328 lines
44 KiB
XML
328 lines
44 KiB
XML
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<mods:title id="58328601A31DA684B6B6B4C73B7E08B0">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:name id="B23DC035A8BC7A7B76437016ABC20095" type="personal">
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<mods:namePart id="43B6BB707D3DFE10E61B28278F25544A">Merle, Didier</mods:namePart>
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<mods:namePart id="E94177CB67D2EEF72D6C9E8CD25D94BF">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="288BC79400CDBA8C66C2AA2EBA91D327">Ledon, Daniel</mods:namePart>
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<mods:affiliation id="21836387D71BCDCF6E70A33042A6FAB1">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:namePart id="3D15C21F21CD3C8D13290B73A0F35321">Goret, Bernard</mods:namePart>
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<mods:affiliation id="634ED60DA6DF5D4E92C8BFFE6C07D93A">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:title id="CD7B73D49EF445DC42EF42A2756BBCAC">Geodiversitas</mods:title>
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<mods:date id="558F717FA8929EADEE88379BD42851E2">2024</mods:date>
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<mods:number id="38E7E4223A67C64E6A188681CD1578A7">2024-10-24</mods:number>
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<taxonomicName id="4C274D4FFFF2E412FC5DFB63FB37FB7C" ID-CoL="CN5MZ" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[997,1206,1205,1231]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="calciacus" status="sp. nov.">
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<emphasis id="B953EADEFFF2E412FC5DFB63FB37FB7C" bold="true" box="[997,1206,1205,1231]" italics="true" pageId="20" pageNumber="513">Timbellus calciacus</emphasis>
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(
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A-D; 26C, D)
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</paragraph>
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<taxonomicName id="4C274D4FFFF2E412FC95FA8CFB81FAC1" ID-CoL="8ZHC9" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[813,1024,1370,1394]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
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<emphasis id="B953EADEFFF2E412FC95FA8CFB81FAC1" box="[813,1024,1370,1394]" italics="true" pageId="20" pageNumber="513">Pterynotus tripteroides</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0A8610DDFFF2E412FBA7FA8CFB4BFAC2" author="MERLE D." box="[1055,1226,1369,1394]" page="185" pageId="20" pageNumber="513" year="1990">
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<bibRefCitation id="EFB64B3DFFF2E412FBA7FA8CFB4BFAC2" author="MERLE D." box="[1055,1226,1369,1394]" pageId="20" pageNumber="513" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">Merle 1990: 185</bibRefCitation>
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</treatmentCitation>
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, pl. 4, fig. 3 only [
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<emphasis id="B953EADEFFF2E412FA33FA8CFA2EFAC2" box="[1419,1455,1370,1393]" italics="true" pageId="20" pageNumber="513">non</emphasis>
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].
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<materialsCitation id="3B4F3C91FFF2E412FC95FA72FBAEF9B8" ID-GBIF-Occurrence="4958671314" collectionCode="MNHN" collectorName="de Chaussy" country="France" location="Val d'Oise" municipality="Falun de Chaussy" pageId="20" pageNumber="513" specimenCount="1" typeStatus="holotype">
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. —
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.
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<collectingCountry id="F330765CFFF2E412FBD0FA72FB2AFA0F" box="[1128,1195,1444,1468]" name="France" pageId="20" pageNumber="513">France</collectingCountry>
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</emphasis>
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• Paris Basin,
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, Chaussy (Les Garennes);
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Falun
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(biozone NP15); middle Eocene (middle Lutetian);
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.F.A90536 (
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<taxonomicName id="4C274D4FFFF2E412FAAFFA0FFAD3FA42" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1303,1362,1497,1521]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
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coll.), H:
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(
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<figureCitation id="131C2A49FFF2E412FCCEFA25FC6BF9B8" box="[886,1002,1523,1547]" captionStart="FIG" captionStartId="21.[132,143,787,804]" captionTargetBox="[132,1455,214,744]" captionTargetId="figure-674@21.[698,1267,213,745]" captionTargetPageId="21" captionText="FIG. 10. — Spiral cords and internal denticles of the outer lip in Timbellus calciacusn. sp.:A, B, holotype,MNHN.F.A90536 (Merle coll.),Chaussy (Les Garennes),Val d’Oise,Paris Basin,France,middle Eocene (Lutetian);C, D, paratype,MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes).Scale bars:3 mm.Credits:L.Cazes." figureDoi="http://doi.org/10.5281/zenodo.13985782" httpUri="https://zenodo.org/record/13985782/files/figure.png" pageId="20" pageNumber="513">Figs 10A, B</figureCitation>
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;
|
||
<figureCitation id="131C2A49FFF2E412FC4DFA25FBA3F9B8" box="[1013,1058,1523,1547]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="20" pageNumber="513">26C</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF2E412FC95F9D8FC2CF919" blockId="20.[811,1457,1444,1706]" pageId="20" pageNumber="513">
|
||
<materialsCitation id="3B4F3C91FFF2E412FC95F9D8FC20F9E9" ID-GBIF-Occurrence="4958671333" pageId="20" pageNumber="513">
|
||
<emphasis id="B953EADEFFF2E412FC95F9D8FC4FF995" bold="true" box="[813,974,1550,1574]" pageId="20" pageNumber="513">
|
||
<typeStatus id="549C886EFFF2E412FC95F9D8FC08F995" box="[813,905,1550,1574]" pageId="20" pageNumber="513" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
<collectingCountry id="F330765CFFF2E412FC28F9D8FC4FF995" box="[912,974,1550,1574]" name="France" pageId="20" pageNumber="513">France</collectingCountry>
|
||
</emphasis>
|
||
• 3 spm; same as for the holotype;
|
||
<collectionCode id="ED36AE09FFF2E412FB45F9D8FACFF996" box="[1277,1358,1550,1573]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="513">MNHN</collectionCode>
|
||
.F.A90537 (
|
||
<figureCitation id="131C2A49FFF2E412FC8CF9FEFC23F9F3" box="[820,930,1576,1600]" captionStart="FIG" captionStartId="21.[132,143,787,804]" captionTargetBox="[132,1455,214,744]" captionTargetId="figure-674@21.[698,1267,213,745]" captionTargetPageId="21" captionText="FIG. 10. — Spiral cords and internal denticles of the outer lip in Timbellus calciacusn. sp.:A, B, holotype,MNHN.F.A90536 (Merle coll.),Chaussy (Les Garennes),Val d’Oise,Paris Basin,France,middle Eocene (Lutetian);C, D, paratype,MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes).Scale bars:3 mm.Credits:L.Cazes." figureDoi="http://doi.org/10.5281/zenodo.13985782" httpUri="https://zenodo.org/record/13985782/files/figure.png" pageId="20" pageNumber="513">Figs 10C, D</figureCitation>
|
||
;
|
||
<figureCitation id="131C2A49FFF2E412FC1EF9FEFC73F9F3" box="[934,1010,1576,1600]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="20" pageNumber="513">26D, H</figureCitation>
|
||
:
|
||
<quantity id="4CDF9B29FFF2E412FC41F9FEFBBCF9F3" box="[1017,1085,1576,1600]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="513" unit="mm" value="12.0">12 mm</quantity>
|
||
),
|
||
<collectionCode id="ED36AE09FFF2E412FBF0F9FFFB19F9F3" box="[1096,1176,1577,1600]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="513">MNHN</collectionCode>
|
||
.F.A90538,
|
||
<collectionCode id="ED36AE09FFF2E412FAB9F9FFFAD0F9F3" box="[1281,1361,1577,1600]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="513">MNHN</collectionCode>
|
||
.F.A90539 (
|
||
<taxonomicName id="4C274D4FFFF2E412FC8BF994FCECF9E9" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[819,877,1602,1626]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4F3C91FFF2E412FC0CF995FB13F9C6" ID-GBIF-Occurrence="4958671399" pageId="20" pageNumber="513">
|
||
1 spm; idem;
|
||
<collectionCode id="ED36AE09FFF2E412FB89F995FB05F9E9" box="[1073,1156,1603,1626]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="513">MNHN</collectionCode>
|
||
.F.R61443 (
|
||
<taxonomicName id="4C274D4FFFF2E412FB4BF994FAACF9E9" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1267,1325,1602,1626]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.), figured specimen of
|
||
<bibRefCitation id="EFB64B3DFFF2E412FC1AF98BFB98F9C6" author="MERLE D." box="[930,1049,1629,1653]" pageId="20" pageNumber="513" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">
|
||
<taxonomicName id="4C274D4FFFF2E412FC1AF98BFC5AF9C6" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[930,987,1629,1653]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
(1990
|
||
</bibRefCitation>
|
||
: pl. 4, fig. 3)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4F3C91FFF2E412FB1EF98BFC2CF919" ID-GBIF-Occurrence="4958671431" pageId="20" pageNumber="513">
|
||
1 spm; Paris Basin, Yvelines, Beynes (Ferme de l’Orme); middle Lutetian;
|
||
<collectionCode id="ED36AE09FFF2E412FB4EF9AEFACBF93C" box="[1270,1354,1656,1679]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="513">MNHN</collectionCode>
|
||
.F.A90535 (
|
||
<taxonomicName id="4C274D4FFFF2E412FC8BF944FCEFF919" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[819,878,1682,1706]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF2E412FC95F917FA2AF940" pageId="20" pageNumber="513" type="etymology">
|
||
<paragraph id="8B9836CCFFF2E412FC95F917FA2AF940" blockId="20.[811,1456,1729,1779]" pageId="20" pageNumber="513">
|
||
<smallCapsWord id="8D7EA010FFF2E412FC95F917FC27F964" baselines="1746,1747" box="[813,934,1729,1752]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="20" pageNumber="513">ETYMOLOGY</smallCapsWord>
|
||
. —
|
||
<emphasis id="B953EADEFFF2E412FC76F917FBA7F96A" box="[974,1062,1729,1753]" italics="true" pageId="20" pageNumber="513">Calciacus</emphasis>
|
||
(Latin adjective): from the ancient name of the village of Chaussy given in 690. Name given in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF2E412FC95F8DDFBEBF835" pageId="20" pageNumber="513" type="materials_examined">
|
||
<paragraph id="8B9836CCFFF2E412FC95F8DDFACDF88F" blockId="20.[813,1455,1802,1853]" pageId="20" pageNumber="513">
|
||
<smallCapsWord id="8D7EA010FFF2E412FC95F8DDFCDCF893" baselines="1820,1820" box="[813,861,1803,1826]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="20" pageNumber="513">TYPE</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF2E412FCDCF8C6FC3EF892" baselines="1821" box="[868,959,1808,1825]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="20" pageNumber="513">HORIZON</smallCapsWord>
|
||
. — Falun de Chaussy (biozone NP15), see
|
||
<bibRefCitation id="EFB64B3DFFF2E412FADEF8DCFC30F88E" author="GELY J. - P. & LORENZ C." pageId="20" pageNumber="513" pagination="713 - 747" refId="ref44561" refString="GELY J. - P. & LORENZ C. 1991. - Analyse sequentielle de l'Eocene et de l'Oligocene du Bassin parisien (France). Revue de l'institut francais du petrole 46 (6): 713 - 747." type="journal article" year="1991">Gély & Lorenz (1991</bibRefCitation>
|
||
: pl. 1), middle Eocene (middle Lutetian).
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF2E412FC95F882FBEBF835" blockId="20.[813,1455,1875,1926]" pageId="20" pageNumber="513">
|
||
<smallCapsWord id="8D7EA010FFF2E412FC95F882FCDCF8DA" baselines="1893,1893" box="[813,861,1876,1899]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="20" pageNumber="513">TYPE</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFF2E412FCDDF88FFC41F8D9" baselines="1894" box="[869,960,1881,1898]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="20" pageNumber="513">LOCALITY</smallCapsWord>
|
||
. —
|
||
<collectingCountry id="F330765CFFF2E412FC4DF882FBB8F8D8" box="[1013,1081,1876,1899]" name="France" pageId="20" pageNumber="513">France</collectingCountry>
|
||
, Paris Basin, Val d’Oise, Chaussy (Les Garennes), see
|
||
<bibRefCitation id="EFB64B3DFFF2E412FC78F8B8FBE7F835" author="FRITEL P. H." box="[960,1126,1902,1926]" pageId="20" pageNumber="513" refId="ref44321" refString="FRITEL P. H. 1910. - Guide geologique et paleontologique de la region parisienne dans un rayon de 100 kilometres. Deyrolle, Paris, 356 p." type="book" year="1910">Fritel (1910: 88)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFF2E412FC95F84BFBDEF859" pageId="20" pageNumber="513" type="distribution">
|
||
<paragraph id="8B9836CCFFF2E412FC95F84BFBDEF859" blockId="20.[812,1455,1949,2026]" pageId="20" pageNumber="513">
|
||
<smallCapsWord id="8D7EA010FFF2E412FC95F84BFC3BF800" baselines="1966,1967" box="[813,954,1949,1972]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="20" pageNumber="513">DISTRIBUTION</smallCapsWord>
|
||
. — Faluns de Chaussy and Calcaire à Orbitolites at La Ferme de l’Orme (biozone NP15), Paris Basin (
|
||
<collectingCountry id="F330765CFFF2E412FAACF861FADBF87C" box="[1300,1370,1975,1999]" name="France" pageId="20" pageNumber="513">France</collectingCountry>
|
||
), middle Eocene (middle Lutetian only).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF586644FFF3E413FF3CFCC5FA30FC88" ID-DOI="http://doi.org/10.5281/zenodo.13985782" ID-Zenodo-Dep="13985782" httpUri="https://zenodo.org/record/13985782/files/figure.png" pageId="21" pageNumber="514" startId="21.[132,143,787,804]" targetBox="[132,1455,214,744]" targetPageId="21" targetType="figure">
|
||
<paragraph id="8B9836CCFFF3E413FF3CFCC5FA30FC88" blockId="21.[132,1457,787,827]" pageId="21" pageNumber="514">
|
||
<smallCapsWord id="8D7EA010FFF3E413FF3CFCC5FF1CFC97" baselines="800,801" box="[132,157,787,804]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="21" pageNumber="514">FIG</smallCapsWord>
|
||
. 10. — Spiral cords and internal denticles of the outer lip in
|
||
<taxonomicName id="4C274D4FFFF3E413FDC4FCC5FC9AFC97" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[636,795,787,804]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="calciacus" status="sp. nov.">
|
||
<emphasis id="B953EADEFFF3E413FDC4FCC5FC9AFC97" box="[636,795,787,804]" italics="true" pageId="21" pageNumber="514">Timbellus calciacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF3E413FCA5FCC5FCCEFC97" box="[797,847,787,804]" pageId="21" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B953EADEFFF3E413FCECFCC5FCE0FC97" bold="true" box="[852,865,787,804]" pageId="21" pageNumber="514">A</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFF3E413FCD2FCC5FCF6FC97" bold="true" box="[874,887,787,804]" pageId="21" pageNumber="514">B</emphasis>
|
||
, holotype,MNHN.F.A90536 (
|
||
<taxonomicName id="4C274D4FFFF3E413FBDDFCC5FB15FC97" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1125,1172,787,804]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.),Chaussy (Les Garennes),Val d’Oise,Paris Basin,France,middle Eocene (Lutetian);
|
||
<emphasis id="B953EADEFFF3E413FD86FCFCFDCDFC88" bold="true" box="[574,588,810,827]" pageId="21" pageNumber="514">C</emphasis>
|
||
,
|
||
<emphasis id="B953EADEFFF3E413FDEDFCFCFDE2FC88" bold="true" box="[597,611,810,827]" pageId="21" pageNumber="514">D</emphasis>
|
||
, paratype,MNHN.F.A90537 (
|
||
<taxonomicName id="4C274D4FFFF3E413FCECFCFCFC02FC88" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[852,899,810,827]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
|
||
coll.), Chaussy (Les Garennes).Scale bars:3 mm.Credits:L.Cazes.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33D6547FFF3E413FF3CFC43FACFFB7D" pageId="21" pageNumber="514" type="description">
|
||
<paragraph id="8B9836CCFFF3E413FF3CFC43FE94FC1E" blockId="21.[130,777,917,1804]" box="[132,277,917,943]" pageId="21" pageNumber="514">
|
||
<smallCapsWord id="8D7EA010FFF3E413FF3CFC43FE94FC1E" baselines="936,936" box="[132,277,917,943]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="21" pageNumber="514">DESCRIPTION</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF3E413FF3CFC63FE1CF8BF" blockId="21.[130,777,917,1804]" pageId="21" pageNumber="514">
|
||
Bulbous protoconch of 1.2 whorls.Teleoconch up to
|
||
<quantity id="4CDF9B29FFF3E413FD01FC63FC86FC7C" box="[697,775,949,975]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="514" unit="mm" value="12.0">12 mm</quantity>
|
||
in height, up to
|
||
<quantity id="4CDF9B29FFF3E413FE80FC02FE0EFC5C" box="[312,399,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="21" pageNumber="514" unit="mm" value="5.5">5.5 mm</quantity>
|
||
in width, biconic in profile, composed of four whorls. Moderately high spire. Last whorl up to 76% of total length. Apical angle 71° including winged varices, up to 55° excluding them. Spiral sculpture with poorly marked primary cords. First whorl: appearance of very weak P1, P2 and P3. Second whorl: weak P1 to P3. Third whorl: appearance of weak s1, s2 and threads covering base of whorl. Fourth whorl: P1 to P6 weak, ADP on siphonal canal, secondary cords undistinguishable from threads; threads covering entire surface of shell. On first whorl, axial sculpture with 8-9 lamellose protovarices. On second whorl: appearance of varices (three varices and one low intervarix between two varices). From third to last whorl, numerous growing microlamellae covering surface of shell and forming delicate, reticulate sculpture covering surface of shell. Intervarical relief very low, but slightly higher on P1-P2. Varices well winged on last two whorls. Appearance of short P1 spine on second whorl. On last whorls, axial ornamentation forming winged varices, rounded in outline, without evidence of spines, even on P1. Subtriangular aperture up to 33% of diameter and up to 74% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly and forming a curved inductura at base. Parietal lip slightly adherent. Outer lip with very weak denticles including D1 to D5. Siphonal canal open, up to 45% of apertural length. Pseudo-umbilicus narrow.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF3E413FF3CF8E4FE9EF8F9" blockId="21.[132,776,1842,2028]" box="[132,287,1842,1868]" pageId="21" pageNumber="514">
|
||
<smallCapsWord id="8D7EA010FFF3E413FF3CF8E4FE9EF8F9" baselines="1861,1861" box="[132,287,1842,1868]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparisons" pageId="21" pageNumber="514">COMPARISONS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFF3E413FF3CF887FACFFB7D" blockId="21.[132,776,1842,2028]" lastBlockId="21.[811,1457,917,1230]" pageId="21" pageNumber="514">
|
||
This small species was never distinguished from its congeners in the Paris Basin, probably mistaken for juveniles of
|
||
<taxonomicName id="4C274D4FFFF3E413FF3EF844FE58F818" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[134,473,1937,1963]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF3E413FF3EF844FE96F818" box="[134,279,1937,1963]" italics="true" pageId="21" pageNumber="514">T. tripteroides</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFF3E413FE9DF847FE50F818" author="LAMARCK J. - B." box="[293,465,1937,1963]" pageId="21" pageNumber="514" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, with which it co-occurs at two localities: Chaussy and La Ferme de l’Orme. With a maximum height of
|
||
<quantity id="4CDF9B29FFF3E413FED9F807FE33F85F" box="[353,434,2001,2028]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="514" unit="mm" value="12.0">12 mm</quantity>
|
||
,
|
||
<taxonomicName id="4C274D4FFFF3E413FE01F804FDACF858" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[441,557,2001,2027]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="calciacus" status="n. sp.">
|
||
<emphasis id="B953EADEFFF3E413FE01F804FDACF858" box="[441,557,2001,2027]" italics="true" pageId="21" pageNumber="514">T. calciacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF3E413FD8DF804FDEFF85F" box="[565,622,2002,2028]" pageId="21" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
|
||
is a very small
|
||
<taxonomicName id="4C274D4FFFF3E413FC94FC43FC12FC1C" box="[812,915,917,943]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFF3E413FC94FC43FC12FC1C" box="[812,915,917,943]" italics="true" pageId="21" pageNumber="514">Timbellus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C274D4FFFF3E413FC19FC43FBE9FC1C" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[929,1128,917,943]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="calciacus" status="sp. nov.">
|
||
<emphasis id="B953EADEFFF3E413FC19FC43FBE9FC1C" box="[929,1128,917,943]" italics="true" pageId="21" pageNumber="514">Timbellus calciacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF3E413FBD7FC43FB28FC1C" box="[1135,1193,917,943]" pageId="21" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
|
||
differs from juveniles of
|
||
<taxonomicName id="4C274D4FFFF3E413FC95FC63FC3AFC7C" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[813,955,949,975]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF3E413FC95FC63FC3AFC7C" box="[813,955,949,975]" italics="true" pageId="21" pageNumber="514">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
in having the aperture completely formed and by having a peculiar reticulate microsculpture (see
|
||
<figureCitation id="131C2A49FFF3E413FAEFFC02FCC7FBBC" captionStart="FIG" captionStartId="16.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-382@16.[698,1267,772,1332]" captionTargetPageId="16" captionText="FIG. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides (Lamarck, 1822): A-D, morphotype with rounded varices and poorly distinguishable P1 spine; E-H, morphotype with easily distinguishable P1 spine: A, original figure of Lamarck (1805: pl. 3, fig. 4); B, C, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E, F, figured specimen of Deshayes (1835: pl. 82, fig. 1-2); G, H, figured specimen of Cossmann & Pissarro (1911: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart." figureDoi="http://doi.org/10.5281/zenodo.13985772" httpUri="https://zenodo.org/record/13985772/files/figure.png" pageId="21" pageNumber="514">Fig. 6C, D</figureCitation>
|
||
). This last character lacks in specimens of
|
||
<taxonomicName id="4C274D4FFFF3E413FAA7FC23FA30FBBD" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[1311,1457,1012,1038]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF3E413FAA7FC23FA30FBBD" box="[1311,1457,1012,1038]" italics="true" pageId="21" pageNumber="514">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
at any growth stage. Specimens of
|
||
<taxonomicName id="4C274D4FFFF3E413FB18FBC3FAB0FB9D" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1184,1329,1044,1070]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
|
||
<emphasis id="B953EADEFFF3E413FB18FBC3FAB0FB9D" box="[1184,1329,1044,1070]" italics="true" pageId="21" pageNumber="514">T. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF3E413FA81FBC3FAF5FB9C" box="[1337,1396,1045,1071]" pageId="21" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
|
||
from the upper Lutetian of
|
||
<collectingRegion id="49E3F82EFFF3E413FBA1FBE2FB11FBFD" box="[1049,1168,1076,1102]" country="France" name="Haute-Normandie" pageId="21" pageNumber="514">Normandy</collectingRegion>
|
||
and the early Bartonian of Loire-Atlantique share with
|
||
<taxonomicName id="4C274D4FFFF3E413FBDFFB83FB5EFBDD" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1127,1247,1108,1134]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="calciacus" status="n. sp.">
|
||
<emphasis id="B953EADEFFF3E413FBDFFB83FB5EFBDD" box="[1127,1247,1108,1134]" italics="true" pageId="21" pageNumber="514">T. calciacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF3E413FB50FB83FAA4FBDC" box="[1256,1317,1109,1135]" pageId="21" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
|
||
a small size, but
|
||
<taxonomicName id="4C274D4FFFF3E413FCE2FBA3FC6CFB3D" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[858,1005,1140,1166]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
|
||
<emphasis id="B953EADEFFF3E413FCE2FBA3FC6CFB3D" box="[858,1005,1140,1166]" italics="true" pageId="21" pageNumber="514">T. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF3E413FC4DFBA3FBB3FB3C" box="[1013,1074,1141,1167]" pageId="21" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
|
||
is double the size (H:
|
||
<quantity id="4CDF9B29FFF3E413FA94FBA2FA10FB3C" box="[1324,1425,1140,1167]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="21" pageNumber="514" unit="mm" value="26.6">26.6 mm</quantity>
|
||
vs
|
||
<quantity id="4CDF9B29FFF3E413FC96FB42FCFEFB1D" box="[814,895,1172,1198]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="21" pageNumber="514" unit="mm" value="12.0">12 mm</quantity>
|
||
). As with
|
||
<taxonomicName id="4C274D4FFFF3E413FC5CFB43FBF0FB1D" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[996,1137,1172,1198]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="tripteroides">
|
||
<emphasis id="B953EADEFFF3E413FC5CFB43FBF0FB1D" box="[996,1137,1172,1198]" italics="true" pageId="21" pageNumber="514">T. tripteroides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C274D4FFFF3E413FBC7FB43FA8EFB1D" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1151,1295,1172,1198]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
|
||
<emphasis id="B953EADEFFF3E413FBC7FB43FA8EFB1D" box="[1151,1295,1172,1198]" italics="true" pageId="21" pageNumber="514">T. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF3E413FAACFB42FACCFB1D" box="[1300,1357,1172,1198]" pageId="21" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
|
||
lacks the peculiar microsculpture found in
|
||
<taxonomicName id="4C274D4FFFF3E413FB2FFB63FA8DFB7D" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1175,1292,1204,1230]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="514" phylum="Mollusca" rank="species" species="calciacus" status="n. sp.">
|
||
<emphasis id="B953EADEFFF3E413FB2FFB63FA8DFB7D" box="[1175,1292,1204,1230]" italics="true" pageId="21" pageNumber="514">T. calciacus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFF3E413FAABFB62FACFFB7D" box="[1299,1358,1204,1230]" pageId="21" pageNumber="514" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |