246 lines
38 KiB
XML
246 lines
38 KiB
XML
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<mods:title id="B739A961ADEBA6AC7BA3CC6FE611C668">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:name id="93305C8CAEA0A375F754568F38B4E3AC" type="personal">
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<mods:namePart id="45114EEE2D795F5F4FF67F372BE10279">Merle, Didier</mods:namePart>
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<mods:namePart id="F6BD2A7133B615D73724DC211985F287">Pacaud, Jean-Michel</mods:namePart>
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<mods:namePart id="DC0BD32119D237CC8349E48A19D2FBEC">Ledon, Daniel</mods:namePart>
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<mods:affiliation id="61BC8BBBE4425EFF08BB6BB23F1416DB">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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<mods:nameIdentifier id="290E60E913CB9CD5767BE7D73C5CF175" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="7BE942E961AB6A5D6B60627451590AA7">Goret, Bernard</mods:namePart>
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<mods:affiliation id="DACD2EC4A87B9F6BD911515F42D745C7">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
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<mods:nameIdentifier id="6967D72D55ED92F12F2DDFF0BC45B8C7" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:title id="4D8BA560689543C9AA979249F7CD54BC">Geodiversitas</mods:title>
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<mods:date id="1CD7C0849354889A629AEF044F9CDDF9">2024</mods:date>
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<mods:number id="626293803DF1D49D3173192B7682FF7D">2024-10-24</mods:number>
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<mods:url id="C85FC0A68E47DD53608809BE6F76D654">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf</mods:url>
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<treatment id="038E87DAFFCCE42DFF56FA45FC84F9DE" ID-DOI="http://doi.org/10.5281/zenodo.13989292" ID-GBIF-Taxon="244735029" ID-Zenodo-Dep="13989292" LSID="urn:lsid:zoobank.org:act:A8FA7CDF-EBB1-4B5C-988E-14A31F6D1689" httpUri="http://treatment.plazi.org/id/038E87DAFFCCE42DFF56FA45FC84F9DE" lastPageId="43" lastPageNumber="535" pageId="42" pageNumber="535" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae" scope_subGenus="Pliocrassimurex">
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<subSubSection id="C33D6547FFCCE42CFF56FA45FD1FFA1D" box="[238,670,1427,1454]" pageId="42" pageNumber="535" type="nomenclature">
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<heading id="D0D081A0FFCCE42CFF56FA45FD1FFA1D" box="[238,670,1427,1454]" centered="true" fontSize="11" level="2" pageId="42" pageNumber="535" reason="2">
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Genus
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<taxonomicName id="4C274D4FFFCCE42CFE83FA45FDC9FA1E" ID-CoL="CN5M2" authority="Merle" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[315,584,1427,1453]" class="Gastropoda" family="Muricidae" genus="Pseudotrophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus" status="n. gen.">
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<emphasis id="B953EADEFFCCE42CFE83FA45FE7EFA1E" box="[315,511,1427,1453]" italics="true" pageId="42" pageNumber="535">Pseudotrophonopsis</emphasis>
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Merle
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26057A5FFCCE42CFDEBFA42FD1FFA1D" box="[595,670,1428,1454]" pageId="42" pageNumber="535" rank="genus">n. gen.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C33D6547FFCCE42CFE3EFA65FC86F9BA" pageId="42" pageNumber="535" type="description">
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<paragraph id="8B9836CCFFCCE42CFE3EFA65FD84FA7E" blockId="42.[238,670,1427,1485]" box="[390,517,1459,1485]" pageId="42" pageNumber="535">
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(
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<figureCitation id="131C2A49FFCCE42CFE37FA65FE5BFA7E" box="[399,474,1459,1485]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann & Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet & Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="42" pageNumber="535">Fig. 22</figureCitation>
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I-L)
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</paragraph>
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<paragraph id="8B9836CCFFCCE42CFF3CFA2EFC86F9BA" blockId="42.[132,775,1528,1545]" box="[132,775,1528,1545]" pageId="42" pageNumber="535">
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<uri id="FFB63ACEFFCCE42CFF3CFA2EFC86F9BA" box="[132,775,1528,1545]" pageId="42" pageNumber="535">
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urn:lsid:zoobank.org:act:
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<uuid id="FF810C19FFCCE42CFEE6FA2EFC86F9BA" box="[350,775,1528,1545]" pageId="42" pageNumber="535">A8FA7CDF-EBB1-4B5C-988E-14A31F6D1689</uuid>
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</uri>
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<subSubSection id="C33D6547FFCCE42CFF3CF9E2FDD0F933" pageId="42" pageNumber="535" type="materials_examined">
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<typeStatus id="549C886EFFCCE42CFF3CF9E2FF32F9FA" box="[132,179,1588,1611]" pageId="42" pageNumber="535">TYPE</typeStatus>
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<smallCapsWord id="8D7EA010FFCCE42CFF00F9EFFF7DF9F9" baselines="1606" box="[184,252,1593,1610]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="42" pageNumber="535">SPECIES</smallCapsWord>
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. —
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<taxonomicName id="4C274D4FFFCCE42CFE99F9E2FDF4F9F8" authority="Pilkington, 1804" authorityName="Pilkington" authorityYear="1804" box="[289,629,1587,1611]" class="Gastropoda" family="Buccinidae" genus="Buccinum" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="species" species="defossum">
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<emphasis id="B953EADEFFCCE42CFE99F9E2FE4EF9F8" box="[289,463,1587,1611]" italics="true" pageId="42" pageNumber="535">Buccinum defossum</emphasis>
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<bibRefCitation id="EFB64B3DFFCCE42CFE6DF9E5FDF4F9F8" author="PILKINGTON W." box="[469,629,1587,1611]" pageId="42" pageNumber="535" pagination="116 - 118" refId="ref48349" refString="PILKINGTON W. 1804. - Description of some fossil shells found in Hampshire. Transactions of the Linnean Society of London 7: 116 - 118." type="journal article" year="1804">Pilkington, 1804</bibRefCitation>
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</taxonomicName>
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, middle Eocene (Bartonian) of
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<collectingCountry id="F330765CFFCCE42CFEAAF998FEE5F9D5" box="[274,356,1614,1638]" name="United Kingdom" pageId="42" pageNumber="535">England</collectingCountry>
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(
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<collectingRegion id="49E3F82EFFCCE42CFEC8F998FE61F9D5" box="[368,480,1614,1638]" country="United Kingdom" name="Hampshire" pageId="42" pageNumber="535">Hampshire</collectingRegion>
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) (
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<figureCitation id="131C2A49FFCCE42CFE4EF998FDB8F9D5" box="[502,569,1614,1638]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann & Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet & Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="42" pageNumber="535">Fig. 22</figureCitation>
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I-K), and late Eocene (Priabonian) of
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<collectingCountry id="F330765CFFCCE42CFEA6F9BFFEDEF933" box="[286,351,1641,1664]" name="France" pageId="42" pageNumber="535">France</collectingCountry>
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(Paris Basin) (
|
||
<figureCitation id="131C2A49FFCCE42CFE57F9BEFDC4F933" box="[495,581,1640,1664]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann & Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet & Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="42" pageNumber="535">Fig. 22L</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33D6547FFCCE42CFF3CF944FED4F8EB" pageId="42" pageNumber="535" type="etymology">
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<paragraph id="8B9836CCFFCCE42CFF3CF944FD3CF96C" blockId="42.[130,776,1587,2027]" pageId="42" pageNumber="535">
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. — Name based on the Greek prefix
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<emphasis id="B953EADEFFCCE42CFDE7F944FD1CF919" box="[607,669,1682,1706]" italics="true" pageId="42" pageNumber="535">pseudo</emphasis>
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||
(false), because the superficial resemblance of the
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<typeStatus id="549C886EFFCCE42CFDBDF97BFDAEF976" box="[517,559,1709,1733]" pageId="42" pageNumber="535">type</typeStatus>
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species with members of the genus
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<taxonomicName id="4C274D4FFFCCE42CFEBAF911FD38F96C" authority="Bucquoy & Dautzenberg, 1882" authorityName="Bucquoy & Dautzenberg" authorityYear="1882" box="[258,697,1735,1759]" class="Gastropoda" family="Muricidae" genus="Trophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus">
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<emphasis id="B953EADEFFCCE42CFEBAF911FEFBF96C" box="[258,378,1735,1759]" italics="true" pageId="42" pageNumber="535">Trophonopsis</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFCCE42CFE38F911FD38F96C" author="BUCQUOY E. & DAUTZENBERG P. & DOLLFUS G." box="[384,697,1735,1759]" pageId="42" pageNumber="535" refId="ref43163" refString="BUCQUOY E., DAUTZENBERG P. & DOLLFUS G. 1882. - Les mollusques marins du Roussillon. Tome Ier. Gastropodes. Bailliere & fils, Paris, 570 p." type="book" year="1882">Bucquoy & Dautzenberg, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFCCE42CFF3CF927FED4F8EB" blockId="42.[130,776,1587,2027]" pageId="42" pageNumber="535">
|
||
<smallCapsWord id="8D7EA010FFCCE42CFF3CF927FF6AF8B4" baselines="1794,1795" box="[132,235,1777,1800]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Included" pageId="42" pageNumber="535">INCLUDED</smallCapsWord>
|
||
<smallCapsWord id="8D7EA010FFCCE42CFF48F920FEB6F8B4" baselines="1795" box="[240,311,1782,1799]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="42" pageNumber="535">SPECIES</smallCapsWord>
|
||
. —
|
||
<taxonomicName id="4C274D4FFFCCE42CFED9F927FD45F8BA" authority="Pilkington, 1804" authorityName="Pilkington" authorityYear="1804" box="[353,708,1776,1801]" class="Gastropoda" family="Buccinidae" genus="Buccinum" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="species" species="defossum">
|
||
<emphasis id="B953EADEFFCCE42CFED9F927FD99F8BB" box="[353,536,1776,1800]" italics="true" pageId="42" pageNumber="535">Buccinum defossum</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFCCE42CFDA6F926FD45F8BA" author="PILKINGTON W." box="[542,708,1776,1801]" pageId="42" pageNumber="535" pagination="116 - 118" refId="ref48349" refString="PILKINGTON W. 1804. - Description of some fossil shells found in Hampshire. Transactions of the Linnean Society of London 7: 116 - 118." type="journal article" year="1804">Pilkington, 1804</bibRefCitation>
|
||
</taxonomicName>
|
||
[synomym:
|
||
<taxonomicName id="4C274D4FFFCCE42CFF7EF8DAFDA7F890" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" box="[198,550,1803,1827]" class="Gastropoda" family="Muricidae" genus="Fusus" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="species" species="sublamellosus">
|
||
<emphasis id="B953EADEFFCCE42CFF7EF8DAFEFFF890" box="[198,382,1803,1827]" italics="true" pageId="42" pageNumber="535">Fusus sublamellosus</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFCCE42CFE3EF8DDFDA7F890" author="DESHAYES G. - P." box="[390,550,1803,1827]" pageId="42" pageNumber="535" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
|
||
</taxonomicName>
|
||
],
|
||
<collectingCountry id="F330765CFFCCE42CFD81F8DDFD0FF890" box="[569,654,1803,1827]" name="United Kingdom" pageId="42" pageNumber="535">England</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E3F82EFFCCE42CFD21F8DDFC86F890" box="[665,775,1803,1827]" country="United Kingdom" name="Hampshire" pageId="42" pageNumber="535">Hampshire</collectingRegion>
|
||
Basin, middle Eocene (Bartonian) and
|
||
<collectingCountry id="F330765CFFCCE42CFDAEF8F0FDDDF88E" box="[534,604,1830,1853]" name="France" pageId="42" pageNumber="535">France</collectingCountry>
|
||
, Paris Basin, late Eocene (Priabonian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFCCE42CFF3CF8BCFB1BFA71" pageId="42" pageNumber="535" type="diagnosis">
|
||
<paragraph id="8B9836CCFFCCE42CFF3CF8BCFB1BFA71" blockId="42.[130,776,1587,2027]" lastBlockId="42.[812,1456,1424,1474]" pageId="42" pageNumber="535">
|
||
<smallCapsWord id="8D7EA010FFCCE42CFF3CF8BCFF6EF833" baselines="1915,1916" box="[132,239,1898,1921]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="42" pageNumber="535">DIAGNOSIS</smallCapsWord>
|
||
. — Smooth multispiral protoconch (
|
||
<figureCitation id="131C2A49FFCCE42CFDEEF8BFFD2FF831" box="[598,686,1897,1922]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann & Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet & Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="42" pageNumber="535">Fig. 22K</figureCitation>
|
||
). Biconic teleoconch with rounded whorls. P6 not atrophied. IP, ADP, MP, ADP present. Primary P1 to P6 present on first whorl (
|
||
<figureCitation id="131C2A49FFCCE42CFD1AF848FD7DF804" box="[674,764,1950,1975]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann & Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet & Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="42" pageNumber="535">Fig. 22K</figureCitation>
|
||
). Coarse secondary cords equal in strength to primary cords. Varices low, formed by one short abaxial lamella. No P1 spine. No shoulder sinus. Posterior sinus shallow. Columellar lip smooth. Internal denticles D1 to D6 split. ID missing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33D6547FFCCE42DFC95FA25FC84F9DE" lastPageId="43" lastPageNumber="536" pageId="42" pageNumber="535" type="discussion">
|
||
<paragraph id="8B9836CCFFCCE42CFC95FA25FC2DF9B8" blockId="42.[811,1456,1523,2027]" box="[813,940,1523,1549]" pageId="42" pageNumber="535">
|
||
<smallCapsWord id="8D7EA010FFCCE42CFC95FA25FC2DF9B8" baselines="1542,1542" box="[813,940,1523,1549]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comments" pageId="42" pageNumber="535">COMMENTS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="8B9836CCFFCCE42DFC95F9C4FC84F9DE" blockId="42.[811,1456,1523,2027]" lastBlockId="43.[130,775,1427,1645]" lastPageId="43" lastPageNumber="536" pageId="42" pageNumber="535">
|
||
This strange species differs from members of
|
||
<taxonomicName id="4C274D4FFFCCE42CFAB8F9C4FCF1F9FE" authority="Merle, 2005 b" authorityName="Merle" authorityYear="2005" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFCCE42CFAB8F9C4FAE2F99F" box="[1280,1379,1554,1580]" italics="true" pageId="42" pageNumber="535">Jsowerbya</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFCCE42CFAD2F9C4FCF1F9FE" author="MERLE D." pageId="42" pageNumber="535" pagination="505 - 517" refId="ref46835" refString="MERLE D. 2005 b. - Jsowerbya, new genus of Muricidae (Mollusca Gastropoda) from the Eocene of the Paris (France) and Hampshire basins (England), with a phylogenetic assessment of its ocenebrine versus ergalataxine affinities. Geobios 38 (4): 505 - 517. https: // doi. org / 10.1016 / j. geobios. 2004.02.002" type="journal article" year="2005">Merle, 2005b</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C274D4FFFCCE42CFC19F9E5FBC7F9FF" authority="Merle" authorityName="Merle" authorityYear="2024" box="[929,1094,1586,1612]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||
<emphasis id="B953EADEFFCCE42CFC19F9E5FC7EF9FF" box="[929,1023,1587,1612]" italics="true" pageId="42" pageNumber="535">Beyregrex</emphasis>
|
||
Merle
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26057A5FFCCE42CFBF6F9E5FB17F9FE" box="[1102,1174,1587,1613]" pageId="42" pageNumber="535" rank="genus">n. gen.</taxonomicNameLabel>
|
||
by having rounded whorls, no shoulder sinus and by having strongly developed secondary cords. Because of its rounded whorls, it resembles members of
|
||
<taxonomicName id="4C274D4FFFCCE42CFCF1F944FC39F91F" authorityName="Merle" authorityYear="2005" box="[841,952,1682,1708]" class="Gastropoda" family="Muricidae" genus="Nucellopsis" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFCCE42CFCF1F944FC39F91F" box="[841,952,1682,1708]" italics="true" pageId="42" pageNumber="535">Nucellopsis</emphasis>
|
||
</taxonomicName>
|
||
, but they are distinguishing by their buccinoid shape and by a later appearance of the primary cords. The phylogenetic analysis (Winclada tree,
|
||
<figureCitation id="131C2A49FFCCE42CFB6BF904FA90F95F" box="[1235,1297,1746,1772]" captionStart="FIG" captionStartId="6.[134,143,1784,1801]" captionTargetBox="[134,618,235,1734]" captionTargetId="graphics-161@6.[134,618,235,1734]" captionTargetPageId="6" captionText="FIG. 1. — Strict Consensus tree of 7 equally parsimonious trees (length 107, ci 0.53, ri 0.78) generated by Winclada and computed using the option Heuristic Analysis (mult* max*), all characters non additive." figureDoi="http://doi.org/10.5281/zenodo.13985762" httpUri="https://zenodo.org/record/13985762/files/figure.png" pageId="42" pageNumber="535">Fig. 1</figureCitation>
|
||
) suggests that it could represent a stem group of
|
||
<taxonomicName id="4C274D4FFFCCE42CFB17F924FA92F8BF" authorityName="Merle" authorityYear="2005" box="[1199,1299,1778,1804]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFCCE42CFB17F924FA92F8BF" box="[1199,1299,1778,1804]" italics="true" pageId="42" pageNumber="535">Jsowerbya</emphasis>
|
||
</taxonomicName>
|
||
. We therefore propose the new genus
|
||
<taxonomicName id="4C274D4FFFCCE42CFB88F8C4FABAF89F" authority="Merle" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" box="[1072,1339,1810,1836]" class="Gastropoda" family="Muricidae" genus="Pseudotrophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus" status="n. gen.">
|
||
<emphasis id="B953EADEFFCCE42CFB88F8C4FB72F89F" box="[1072,1267,1810,1836]" italics="true" pageId="42" pageNumber="535">Pseudotrophonopsis</emphasis>
|
||
Merle
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26057A5FFCCE42CFAFFF8C4FA12F89F" box="[1351,1427,1810,1836]" pageId="42" pageNumber="535" rank="genus">n. gen.</taxonomicNameLabel>
|
||
as
|
||
<taxonomicName id="4C274D4FFFCCE42CFC96F8E4FBD2F8FF" authority="(Pilkington, 1804)" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" baseAuthorityName="Pilkington" baseAuthorityYear="1804" box="[814,1107,1841,1868]" class="Gastropoda" family="Muricidae" genus="Pseudotrophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="species" species="defossa" status="n. comb.">
|
||
<emphasis id="B953EADEFFCCE42CFC96F8E4FC09F8F8" box="[814,904,1841,1867]" italics="true" pageId="42" pageNumber="535">P. defossa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFCCE42CFC2BF8E7FBCBF8FF" author="PILKINGTON W." box="[915,1098,1841,1868]" pageId="42" pageNumber="535" pagination="116 - 118" refId="ref48349" refString="PILKINGTON W. 1804. - Description of some fossil shells found in Hampshire. Transactions of the Linnean Society of London 7: 116 - 118." type="journal article" year="1804">Pilkington, 1804</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26057A5FFCCE42CFBE4F8E4FB3BF8F8" box="[1116,1210,1841,1868]" pageId="42" pageNumber="535" rank="species">n. comb.</taxonomicNameLabel>
|
||
cannot be attributed to
|
||
<taxonomicName id="4C274D4FFFCCE42CFC93F887FC0FF8D8" authorityName="Merle" authorityYear="2005" box="[811,910,1873,1899]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B953EADEFFCCE42CFC93F887FC0FF8D8" box="[811,910,1873,1899]" italics="true" pageId="42" pageNumber="535">Jsowerbya</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C274D4FFFCCE42CFC15F884FBD2F8D8" authority="Merle" authorityName="Merle" authorityYear="2024" box="[941,1107,1873,1899]" class="Gastropoda" family="Muricidae" genus="Beyregrex" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="genus" status="gen. nov.">
|
||
<emphasis id="B953EADEFFCCE42CFC15F884FB8AF8D8" box="[941,1035,1874,1899]" italics="true" pageId="42" pageNumber="535">Beyregrex</emphasis>
|
||
Merle
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26057A5FFCCE42CFBE4F884FB22F8DF" box="[1116,1187,1874,1900]" pageId="42" pageNumber="535" rank="genus">n. gen.</taxonomicNameLabel>
|
||
<taxonomicName id="4C274D4FFFCCE42CFB1DF887FA2CF8D8" authorityName="Merle & Pacaud & Ledon & Goret" authorityYear="2024" baseAuthorityName="Pilkington" baseAuthorityYear="1804" box="[1189,1453,1873,1899]" class="Gastropoda" family="Muricidae" genus="Pseudotrophonopsis" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="species" species="defossa" status="comb. nov.">
|
||
<emphasis id="B953EADEFFCCE42CFB1DF887FA2CF8D8" box="[1189,1453,1873,1899]" italics="true" pageId="42" pageNumber="535">Pseudotrophonopsis defossa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26057A5FFCCE42CFC93F8A4FC08F838" box="[811,905,1905,1932]" pageId="42" pageNumber="535" rank="species">n. comb.</taxonomicNameLabel>
|
||
occurs in the Bartonian from
|
||
<collectingRegion id="49E3F82EFFCCE42CFB7EF8A7FABCF838" box="[1222,1341,1905,1931]" country="United Kingdom" name="Hampshire" pageId="42" pageNumber="535">Hampshire</collectingRegion>
|
||
(
|
||
<collectingCountry id="F330765CFFCCE42CFAF2F8A7FA27F838" box="[1354,1446,1905,1931]" name="United Kingdom" pageId="42" pageNumber="535">England</collectingCountry>
|
||
) and is known in the Priabonian from the Paris Basin, where it was identified as
|
||
<taxonomicName id="4C274D4FFFCCE42CFBB8F864FAFAF87F" authority="Deshayes, 1835" authorityName="Deshayes" authorityYear="1835" box="[1024,1403,1969,1996]" class="Gastropoda" family="Muricidae" genus="Fusus" kingdom="Animalia" order="Neogastropoda" pageId="42" pageNumber="535" phylum="Mollusca" rank="species" species="sublamellosus">
|
||
<emphasis id="B953EADEFFCCE42CFBB8F864FB46F878" box="[1024,1223,1969,1995]" italics="true" pageId="42" pageNumber="535">Fusus sublamellosus</emphasis>
|
||
<bibRefCitation id="EFB64B3DFFCCE42CFB76F867FAFAF87F" author="DESHAYES G. - P." box="[1230,1403,1969,1996]" pageId="42" pageNumber="535" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel & Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes, 1835</bibRefCitation>
|
||
</taxonomicName>
|
||
. The protoconch of the English and French populations (
|
||
<figureCitation id="131C2A49FFCCE42CFAF2F807FA26F858" box="[1354,1447,2001,2027]" captionStart="FIG" captionStartId="40.[132,143,1849,1866]" captionTargetBox="[166,1424,259,1693]" captionTargetId="figure-226@40.[703,1243,828,1398]" captionTargetPageId="40" captionText="FIG. 22. — Species of the informal group Jsowerbyia-Beyregrex n. gen.-Pseudotrophonopsis n. gen.: A, B, Jsowerbya sexdentata (Sowerby, 1823), BMNH GG22673 (Burton coll.), Colwell Bay, Wight Island, England, late Eocene (Priabonian); C, D, Beyregrex sarroniensis (Carez, 1870), n. comb., MNHN.F.J02576 (figured specimen of Cossmann & Pissarro 1911: pl. 35, fig.169-14), Sarron, Oise, Paris Basin, France, early Eocene (Ypresian); E, F, B. hantoniensis (Edwards, 1866),n. comb.,Morton coll., Middle Headon Brockenhurst bed, Whitecliff Bay, Wight Island, England,late Eocene (Priabonian); G, H, B. pereger (Beyrich, 1854), n. comb.,MNHN.F.A90561 (Lozouet & Maestrati coll.), Auvers-Saint-Georges,Essonne,Paris Basin,France, early Oligocene (Rupelian); I-L, Pseudotrophonopsis defossa (Pilkington, 1804), n. comb.: I, J, MNHN.F.J17634 (Cossmann coll.), Barton-on-Sea, Hampshire, England, middle Eocene (Bartonian); K, A. Smith coll. (juvenile specimen), upper Barton bed H, Barton Highcliffe; L, MNHN.F.A90560 (Munier-Chalmas coll.), Le Vouast, Oise, late Eocene (Priabonian). Scale bars: A-J, L, 5 mm; K, 1 mm. Credits: A, B, D. Serrette (MNHN); C, D, G-J, L, P. Loubry (MNHN/CNRS); E-F, K, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985815" httpUri="https://zenodo.org/record/13985815/files/figure.png" pageId="42" pageNumber="535">Fig. 22K</figureCitation>
|
||
) is similar and the teleoconch of the French population is only less thick (personal observation DM). Therefore,we believe that they belong to the same species.
|
||
<bibRefCitation id="EFB64B3DFFCDE42DFE63FA05FDF6FA5E" author="GLIBERT M." box="[475,631,1491,1517]" pageId="43" pageNumber="536" pagination="1 - 179" refId="ref44626" refString="GLIBERT M. 1963. - Les Muricacea et les Buccinacea du Cenozoique etranger des collections de l'Institut royal des Sciences naturelles de Belgique. Memoires de l'Institut royal des Sciences naturelles de Belgique serie 2, 74: 1 - 179." type="journal article" year="1963">Glibert (1963)</bibRefCitation>
|
||
recorded this species in the Bartonian from the Paris Basin, but we cannot confirm this record based on material in the examined collections (MNHN and private collections). It is possible that this species was mistaken for
|
||
<taxonomicName id="4C274D4FFFCDE42DFECCF984FC80F9DE" authority="(Deshayes, 1865)" baseAuthorityName="Deshayes" baseAuthorityYear="1865" box="[372,769,1618,1645]" class="Gastropoda" family="Muricidae" genus="Jsowerbya" kingdom="Animalia" order="Neogastropoda" pageId="43" pageNumber="536" phylum="Mollusca" rank="species" species="depauperata">
|
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<emphasis id="B953EADEFFCDE42DFECCF984FDCDF9DF" box="[372,588,1618,1644]" italics="true" pageId="43" pageNumber="536">Jsowerbya depauperata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB64B3DFFCDE42DFDEFF984FD7BF9DE" author="DESHAYES G. - P." box="[599,762,1618,1645]" pageId="43" pageNumber="536" pagination="101 - 107" refId="ref44001" refString="DESHAYES G. - P. 1865. - Description des Animaux sans vertebres decouverts dans le bassin de Paris pour servir de supplement a la description des coquilles fossiles des environs de Paris, comprenant une revue generale de toutes les especes actuellement connues. Tome 3. Bailliere, Paris, Livraisons 45 - 46, p. 201 - 288; Atlas 2, pl. 86 - 93 [2 January]; Livraisons 47 - 48, p. 289 - 424, Atlas 2, pl. 94 - 100 [10 June]; Livraisons 49 - 50, p. 425 - 668; Atlas 2, pl. 101 - 107 [15 December]." type="journal article" year="1865">Deshayes, 1865</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |