId="18" pageNumber="319">-</td>
</tr>
</table>
</paragraph>
<subSubSection id="C31581D8821CFFC11BF8FBB5FBF75CE5" box="[199,1067,1142,1167]" pageId="18" pageNumber="319" type="distribution">
<paragraph id="8BB0D253821CFFC11BF8FBB5FBF75CE5" blockId="18.[151,1437,998,2031]" box="[199,1067,1142,1167]" pageId="18" pageNumber="319">
<emphasis id="B97B0E41821CFFC11BF8FBB5FE855CE4" bold="true" box="[199,345,1142,1167]" pageId="18" pageNumber="319">Distribution</emphasis>
. 
<collectingCountry id="F31892C3821CFFC11A58FBB4FDC45CE4" box="[359,536,1142,1167]" name="New Caledonia" pageId="18" pageNumber="319">New Caledonia</collectingCountry>
, 
<collectingCountry id="F31892C3821CFFC1191DFBB5FDA75CE4" box="[546,635,1142,1167]" name="Norfolk Island" pageId="18" pageNumber="319">Norfolk</collectingCountry>
Ridge; Loyalty Islands in 
<quantity id="4CF77FB6821CFFC1189AFBB5FBFB5CE5" box="[933,1063,1142,1167]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="9.7" metricValueMin="4.6" pageId="18" pageNumber="319" unit="m" value="715.0" valueMax="970.0" valueMin="460.0">
460– 
<specimenCount id="9D0919DA821CFFC118E2FBB5FBFB5CE5" box="[989,1063,1142,1167]" pageId="18" pageNumber="319" type="generic">970 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C31581D8821CFFC11BF8FB59FE105CBC" pageId="18" pageNumber="319" type="etymology">
<paragraph id="8BB0D253821CFFC11BF8FB59FAC55CD9" blockId="18.[151,1437,998,2031]" box="[199,1305,1178,1203]" pageId="18" pageNumber="319">
<emphasis id="B97B0E41821CFFC11BF8FB59FE9A5CD8" bold="true" box="[199,326,1178,1203]" pageId="18" pageNumber="319">Etymology</emphasis>
. The name 
<taxonomicName id="4C0FA9D0821CFFC11AF5FB58FD5F5CD8" box="[458,643,1179,1203]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="neocaledonensis">
<emphasis id="B97B0E41821CFFC11AF5FB58FD5F5CD8" box="[458,643,1179,1203]" italics="true" pageId="18" pageNumber="319">neocaledonensis</emphasis>
</taxonomicName>
refers to the species 
<typeStatus id="54B46CF1821CFFC11850FB58FC435CD8" box="[879,927,1179,1203]" pageId="18" pageNumber="319">type</typeStatus>
locality, New Caledonian waters.
</paragraph>
<paragraph id="8BB0D253821CFFC11BF8FB7DFE105CBC" blockId="18.[151,1437,998,2031]" box="[199,460,1214,1239]" pageId="18" pageNumber="319">
<emphasis id="B97B0E41821CFFC11BF8FB7DFE945CBC" bold="true" box="[199,328,1214,1239]" pageId="18" pageNumber="319">Coloration</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31581D8821CFFC01BF8FB21FEA259BB" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319" type="materials_examined">
<paragraph id="8BB0D253821CFFC11BF8FB21FBF75C91" blockId="18.[151,1437,998,2031]" box="[199,1067,1250,1275]" pageId="18" pageNumber="319">
<emphasis id="B97B0E41821CFFC11BF8FB21FE815C90" bold="true" box="[199,349,1250,1275]" pageId="18" pageNumber="319">
<typeStatus id="54B46CF1821CFFC11BF8FB21FEDE5C90" box="[199,258,1250,1275]" pageId="18" pageNumber="319">Type</typeStatus>
locality
</emphasis>
. 
<collectingCountry id="F31892C3821CFFC11A53FB20FDC05C90" box="[364,540,1250,1275]" name="New Caledonia" pageId="18" pageNumber="319">New Caledonia</collectingCountry>
, 
<collectingCountry id="F31892C3821CFFC11918FB21FD5C5C90" box="[551,640,1250,1275]" name="Norfolk Island" pageId="18" pageNumber="319">Norfolk</collectingCountry>
Ridge, 
<geoCoordinate id="EE3BB494821CFFC119E7FB21FC975C90" box="[728,843,1250,1275]" direction="south" orientation="latitude" pageId="18" pageNumber="319" precision="92" value="-22.996666">22°59.8’S</geoCoordinate>
, 
<geoCoordinate id="EE3BB494821CFFC11868FB21FC065C90" box="[855,986,1250,1275]" direction="east" orientation="longitude" pageId="18" pageNumber="319" precision="92" value="167.40334">167°24.2’E</geoCoordinate>
, 
<quantity id="4CF77FB6821CFFC118DAFB21FBF75C91" box="[997,1067,1250,1275]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="18" pageNumber="319" unit="m" value="560.0">560 m</quantity>
</paragraph>
<paragraph id="8BB0D253821CFFC11BF8FAC5FB9A5DC5" blockId="18.[151,1437,998,2031]" pageId="18" pageNumber="319">
Allotype (female, MNHN-NA-12000) (
<figureCitation id="1334CED6821CFFC119B4FAC5FD025D74" box="[651,734,1286,1311]" captionStart="FIGURE 13" captionStartId="15.[151,250,1880,1902]" captionTargetBox="[232,1345,492,1851]" captionTargetId="figure@15.[222,1365,481,1859]" captionTargetPageId="15" captionText="FIGURE 13. Spongiocaris neocaledonensis sp. nov. Female allotype, MNHN Na- 12000. A, carapace and abdomen, lateral view; B, carapace, dorsal view; C, fifth, sixth pleomeres, telson and uropods; D, thoracic sternites; E, egg; F; dactylus, propodus and carpus third pereiopod. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240473/files/figure.png" pageId="18" pageNumber="319">Figs.13</figureCitation>
A–D). Sexual differences from the 
<typeStatus id="54B46CF1821CFFC11F48FAC5FB055D74" box="[1143,1241,1286,1311]" pageId="18" pageNumber="319" type="holotype">holotype</typeStatus>
include: inflated carapace; broader thoracic sternites with fourth, fifth bearing 2 rounded projections, others unarmed; no ventral median spine on fifth abdominal somite; abdominal pleura expanded, overlapping each other; posterior margin of fifth abdominal somite with 5 spines; sixth abdominal somite with row of 3 dorsal spines, 9 posterior margin spines; third pereiopod completely glabrous; pleopods much larger and more setose.
</paragraph>
<paragraph id="8BB0D253821CFFC11BF8FA79FC785E98" blockId="18.[151,1437,998,2031]" pageId="18" pageNumber="319">
Five 
<typeStatus id="54B46CF1821CFFC11BC0FA78FEB75DB8" box="[255,363,1467,1491]" pageId="18" pageNumber="319" type="paratype">paratypes</typeStatus>
and 5 additional specimens agree well with the 
<typeStatus id="54B46CF1821CFFC118BDFA79FC385DB8" box="[898,996,1466,1491]" pageId="18" pageNumber="319" type="holotype">holotype</typeStatus>
and allotype and show the same sexual differences. The spination of the third pereiopods varies in the males (
<figureCitation id="1334CED6821CFFC11884FA1DFBCB5D9C" box="[955,1047,1502,1527]" captionStart="FIGURE 14" captionStartId="16.[151,250,1651,1673]" captionTargetBox="[248,1332,196,1611]" captionTargetId="figure@16.[237,1349,193,1629]" captionTargetPageId="16" captionText="FIGURE 14. Spongiocaris neocaledonensis sp. nov. A – E, I, J, male holotype, MNHN Na- 12000; F, male paratype, NMNH Na- 12003; G, male paratype, MNHN Na- 12005; H, male paratype, MNHN Na- 12006. A, first maxilliped; B, second maxilliped; C, third maxilliped; D, first pereiopod; E, second pereiopod; F – H, third pereiopod; I; fourth pereiopod; J, fifth pereiopod. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/240474/files/figure.png" pageId="18" pageNumber="319">Figs. 14</figureCitation>
F–H) with larger males being less spinose and the meral distdorsal ridge ending in 1 or 2 spines. Spination of the carapace is variable with some specimens bearing hepatic, superior and inferior orbital spines and others lacking an antennal spine. Other variation in spination include: 4–6 dorsal, 0–1 ventral and 0–2 lateral spines on the rostrum; 4–10 cervical groove spines; 4– 6 outer margin spines on scaphocerite; 8–16 ventral movable spines on the propodi of the fourth and fifth pereiopods; 4–9 spines on the telson’s median carinae; 5–10 spines on telson’s lateral border; 6–12 posteror teeth on telson; and 8–14 teeth on outer margin of uropodal exopodite. This slight variation in body spination probably reflect allometric growth changes and normal variation in the species.
</paragraph>
<paragraph id="8BB0D253821CFFC11BF8F93DFAD35F34" blockId="18.[151,1437,998,2031]" pageId="18" pageNumber="319">
<emphasis id="B97B0E41821CFFC11BF8F93DFF235F7C" bold="true" box="[199,255,1790,1815]" pageId="18" pageNumber="319">Host</emphasis>
. Three females were taken from the hexactinellid sponge 
<emphasis id="B97B0E41821CFFC118EAF93DFB3C5F7D" box="[981,1248,1790,1814]" italics="true" pageId="18" pageNumber="319">Regardella okinoseana</emphasis>
Ijima, 1901 at MUSORSTOM 4 station 215 and another female from the same sponge species at BIOCAL station 46. The second largest male (MNHN-NA 12003) was collected on a gorgonian octocoral by the CALSUB expedition.
</paragraph>
<paragraph id="8BB0D253821CFFC01BF8F8A9FEA259BB" blockId="18.[151,1437,998,2031]" lastBlockId="19.[151,1437,151,1112]" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">
<emphasis id="B97B0E41821CFFC11BF8F8A9FE6C5FE8" bold="true" box="[199,432,1898,1923]" pageId="18" pageNumber="319">Systematic position</emphasis>
. 
<taxonomicName id="4C0FA9D0821CFFC11A80F8A8FCC95FE8" box="[447,789,1899,1923]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="neocaledonensis">
<emphasis id="B97B0E41821CFFC11A80F8A8FCC95FE8" box="[447,789,1899,1923]" italics="true" pageId="18" pageNumber="319">Spongiocaris neocaledonensis</emphasis>
</taxonomicName>
closely follows the definition of the genus 
<taxonomicName id="4C0FA9D0821CFFC11E37F8A8FA405FE9" box="[1288,1436,1899,1922]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B97B0E41821CFFC11E37F8A8FA405FE9" box="[1288,1436,1899,1922]" italics="true" pageId="18" pageNumber="319">Spongiocaris</emphasis>
</taxonomicName>
given by 
<bibRefCitation id="EF9EAFA2821CFFC11A3BF84CFDDE5FCC" author="Bruce" box="[260,514,1934,1959]" pageId="18" pageNumber="319" refString="Bruce A. J. &amp; Baba, K. (1973) Spongiocaris, a new genus of stenopodidean shrimp from New Zealand and South African waters, with a description of two new species (Decapoda Natantia, Stenopodidea). Crustaceana, 25, 153 - 170." type="journal article" year="1973">Bruce &amp; Baba (1973)</bibRefCitation>
. The genus 
<taxonomicName id="4C0FA9D0821CFFC119AFF84CFCF85FCD" box="[656,804,1935,1958]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B97B0E41821CFFC119AFF84CFCF85FCD" box="[656,804,1935,1958]" italics="true" pageId="18" pageNumber="319">Spongiocaris</emphasis>
</taxonomicName>
needs to be revised (
<bibRefCitation id="EF9EAFA2821CFFC11F22F84DFACE5FCC" author="Saito" box="[1053,1298,1934,1959]" pageId="18" pageNumber="319" refString="Saito, T. &amp; Komai, T. (2008) A review of species of the genera Spongicola de Haan, 1844 and Paraspongicola de Saint Laurent &amp; Cleva, 1981 (Crustacea, Decapoda, Stenopodidea, Spongicolidae). Zoosystema, 30 (1), 87 - 147." type="journal article" year="2008">Saito &amp; Komai 2008</bibRefCitation>
; 
<bibRefCitation id="EF9EAFA2821CFFC11E1EF84CFA495FCC" author="Goy" box="[1313,1429,1935,1959]" pageId="18" pageNumber="319" refString="Goy, J. W. (2010) Infraorder Stenopodidea Claus 1872. In: Schram, F. R., von Vaupel Klein, J. C., Forest, J. &amp; Charmantier- Daures, M. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, Decapoda, Volume 9 Part A Eucarida: Euphausiacea, Amphionidacea, and Decapoda (partim). Vol. 9 A, 215 - 265. Brill, Leiden, Netherlands." type="journal article" year="2010">Goy 2010</bibRefCitation>
) since some members assigned to other spongicolid genera actually need to be transferred to 
<taxonomicName id="4C0FA9D0821CFFC11F8AF870FA955FA1" box="[1205,1353,1971,1994]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B97B0E41821CFFC11F8AF870FA955FA1" box="[1205,1353,1971,1994]" italics="true" pageId="18" pageNumber="319">Spongiocaris</emphasis>
</taxonomicName>
. These species include 
<taxonomicName id="4C0FA9D0821CFFC11A69F815FD3C5F84" authority="Kubo, 1942" authorityName="Kubo" authorityYear="1942" box="[342,736,2006,2031]" class="Malacostraca" family="Spongicolidae" genus="Spongicola" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B97B0E41821CFFC11A69F815FD985F85" box="[342,580,2006,2030]" italics="true" pageId="18" pageNumber="319">Spongicola japonica</emphasis>
<bibRefCitation id="EF9EAFA2821CFFC1196DF815FD3C5F84" author="Kubo" box="[594,736,2006,2031]" pageId="18" pageNumber="319" refString="Kubo, I. (1942) A new commensal shrimp, Spongicola japonica, n. sp. Annotationes Zoologicae Japonenses, 21 (2), 90 - 94." type="journal article" year="1942">
<collectingCountry id="F31892C3821CFFC1196DF815FD4F5F84" box="[594,659,2006,2031]" name="Cuba" pageId="18" pageNumber="319">Kubo</collectingCountry>
, 1942
</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C0FA9D0821CFFC119CCF815FABD5F84" authority="Ortiz, Gomez &amp; Lalana, 1994" authorityName="Ortiz, Gomez &amp; Lalana" authorityYear="1994" box="[755,1377,2006,2031]" class="Malacostraca" family="Spongicolidae" genus="Spongicola" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="cubanica">
<emphasis id="B97B0E41821CFFC119CCF815FC395F85" box="[755,997,2006,2030]" italics="true" pageId="18" pageNumber="319">Spongicola cubanica</emphasis>
<bibRefCitation id="EF9EAFA2821CFFC118CBF815FABD5F84" author="Ortiz" box="[1012,1377,2006,2031]" pageId="18" pageNumber="319" refString="Ortiz. M., Gomez, O. &amp; Lalana, R. (1994) Un nuevo camaron (Stenopodidea) asociado a una esponja silicea (Hexactinellida) de Cuba. Caribbean Journal of Science, 30, 189 - 196." type="journal article" year="1994">Ortiz, Gómez &amp; Lalana, 1994</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C0FA9D0821DFFC01BA8FF5BFDA758DB" authority="Caullery, 1896" authorityName="Caullery" authorityYear="1896" box="[151,635,151,176]" class="Malacostraca" family="Spongicolidae" genus="Spongicoloides" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="koehleri">
<emphasis id="B97B0E41821DFFC01BA8FF5BFE7058DB" box="[151,428,152,176]" italics="true" pageId="19" pageNumber="320">Spongicoloides koehleri</emphasis>
(
<bibRefCitation id="EF9EAFA2821DFFC01AFDFF54FDAF58DB" author="Caullery" box="[450,627,151,176]" pageId="19" pageNumber="320" refString="Caullery, M. (1896) Resultats scientifiques de la Campagne du &quot; Caudan &quot; dans la Golfe de Gascogne. Aout-Septembre 1885. Crustaces Schizopodes et Decapodes. Annales des Universite Lyons. 2, 365 - 419." type="journal article" year="1896">Caullery, 1896</bibRefCitation>
)
</taxonomicName>
. A revision of the genera 
<taxonomicName id="4C0FA9D0821DFFC018F8FF5BFAB958DB" authority="Bruce &amp; Baba, 1973" authorityName="Bruce &amp; Baba" authorityYear="1973" box="[967,1381,151,176]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B97B0E41821DFFC018F8FF5BFB8758C4" box="[967,1115,152,175]" italics="true" pageId="19" pageNumber="320">Spongiocaris</emphasis>
<bibRefCitation id="EF9EAFA2821DFFC01F56FF5BFAB958DB" author="Bruce" box="[1129,1381,151,176]" pageId="19" pageNumber="320" refString="Bruce A. J. &amp; Baba, K. (1973) Spongiocaris, a new genus of stenopodidean shrimp from New Zealand and South African waters, with a description of two new species (Decapoda Natantia, Stenopodidea). Crustaceana, 25, 153 - 170." type="journal article" year="1973">Bruce &amp; Baba, 1973</bibRefCitation>
</taxonomicName>
and 
<emphasis id="B97B0E41821DFFC01BA8FF7FFEE858BF" box="[151,308,188,212]" italics="true" pageId="19" pageNumber="320">Spongicolides</emphasis>
<bibRefCitation id="EF9EAFA2821DFFC01A01FF7EFE0058BF" author="Hansen" box="[318,476,188,212]" pageId="19" pageNumber="320" refString="Hansen, H. J. (1908) Crustacea Malacostraca I. The Danish Ingolf-Expedition, 3 (2), 1 - 120." type="journal article" year="1908">Hansen, 1908</bibRefCitation>
is in progress to affect these transfers. That said, 
<taxonomicName id="4C0FA9D0821DFFC01F17FF7FFAA358BF" box="[1064,1407,188,212]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="neocaledonensis">
<emphasis id="B97B0E41821DFFC01F17FF7FFAA358BF" box="[1064,1407,188,212]" italics="true" pageId="19" pageNumber="320">Spongiocaris neocaledonensis</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C0FA9D0821DFFC01A6AFF23FD9E589C" box="[341,578,224,248]" class="Malacostraca" family="Spongicolidae" genus="Spongicola" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B97B0E41821DFFC01A6AFF23FD9E589C" box="[341,578,224,248]" italics="true" pageId="19" pageNumber="320">Spongicola japonica</emphasis>
</taxonomicName>
in rostral, carapacial and telsonal spination, while the new species’ eye pigmentation, morphology of the third pereiopods approaches the conditions seen in 
<taxonomicName id="4C0FA9D0821DFFC01F5BFEC7FAA95977" box="[1124,1397,260,284]" class="Malacostraca" family="Spongicolidae" genus="Spongicoloides" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="koehleri">
<emphasis id="B97B0E41821DFFC01F5BFEC7FAA95977" box="[1124,1397,260,284]" italics="true" pageId="19" pageNumber="320">Spongicoloides koehleri</emphasis>
</taxonomicName>
. In overall appearance 
<taxonomicName id="4C0FA9D0821DFFC01A4EFEEBFD9B592B" box="[369,583,296,320]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="neocaledonensis">
<emphasis id="B97B0E41821DFFC01A4EFEEBFD9B592B" box="[369,583,296,320]" italics="true" pageId="19" pageNumber="320">S. neocaledonensis</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C0FA9D0821DFFC019F9FEEBFC0B592B" authority="Goy, 2010" authorityName="Goy" authorityYear="2010" box="[710,983,296,320]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="B97B0E41821DFFC019F9FEEBFC9B592B" box="[710,839,296,320]" italics="true" pageId="19" pageNumber="320">S. yaldwyni</emphasis>
(
<bibRefCitation id="EF9EAFA2821DFFC01868FEEBFC13592B" author="Goy" box="[855,975,296,320]" pageId="19" pageNumber="320" refString="Goy, J. W. (2010) Infraorder Stenopodidea Claus 1872. In: Schram, F. R., von Vaupel Klein, J. C., Forest, J. &amp; Charmantier- Daures, M. (Eds.), Treatise on Zoology - Anatomy, Taxonomy, Biology - The Crustacea, Decapoda, Volume 9 Part A Eucarida: Euphausiacea, Amphionidacea, and Decapoda (partim). Vol. 9 A, 215 - 265. Brill, Leiden, Netherlands." type="journal article" year="2010">Goy, 2010</bibRefCitation>
)
</taxonomicName>
and its branchial formula is the same as 
<taxonomicName id="4C0FA9D0821DFFC01BA8FE8FFEC2590F" box="[151,286,332,356]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="B97B0E41821DFFC01BA8FE8FFEC2590F" box="[151,286,332,356]" italics="true" pageId="19" pageNumber="320">S. yaldwyni</emphasis>
</taxonomicName>
, except for the presence of an epipod on the third maxillipeds. The new species can be easily distinguished from the other seven species in the genus by the segmentation of the carpus of the last two pairs of pereiopods, the presence of the third maxilliped’s setiferous organ, and the dense fringe of setal brushes on the third pereiopod propodus.
</paragraph>
</subSubSection>
<subSubSection id="C31581D8821DFFC01BF8FE1FFCE95C3C" pageId="19" pageNumber="320" type="discussion">
<paragraph id="8BB0D253821DFFC01BF8FE1FFB995A9B" blockId="19.[151,1437,151,1112]" pageId="19" pageNumber="320">
<emphasis id="B97B0E41821DFFC01BF8FE1FFEE8599E" bold="true" box="[199,308,476,501]" pageId="19" pageNumber="320">Remarks</emphasis>
. 
<taxonomicName id="4C0FA9D0821DFFC01A79FE1FFD42599F" box="[326,670,476,500]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="neocaledonensis">
<emphasis id="B97B0E41821DFFC01A79FE1FFD42599F" box="[326,670,476,500]" italics="true" pageId="19" pageNumber="320">Spongiocaris neocaledonensis</emphasis>
</taxonomicName>
is the only member of the genus with well developed setiferous organs on the third maxillipeds and first pereiopods; approaching the condition of many members of the family 
<taxonomicName id="4C0FA9D0821DFFC01BA8FDE7FEEF5A56" box="[151,307,548,573]" class="Malacostraca" family="Stenopodidae" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="family">Stenopodidae</taxonomicName>
. All other members of the genus 
<taxonomicName id="4C0FA9D0821DFFC01993FDE7FC9C5A50" box="[684,832,548,571]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B97B0E41821DFFC01993FDE7FC9C5A50" box="[684,832,548,571]" italics="true" pageId="19" pageNumber="320">Spongiocaris</emphasis>
</taxonomicName>
lack these structures, except 
<taxonomicName id="4C0FA9D0821DFFC01FB2FDE7FA405A57" box="[1165,1436,548,572]" class="Malacostraca" family="Spongicolidae" genus="Spongicoloides" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="koehleri">
<emphasis id="B97B0E41821DFFC01FB2FDE7FA405A57" box="[1165,1436,548,572]" italics="true" pageId="19" pageNumber="320">Spongicoloides koehleri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0FA9D0821DFFC01BF7FD8BFE6D5A34" box="[200,433,584,608]" class="Malacostraca" family="Spongicolidae" genus="Spongicola" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B97B0E41821DFFC01BF7FD8BFE6D5A34" box="[200,433,584,608]" italics="true" pageId="19" pageNumber="320">Spongicola japonica</emphasis>
</taxonomicName>
which bear rudimentary setiferous organs on the first pereiopods. The main function of these setiferous organs in shallow water stenopodideans is for self-grooming (Bauer 1981, 1989). The function of these structures in a deep water hexactinellid sponge commensal is hard to explain. Perhaps in combination with the dense setal brushes on the third pereiopods, these setiferous organs are used to sweep silty deposits off the shrimp’s body as well as the internal sponge chamber in which the shrimp occupies.
</paragraph>
<paragraph id="8BB0D253821DFFC01BF8FD3FFCE95C3C" blockId="19.[151,1437,151,1112]" pageId="19" pageNumber="320">
The bathymetric range of the new species is from 
<quantity id="4CF77FB6821DFFC019CCFD3FFC545B7F" box="[755,904,764,789]" metricMagnitude="2" metricUnit="m" metricValue="7.13" metricValueMax="9.66" metricValueMin="4.6" pageId="19" pageNumber="320" unit="m" value="713.0" valueMax="966.0" valueMin="460.0">460 to 966 m</quantity>
, which is similar to that of 
<taxonomicName id="4C0FA9D0821DFFC01F8AFD3FFA405B7F" box="[1205,1436,764,788]" class="Malacostraca" family="Spongicolidae" genus="Spongicola" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B97B0E41821DFFC01F8AFD3FFA405B7F" box="[1205,1436,764,788]" italics="true" pageId="19" pageNumber="320">Spongicola japonica</emphasis>
</taxonomicName>
at 
<quantity id="4CF77FB6821DFFC01B8CFCE3FEEA5B5C" box="[179,310,800,824]" metricMagnitude="2" metricUnit="m" metricValue="5.535" metricValueMax="8.07" metricValueMin="3.0" pageId="19" pageNumber="320" unit="m" value="553.5" valueMax="807.0" valueMin="300.0">
300– 
<specimenCount id="9D0919DA821DFFC01BD3FCE3FEEA5B5C" box="[236,310,800,824]" pageId="19" pageNumber="320" type="generic">807 m</specimenCount>
</quantity>
. Collection of deep water hexactinellid sponges and careful examination of their internal chambers can yield commensals, as evidenced by the capture of four specimens of 
<taxonomicName id="4C0FA9D0821DFFC018E9FC87FAF75B37" box="[982,1323,836,860]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="neocaledonensis">
<emphasis id="B97B0E41821DFFC018E9FC87FAF75B37" box="[982,1323,836,860]" italics="true" pageId="19" pageNumber="320">Spongiocaris neocaledonensis</emphasis>
</taxonomicName>
from two specimens of the hexactinellid sponge 
<emphasis id="B97B0E41821DFFC01976FCABFC965BEB" box="[585,842,872,896]" italics="true" pageId="19" pageNumber="320">Regardella okinoseana</emphasis>
. The unique 
<typeStatus id="54B46CF1821DFFC018E0FCA4FB9D5BEB" box="[991,1089,871,896]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C0FA9D0821DFFC01F59FCABFB3B5BEB" box="[1126,1255,872,896]" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="yaldwyni">
<emphasis id="B97B0E41821DFFC01F59FCABFB3B5BEB" box="[1126,1255,872,896]" italics="true" pageId="19" pageNumber="320">S. yaldwyni</emphasis>
</taxonomicName>
was taken from 
<emphasis id="B97B0E41821DFFC01BA8FC4FFE415BCF" box="[151,413,908,932]" italics="true" pageId="19" pageNumber="320">Regardella okiniseana</emphasis>
like the new species. Use of deepsea submersibles can uncover new commensal relationships that are normally lost to trawling or dredging sampling methods. The capture of a male 
<taxonomicName id="4C0FA9D0821DFFC01EB8FC73FE8C5B87" class="Malacostraca" family="Spongicolidae" genus="Spongiocaris" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="neocaledonensis">
<emphasis id="B97B0E41821DFFC01EB8FC73FE8C5B87" italics="true" pageId="19" pageNumber="320">S. neocaledonensis</emphasis>
</taxonomicName>
on a gorgorian octocoral was accomplished by the use of the deepsea submersible “Cyana” during the CALSUB expedition. This represents the first record of a stenopodidean shrimp in a commensal relationship with an alcyonarian. Continued future use of deepsea submersibles for invertebrate collections may reveal other new and interesting symbiotic associations such as this one.
</paragraph>
</subSubSection>
</treatment>
</document>