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<document id="AF1B09619D4DDE25D2412213B3E1364D" ID-DOI="10.5281/zenodo.3677235" ID-GBIF-Dataset="1ee19d5d-27cf-4c93-989c-3fe3eb467203" ID-ISSN="1942-1354" ID-Zenodo-Dep="3677235" ID-ZooBank="BA35690A-FC73-4E5A-A805-FE9550275FEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582218729488" checkinUser="felipe" docAuthor="Cong, Qian, Zhang, Jing, Shen, Jinhui &amp; Grishin, Nick V." docDate="2019" docId="038E29227E6CFFE0FF3F7110DBABFA0D" docLanguage="en" docName="InsectaMundi.0731.1-56.pdf.imf" docOrigin="Insecta Mundi 731 (731)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Tava Grishin 2019, new genus" docType="treatment" docUuid="A4C0707B-1BB7-410C-9682-7ACC79306988" docUuidSource="ZooBank" docVersion="11" lastPageNumber="29" masterDocId="FFB7515A7E72FFFEFFFF732ADF1EFFFB" masterDocTitle="Fifty new genera of Hesperiidae (Lepidoptera)" masterLastPageNumber="56" masterPageNumber="1" pageNumber="29" updateTime="1698720609646" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="80E2636403EF3225D4EF76ECC691E3F3">Fifty new genera of Hesperiidae (Lepidoptera)</mods:title>
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<taxonomicName id="4C27E3B77E6CFFE0FF3F7110DE69FDA9" authority="Grishin" authorityName="Grishin" authorityYear="2019" box="[192,375,570,594]" class="Insecta" family="Hesperiidae" genus="Tava" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95344267E6CFFE0FF3F7110DE1BFDAA" bold="true" box="[192,261,570,593]" italics="true" pageId="30" pageNumber="29">Tava</emphasis>
Grishin
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,
<taxonomicNameLabel id="A260F95D7E6CFFE0FE7A7110DD09FDA9" box="[389,535,570,594]" pageId="30" pageNumber="29" rank="genus">new genus</taxonomicNameLabel>
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http://zoobank.org/
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<typeStatus id="549C26967E6CFFE0FF3F7186DE1AFD3F" box="[192,260,684,708]" pageId="30" pageNumber="29">Type</typeStatus>
species.
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<taxonomicName id="4C27E3B77E6CFFE0FE817186DDCAFD3F" authority="Schaus, 1902" authorityName="Schaus" authorityYear="1902" box="[382,724,684,708]" class="Insecta" family="Cerambycidae" genus="Phanis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="tavola">
<emphasis id="B95344267E6CFFE0FE817186DD39FD3F" box="[382,551,684,708]" italics="true" pageId="30" pageNumber="29">Phanis tavola</emphasis>
Schaus, 1902
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.
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<emphasis id="B95344267E6CFFE0FF3F71F6DE4CFD0F" bold="true" box="[192,338,732,756]" pageId="30" pageNumber="29">Definition.</emphasis>
A genus without apparent close relatives, although in the same clade with
<taxonomicName id="4C27E3B77E6CFFE0FB2C71F6DE12FCEE" ID-CoL="93P6P" authority="Godman, [1901]" authorityName="Godman" authorityYear="1901" class="Insecta" family="Hesperiidae" genus="Phanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E6CFFE0FB2C71F6DA32FD0F" box="[1235,1324,732,756]" italics="true" pageId="30" pageNumber="29">Phanes</emphasis>
Godman, [1901]
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(
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species
<taxonomicName id="4C27E3B77E6CFFE0FE4C71D6DC0CFCEE" ID-CoL="94K6T" authority="Geyer, 1832" authorityName="Geyer" authorityYear="1832" box="[435,786,764,789]" class="Insecta" family="Hesperiidae" genus="Thracides" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="aletes">
<emphasis id="B95344267E6CFFE0FE4C71D6DD66FCEF" box="[435,632,764,788]" italics="true" pageId="30" pageNumber="29">Thracides aletes</emphasis>
Geyer, 1832
</taxonomicName>
), where it was placed previously but not monophyletic with it,
<taxonomicName id="4C27E3B77E6CFFE0FEDF7036DD01FCCF" ID-CoL="926CX" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[288,543,796,820]" class="Insecta" family="Hesperiidae" genus="Cumbre" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E6CFFE0FEDF7036DE9FFCCF" box="[288,385,796,820]" italics="true" pageId="30" pageNumber="29">Cumbre</emphasis>
<bibRefCitation id="EFB6E5C57E6CFFE0FE767037DD01FCCF" author="Evans" box="[393,543,796,820]" firstAuthor="Evans" pageId="30" pageNumber="29" pagination="54 - 88" refId="ref27591" refString="Evans, W. H. 1955. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History); London. v + 499 p., pl. 54 - 88" type="book chapter" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
(
<typeStatus id="549C26967E6CFFE0FDD17037DD7AFCCE" box="[558,612,797,821]" pageId="30" pageNumber="29">type</typeStatus>
species
<taxonomicName id="4C27E3B77E6CFFE0FD367036DB32FCCE" authority="Schaus, 1902" authorityName="Schaus" authorityYear="1902" box="[713,1068,796,821]" class="Insecta" family="Cerambycidae" genus="Phanis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="cumbre">
<emphasis id="B95344267E6CFFE0FD367036DC9EFCCF" box="[713,896,796,820]" italics="true" pageId="30" pageNumber="29">Phanis cumbre</emphasis>
Schaus, 1902
</taxonomicName>
),
<taxonomicName id="4C27E3B77E6CFFE0FBBF7037DA45FCCF" ID-CoL="92CVN" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" box="[1088,1371,796,820]" class="Insecta" family="Hesperiidae" genus="Lamponia" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E6CFFE0FBBF7037DBA3FCCF" box="[1088,1213,797,820]" italics="true" pageId="30" pageNumber="29">Lamponia</emphasis>
<bibRefCitation id="EFB6E5C57E6CFFE0FB3A7037DA45FCCF" author="Evans" box="[1221,1371,796,820]" firstAuthor="Evans" pageId="30" pageNumber="29" pagination="54 - 88" refId="ref27591" refString="Evans, W. H. 1955. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History); London. v + 499 p., pl. 54 - 88" type="book chapter" year="1955">Evans, 1955</bibRefCitation>
</taxonomicName>
(
<typeStatus id="549C26967E6CFFE0FA957037DABEFCCE" box="[1386,1440,797,821]" pageId="30" pageNumber="29">type</typeStatus>
species
<taxonomicName id="4C27E3B77E6CFFE0FEE57017DDA1FCAF" ID-CoL="92CVN" authority="Hewitson, 1876" authorityName="Hewitson" authorityYear="1876" box="[282,703,828,852]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="lamponia">
<emphasis id="B95344267E6CFFE0FEE57017DEE3FCAF" box="[282,509,828,852]" italics="true" pageId="30" pageNumber="29">Hesperia lamponia</emphasis>
Hewitson, 1876
</taxonomicName>
), and
<taxonomicName id="4C27E3B77E6CFFE0FCFB7016DB29FCAE" ID-CoL="92JRZ" authority="Godman, [1900]" authorityName="Godman" authorityYear="1900" box="[772,1079,828,853]" class="Insecta" family="Hesperiidae" genus="Pheraeus" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B95344267E6CFFE0FCFB7016DC6DFCAF" box="[772,883,828,852]" italics="true" pageId="30" pageNumber="29">Pheraeus</emphasis>
Godman, [1900]
</taxonomicName>
(
<typeStatus id="549C26967E6CFFE0FBBB7017DB67FCAE" box="[1092,1145,829,853]" pageId="30" pageNumber="29">type</typeStatus>
species
<taxonomicName id="4C27E3B77E6CFFE0FB277017DE76FC8E" ID-CoL="9C3XQ" authority="Mabille, 1891" authorityName="Mabille" authorityYear="1891" class="Insecta" family="Hesperiidae" genus="Carystus" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="epidius">
<emphasis id="B95344267E6CFFE0FB277017DA81FCAF" box="[1240,1439,828,852]" italics="true" pageId="30" pageNumber="29">Carystus epidius</emphasis>
Mabille, 1891
</taxonomicName>
) among others (
<figureCitation id="131C84B17E6CFFE0FDCD7077DDA9FC8E" box="[562,695,861,885]" captionStart-0="Figure 11" captionStart-1="Figure 12" captionStartId-0="29.[192,275,1330,1352]" captionStartId-1="31.[192,275,1330,1352]" captionTargetBox-0="[195,1438,214,1295]" captionTargetBox-1="[194,1440,214,1295]" captionTargetId-0="figure@29.[194,1442,213,1296]" captionTargetId-1="figure@31.[192,1440,213,1296]" captionTargetPageId-0="29" captionTargetPageId-1="31" captionText-0="Figure 11. Nuclear genome and Z chromosome trees of Moncina and relatives. See Fig. 2 legend for notations." captionText-1="Figure 12. Mitochondrial genome and COI barcode trees of Moncina and relatives. See Fig. 2 legend for notations." figureDoi-0="http://doi.org/10.5281/zenodo.3677257" figureDoi-1="http://doi.org/10.5281/zenodo.3677259" httpUri-0="https://zenodo.org/record/3677257/files/figure.png" httpUri-1="https://zenodo.org/record/3677259/files/figure.png" pageId="30" pageNumber="29">Fig. 1112</figureCitation>
). Keys to J.
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<bibRefCitation id="EFB6E5C57E6CFFE0FC5A7077DB55FC8F" author="Evans" box="[933,1099,860,884]" firstAuthor="Evans" pageId="30" pageNumber="29" pagination="54 - 88" refId="ref27591" refString="Evans, W. H. 1955. A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History); London. v + 499 p., pl. 54 - 88" type="book chapter" year="1955">Evans (1955)</bibRefCitation>
. Similar to many species in this group of relatives in having characteristic nearly square in many specimens, semi-hyaline spots near the bases of cells M
<subScript id="17A39A717E6CFFE0FE0E7086DEE4FC41" attach="left" box="[497,506,940,954]" fontSize="6" pageId="30" pageNumber="29">3</subScript>
-CuA
<subScript id="17A39A717E6CFFE0FDC67086DD5CFC41" attach="left" box="[569,578,940,954]" fontSize="6" pageId="30" pageNumber="29">1</subScript>
and CuA
<subScript id="17A39A717E6CFFE0FD487086DDDEFC41" attach="left" box="[695,704,940,954]" fontSize="6" pageId="30" pageNumber="29">1</subScript>
-CuA
<subScript id="17A39A717E6CFFE0FD007086DC16FC41" attach="left" box="[767,776,940,954]" fontSize="6" pageId="30" pageNumber="29">2</subScript>
on forewing. Differs from them by the combination of the following characters: antennae about 2/3 of costa length, nudum of 11 segments, palpi longer than wide in ventral view, 3rd segment short and conical, mid-tibiae with spines, forewing discal cell with well-developed recurrent vein, stigma V-shaped, inconspicuous, at the base of CuA
<subScript id="17A39A717E6CFFE0FB707726DB89FBE1" attach="left" box="[1167,1175,1036,1050]" fontSize="6" pageId="30" pageNumber="29">1</subScript>
-CuA
<subScript id="17A39A717E6CFFE0FB297726DBC1FBE1" attach="left" box="[1238,1247,1036,1050]" fontSize="6" pageId="30" pageNumber="29">2</subScript>
cell along veins, wings without apical spots, rufous brown below, hindwings mostly unspotted, in some specimens with a trace of small postdiscal yellowish spots (
<bibRefCitation id="EFB6E5C57E6CFFE0FD547717DC3FFBAE" author="Bell" box="[683,801,1085,1109]" firstAuthor="Bell" pageId="30" pageNumber="29" pagination="1 - 5" refId="ref26604" refString="Bell, E. L. 1940. A new genus and some new species of neotropical Hesperiidae (Lepidoptera, Rhopalocera). American Museum Novitates 1064: 1 - 5." type="journal article" year="1940">Bell 1940</bibRefCitation>
). Genitalia as figured by
<bibRefCitation id="EFB6E5C57E6CFFE0FBB67717DBD9FBAE" author="Bell" box="[1097,1223,1085,1109]" firstAuthor="Bell" pageId="30" pageNumber="29" pagination="1 - 5" refId="ref26604" refString="Bell, E. L. 1940. A new genus and some new species of neotropical Hesperiidae (Lepidoptera, Rhopalocera). American Museum Novitates 1064: 1 - 5." type="journal article" year="1940">Bell (1940</bibRefCitation>
: fig. 6), distinctive in the shape of harpe, rather straight, only slightly upturned and narrowing to a point, ampulla with a small tooth, penis wider than harpe, longer than valva, tegumen and uncus shorter than harpe. In DNA, a combination of the following base pairs is diagnostic: aly671.22.3:T167A, aly374.12.1:G1087C, aly2874.22.6:A1535G, aly671.22.3:A155T, aly420.67.1:A1631T.
</paragraph>
</subSubSection>
<subSubSection id="C33DCBBF7E6CFFE0FF3F77C7DC28FA7D" pageId="30" pageNumber="29" type="etymology">
<paragraph id="8B9898347E6CFFE0FF3F77C7DECBFADE" blockId="30.[192,1440,1261,1318]" pageId="30" pageNumber="29">
<emphasis id="B95344267E6CFFE0FF3F77C7DE45FAFE" bold="true" box="[192,347,1261,1285]" pageId="30" pageNumber="29">Etymology.</emphasis>
The name is a feminine noun in the nominative singular, derived from the beginning of the
<typeStatus id="549C26967E6CFFE0FF107624DE3AFADD" box="[239,292,1294,1318]" pageId="30" pageNumber="29">type</typeStatus>
species name.
</paragraph>
<paragraph id="8B9898347E6CFFE0FF3F7617DDDDFAAE" blockId="30.[192,707,1341,1366]" box="[192,707,1341,1366]" pageId="30" pageNumber="29">
<emphasis id="B95344267E6CFFE0FF3F7617DEAEFAAE" bold="true" box="[192,432,1341,1365]" pageId="30" pageNumber="29">Species included.</emphasis>
Only the
<typeStatus id="549C26967E6CFFE0FDD67614DD40FAAD" box="[553,606,1342,1366]" pageId="30" pageNumber="29">type</typeStatus>
species.
</paragraph>
<paragraph id="8B9898347E6CFFE0FF3F7647DC28FA7D" blockId="30.[192,822,1389,1414]" box="[192,822,1389,1414]" pageId="30" pageNumber="29">
<emphasis id="B95344267E6CFFE0FF3F7647DE62FA7E" bold="true" box="[192,380,1389,1413]" pageId="30" pageNumber="29">Parent taxon.</emphasis>
Subtribe Moncina A.
<bibRefCitation id="EFB6E5C57E6CFFE0FD787644DC2DFA7D" author="Warren, A. D. &amp; J. R. Ogawa &amp; A. V. Z. Brower" box="[647,819,1390,1414]" journalOrPublisher="Phylogenetic relationships of subfamilies and circumscription of tribes in the family Hesperiidae (Lepidoptera: Hesperioidea). Cladistics" pageId="30" pageNumber="29" pagination="642 - 676" part="24" refId="ref28580" refString="Warren, A. D., J. R. Ogawa, and A. V. Z. Brower. 2008. Phylogenetic relationships of subfamilies and circumscription of tribes in the family Hesperiidae (Lepidoptera: Hesperioidea). Cladistics 24 (5): 642 - 676." type="journal article" year="2008">Warren, 2008</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33DCBBF7E6CFFE0FF3F76B7DBABFA0D" pageId="30" pageNumber="29" type="discussion">
<paragraph id="8B9898347E6CFFE0FF3F76B7DBABFA0D" blockId="30.[192,1440,1437,1526]" pageId="30" pageNumber="29">
<emphasis id="B95344267E6CFFE0FF3F76B7DE47FA4E" bold="true" box="[192,345,1437,1461]" pageId="30" pageNumber="29">Comments.</emphasis>
We sequenced a
<typeStatus id="549C26967E6CFFE0FDDB76B4DD9AFA4D" box="[548,644,1438,1462]" pageId="30" pageNumber="29" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C27E3B77E6CFFE0FD5A76B7DC53FA4E" authorityName="Schaus" authorityYear="1902" box="[677,845,1437,1461]" class="Insecta" family="Cerambycidae" genus="Phanis" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="tavola">
<emphasis id="B95344267E6CFFE0FD5A76B7DC53FA4E" box="[677,845,1437,1461]" italics="true" pageId="30" pageNumber="29">Phanis tavola</emphasis>
</taxonomicName>
and the
<typeStatus id="549C26967E6CFFE0FC4C76B4DB05FA4D" box="[947,1051,1438,1462]" pageId="30" pageNumber="29" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C27E3B77E6CFFE0FBC376B7DA83FA4D" authority="Bell, 1940" authorityName="Bell" authorityYear="1940" box="[1084,1437,1437,1462]" class="Insecta" family="Hesperiidae" genus="Phanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="B95344267E6CFFE0FBC376B7DA04FA4E" box="[1084,1306,1437,1461]" italics="true" pageId="30" pageNumber="29">Phanes hoffmanni</emphasis>
<bibRefCitation id="EFB6E5C57E6CFFE0FADF76B4DA83FA4D" author="Bell" box="[1312,1437,1438,1462]" firstAuthor="Bell" pageId="30" pageNumber="29" pagination="1 - 5" refId="ref26604" refString="Bell, E. L. 1940. A new genus and some new species of neotropical Hesperiidae (Lepidoptera, Rhopalocera). American Museum Novitates 1064: 1 - 5." type="journal article" year="1940">Bell, 1940</bibRefCitation>
</taxonomicName>
. Analysis of these genomic sequences confirms the synonymy previously suggested for these two taxa (
<bibRefCitation id="EFB6E5C57E6CFFE0FF3776F4DE7DFA0D" author="Mielke" box="[200,355,1502,1526]" firstAuthor="Mielke" pageId="30" pageNumber="29" pagination="759 - 776" refId="ref27835" refString="Mielke, C. G. C. 1995. Papilionoidea e Hesperioidea (Lepidoptera) de Curitiba e seus arrededores, Parana, Brasil, com notas taxonomicas sobre Hesperiidae. Revista brasileira de Zoologia 11 (4): 759 - 776." type="journal article" year="1995">Mielke 1995</bibRefCitation>
). For instance, their COI barcodes differ by only one position (0.15%).
</paragraph>
</subSubSection>
</treatment>
</document>