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<document id="31A8E7EADF29443A632E432DB491B3B2" ID-DOI="10.11646/zootaxa.5040.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531538" ID-ZooBank="E862E329-68CC-4D90-862F-32FC75A36C30" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632749308219" checkinUser="plazi" docAuthor="Acosta, Luis E." docDate="2021" docId="038DB61AFF83BE0700D0F9A9F4FFFC38" docLanguage="en" docName="zootaxa.5040.3.7.pdf" docOrigin="Zootaxa 5040 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthopachylus Roewer 1913" docType="treatment" docVersion="3" lastPageNumber="438" masterDocId="FFB4CE62FF82BE050047FFD7F667FFF1" masterDocTitle="The return of a forgotten harvestman: revalidation of Gonyleptes robustus Holmberg, 1876, as the second species of Acanthopachylus Roewer, 1913 (Arachnida, Opiliones, Gonyleptidae)" masterLastPageNumber="438" masterPageNumber="428" pageNumber="428" updateTime="1699415529856" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="62E849B45BBA4653004C3A0EA39F9BD2">The return of a forgotten harvestman: revalidation of Gonyleptes robustus Holmberg, 1876, as the second species of Acanthopachylus Roewer, 1913 (Arachnida, Opiliones, Gonyleptidae)</mods:title>
</mods:titleInfo>
<mods:name id="410A609E3C0044B9AB058BA47A75E2E9" type="personal">
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<mods:roleTerm id="D557AC5E39544694D7D8B8FD5A62FCD6">Author</mods:roleTerm>
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<mods:namePart id="15DD08BA83F1F213CE7D2B3A70A11E26">Acosta, Luis E.</mods:namePart>
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<mods:typeOfResource id="5026A55077CA91B03EC902D0E54244DF">text</mods:typeOfResource>
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<mods:title id="23BD22398D8E2D44AFB3F12D8CED2C56">Zootaxa</mods:title>
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<mods:part id="5F55A0C7D17ADD54AD83D463AC30BE15">
<mods:date id="803BB005A91DF26ACE307A088491A23C">2021</mods:date>
<mods:detail id="1EBD8B1B22ADC948487DC03A009E38A2" type="pubDate">
<mods:number id="556A38C5647CAB4206602F3099BE43A2">2021-09-22</mods:number>
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<mods:detail id="F9D2B5F8BCBF5CB804B4F269ABF98AF7" type="volume">
<mods:number id="46FD83CE6217EE74015665E73C497458">5040</mods:number>
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<mods:detail id="2599218C0591775D21D133E84D6FE3CD" type="issue">
<mods:number id="FC207CDDDF38DF299EFE54D9AB7F2D17">3</mods:number>
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<mods:start id="8AD693D14095A7F74309C491B12FF5F7">428</mods:start>
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<mods:classification id="ADF5BC3B2425CBC572B809067238A214">journal article</mods:classification>
<mods:identifier id="C78A2CD09AD75B585B306414DBB76EC6" type="DOI">10.11646/zootaxa.5040.3.7</mods:identifier>
<mods:identifier id="B6CAA9BDD13AB19DC494B134BD988168" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1621204CBD2DC3942B718A038FC6EE6E" type="Zenodo-Dep">5531538</mods:identifier>
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<treatment id="038DB61AFF83BE0700D0F9A9F4FFFC38" ID-DOI="http://doi.org/10.5281/zenodo.5531548" ID-Zenodo-Dep="5531548" LSID="urn:lsid:plazi:treatment:038DB61AFF83BE0700D0F9A9F4FFFC38" httpUri="http://treatment.plazi.org/id/038DB61AFF83BE0700D0F9A9F4FFFC38" lastPageId="2" pageId="1">
<subSubSection id="C33E5487FF83BE0400D0F9A9F5B7F94C" pageId="1" type="nomenclature">
<paragraph id="8B9B070CFF83BE0400D0F9A9F47FF969" blockId="1.[151,976,1662,1725]" box="[151,536,1662,1689]" pageId="1">
<heading id="D0D3B060FF83BE0400D0F9A9F47FF969" bold="true" box="[151,536,1662,1689]" fontSize="11" level="1" pageId="1" reason="1">
<taxonomicName id="4C247C8FFF83BE0400D0F9A9F47FF969" ID-CoL="7NMM8" authority="Roewer, 1913" authorityName="Roewer" authorityYear="1913" box="[151,536,1662,1689]" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF83BE0400D0F9A9F47FF969" bold="true" box="[151,536,1662,1689]" pageId="1">
<emphasis id="B950DB1EFF83BE0400D0F9A9F70EF968" bold="true" box="[151,361,1662,1689]" italics="true" pageId="1">Acanthopachylus</emphasis>
<bibRefCitation id="EFB57AFDFF83BE040136F9A8F47FF969" author="Roewer, C. F." box="[369,536,1662,1689]" pageId="1" pagination="4 - 5" refId="ref7023" refString="Roewer, C. F. (1913) Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, 79 A (4 - 5), 1 - 472, Pl. Ia, b." type="journal article" year="1913">Roewer, 1913</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9B070CFF83BE0400D0F975F5B7F94C" blockId="1.[151,976,1662,1725]" box="[151,976,1698,1725]" pageId="1">
<uri id="FFB50B0EFF83BE0400D0F975F5B7F94C" box="[151,976,1698,1725]" pageId="1">
urn:lsid:zoobank.org:act:
<uuid id="FF823DD9FF83BE0401ECF975F5B7F94C" box="[427,976,1698,1725]" pageId="1">B64F3F55-8C88-4EBA-A6E6-AD95CE6088E6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33E5487FF83BE0400D0F930F29EF8AD" pageId="1" type="reference_group">
<paragraph id="8B9B070CFF83BE0400D0F930F52DF90E" blockId="1.[151,1273,1767,1884]" box="[151,842,1767,1791]" pageId="1">
<treatmentCitationGroup id="AB342022FF83BE0400D0F930F52DF90E" box="[151,842,1767,1791]" pageId="1">
<treatmentCitation id="0A85211DFF83BE0400D0F930F7D9F90E" author="Kirby, W." box="[151,446,1767,1791]" page="450" pageId="1" year="1819">
<taxonomicName id="4C247C8FFF83BE0400D0F930F7D9F90E" ID-CoL="7PHC5" authority=": Kirby 1819: 450" authorityName="Kirby" authorityPageNumber="450" authorityYear="1819" box="[151,446,1767,1791]" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF83BE0400D0F930F76EF90E" box="[151,265,1767,1791]" italics="true" pageId="1">Gonyleptes</emphasis>
:
<bibRefCitation id="EFB57AFDFF83BE040150F930F7D9F90E" author="Kirby, W." box="[279,446,1767,1791]" pageId="1" pagination="375 - 453" refId="ref6132" refString="Kirby, W. (1819) A century of Insects, including several new genera described from his cabinet. The Transactions of the Linnean Society of London, 12, 375 - 453, pls. XXI - XXII. https: // doi. org / 10.1111 / j. 1095 - 8339.1817. tb 00239. x" type="journal article" year="1819">Kirby 1819: 450</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part);
<treatmentCitation id="0A85211DFF83BE040263F930F48DF90E" author="Holmberg, E. L." box="[548,746,1767,1791]" page="28" pageId="1" year="1876">
<bibRefCitation id="EFB57AFDFF83BE040263F930F48DF90E" author="Holmberg, E. L." box="[548,746,1767,1791]" pageId="1" pagination="1 - 30" refId="ref6017" refString="Holmberg, E. L. (1876) Aracnidos argentinos. Anales de Agricultura de la Republica Argentina, 4, 1 - 30. [separatum]" type="journal article" year="1876">Holmberg 1876: 28</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B070CFF83BE0400D0F8D1F531F8EF" blockId="1.[151,1273,1767,1884]" box="[151,854,1798,1822]" pageId="1">
<treatmentCitationGroup id="AB342022FF83BE0400D0F8D1F531F8EF" box="[151,854,1798,1822]" pageId="1">
<treatmentCitation id="0A85211DFF83BE0400D0F8D1F7A0F8EF" author="Holmberg, E. L." box="[151,455,1798,1822]" page="70" pageId="1" year="1878">
<taxonomicName id="4C247C8FFF83BE0400D0F8D1F7A0F8EF" ID-CoL="7PRBX" authority=": Holmberg 1878: 70" authorityName="Holmberg" authorityPageNumber="70" authorityYear="1878" box="[151,455,1798,1822]" class="Arachnida" family="Gonyleptidae" genus="Pachylus" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF83BE0400D0F8D1F693F8EF" box="[151,244,1798,1822]" italics="true" pageId="1">Pachylus</emphasis>
:
<bibRefCitation id="EFB57AFDFF83BE040146F8D1F7A0F8EF" author="Holmberg, E. L." box="[257,455,1798,1822]" pageId="1" pagination="69 - 74" refId="ref6046" refString="Holmberg, E. L. (1878) Notas aracnologicas. Sobre los solpugidos argentinos. El Naturalista Argentino, 1 (3), 69 - 74." type="journal article" year="1878">Holmberg 1878: 70</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part);
<treatmentCitation id="0A85211DFF83BE04026AF8D1F490F8EF" author="Sorensen, W." box="[557,759,1798,1822]" page="639" pageId="1" year="1884">
<bibRefCitation id="EFB57AFDFF83BE04026AF8D1F490F8EF" author="Sorensen, W." box="[557,759,1798,1822]" pageId="1" pagination="555 - 646" refId="ref7266" refString="Sorensen, W. (1884) Opiliones Laniatores (Gonyleptides W. S. olim) Musei Hauniensis. Naturhistorisk Tidsskrift, Series 3, 14, 555 - 646." type="journal article" year="1884">Sørensen 1884: 639</bibRefCitation>
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B070CFF83BE0400D0F8F3F5BAF8CD" blockId="1.[151,1273,1767,1884]" box="[151,989,1828,1853]" pageId="1">
<treatmentCitationGroup id="AB342022FF83BE0400D0F8F3F5BAF8CD" box="[151,989,1828,1853]" pageId="1">
<treatmentCitation id="0A85211DFF83BE0400D0F8F3F78AF8CD" author="Roewer, C. F." box="[151,493,1828,1853]" page="50" pageId="1" year="1913">
<taxonomicName id="4C247C8FFF83BE0400D0F8F3F78AF8CD" authority="Roewer 1913: 50" authorityName="Roewer" authorityPageNumber="50" authorityYear="1913" box="[151,493,1828,1853]" class="Arachnida" family="Gonyleptidae" genus="Acanthpachylus" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF83BE0400D0F8F3F75EF8CD" box="[151,313,1828,1852]" italics="true" pageId="1">Acanthpachylus</emphasis>
<bibRefCitation id="EFB57AFDFF83BE040178F8F2F78AF8CD" author="Roewer, C. F." box="[319,493,1828,1853]" pageId="1" pagination="4 - 5" refId="ref7023" refString="Roewer, C. F. (1913) Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, 79 A (4 - 5), 1 - 472, Pl. Ia, b." type="journal article" year="1913">Roewer 1913: 50</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part). Incorrect original spelling (see below).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9B070CFF83BE0400D0F894F29EF8AD" blockId="1.[151,1273,1767,1884]" box="[151,1273,1859,1884]" pageId="1">
<treatmentCitationGroup id="AB342022FF83BE0400D0F894F29EF8AD" box="[151,1273,1859,1884]" pageId="1">
<treatmentCitation id="0A85211DFF83BE0400D0F894F46AF8AA" author="Roewer, C. F." box="[151,525,1859,1883]" page="411" pageId="1" year="1923">
<taxonomicName id="4C247C8FFF83BE0400D0F894F46AF8AA" ID-CoL="7NMM8" authority=": Roewer 1923: 411" authorityName=": Roewer" authorityPageNumber="411" authorityYear="1923" box="[151,525,1859,1883]" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF83BE0400D0F894F722F8AA" box="[151,325,1859,1883]" italics="true" pageId="1">Acanthopachylus</emphasis>
:
<bibRefCitation id="EFB57AFDFF83BE040114F893F46AF8AA" author="Roewer, C. F." box="[339,525,1859,1883]" pageId="1" refId="ref7061" refString="Roewer, C. F. (1923) Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. G. Fischer Verlag, Jena, 1116 pp." type="book" year="1923">Roewer 1923: 411</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(in part);
<treatmentCitation id="0A85211DFF83BE040234F894F525F8AA" author="Ringuelet, R. A." box="[627,834,1859,1883]" page="274" pageId="1" year="1959">
<bibRefCitation id="EFB57AFDFF83BE040234F894F525F8AA" author="Ringuelet, R. A." box="[627,834,1859,1883]" pageId="1" pagination="127 - 439" refId="ref6977" refString="Ringuelet, R. A. (1959) Los aracnidos argentinos del orden Opiliones. Revista del Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Cs. Zool., 5 (2), 127 - 439, pls. I - XX." type="journal article" year="1959">Ringuelet 1959: 274</bibRefCitation>
</treatmentCitation>
[synonymy];
<treatmentCitation id="0A85211DFF83BE040397F893F216F8AA" author="Kury, A. B." box="[976,1137,1859,1884]" page="153" pageId="1" year="2003">
<bibRefCitation id="EFB57AFDFF83BE040397F893F216F8AA" author="Kury, A. B." box="[976,1137,1859,1884]" pageId="1" pagination="5 - 337" refId="ref6264" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, Volumen especial monografico, 1, 5 - 337." type="journal article" year="2003">Kury 2003: 153</bibRefCitation>
</treatmentCitation>
[synonymy].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33E5487FF83BE0700D0F85EF2DBFDCC" lastPageId="2" pageId="1" type="type_taxon">
<paragraph id="8B9B070CFF83BE0400D0F85EF5A9F852" blockId="1.[151,1436,1929,2027]" box="[151,974,1929,1955]" pageId="1">
<emphasis id="B950DB1EFF83BE0400D0F85EF755F852" bold="true" box="[151,306,1929,1955]" pageId="1">
<typeStatus id="549FB9AEFF83BE0400D0F85EF6B5F852" box="[151,210,1929,1955]" pageId="1">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C247C8FFF83BE04017EF85EF4DEF852" ID-CoL="9WYSJ" authority="Kirby, 1819" authorityName="Kirby" authorityYear="1819" box="[313,697,1929,1955]" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="species" species="aculeatus">
<emphasis id="B950DB1EFF83BE04017EF85EF44EF852" box="[313,553,1929,1955]" italics="true" pageId="1">Gonyleptes aculeatus</emphasis>
<bibRefCitation id="EFB57AFDFF83BE040277F85EF4DEF852" author="Kirby, W." box="[560,697,1929,1955]" pageId="1" pagination="375 - 453" refId="ref6132" refString="Kirby, W. (1819) A century of Insects, including several new genera described from his cabinet. The Transactions of the Linnean Society of London, 12, 375 - 453, pls. XXI - XXII. https: // doi. org / 10.1111 / j. 1095 - 8339.1817. tb 00239. x" type="journal article" year="1819">Kirby, 1819</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8B9B070CFF83BE040080F87AF77EF81A" blockId="1.[151,1436,1929,2027]" pageId="1">
<emphasis id="B950DB1EFF83BE040080F87AF7F2F836" bold="true" box="[199,405,1965,1991]" pageId="1">Included species.</emphasis>
<taxonomicName id="4C247C8FFF83BE0401D8F87AF51DF836" ID-CoL="7QBMT" authority="(Kirby, 1819)" authorityName=": Roewer" authorityYear="1923" baseAuthorityName="Kirby" baseAuthorityYear="1819" box="[415,890,1965,1991]" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="species" species="aculeatus">
<emphasis id="B950DB1EFF83BE0401D8F87AF4B3F836" box="[415,724,1965,1991]" italics="true" pageId="1">Acanthopachylus aculeatus</emphasis>
(
<bibRefCitation id="EFB57AFDFF83BE0402A1F87AF515F836" author="Kirby, W." box="[742,882,1965,1991]" pageId="1" pagination="375 - 453" refId="ref6132" refString="Kirby, W. (1819) A century of Insects, including several new genera described from his cabinet. The Transactions of the Linnean Society of London, 12, 375 - 453, pls. XXI - XXII. https: // doi. org / 10.1111 / j. 1095 - 8339.1817. tb 00239. x" type="journal article" year="1819">Kirby, 1819</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C247C8FFF83BE0403F1F879F368F839" ID-CoL="7WCYD" authority="(Holmberg, 1876)" authorityName="Acosta" authorityYear="2021" baseAuthorityName="Holmberg" baseAuthorityYear="1876" box="[950,1295,1965,1992]" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="species" species="robustus" status="comb. nov.">
<emphasis id="B950DB1EFF83BE0403F1F879F25FF836" box="[950,1080,1965,1992]" italics="true" pageId="1">A. robustus</emphasis>
(
<bibRefCitation id="EFB57AFDFF83BE04040DF87AF361F839" author="Holmberg, E. L." box="[1098,1286,1965,1992]" pageId="1" pagination="1 - 30" refId="ref6017" refString="Holmberg, E. L. (1876) Aracnidos argentinos. Anales de Agricultura de la Republica Argentina, 4, 1 - 30. [separatum]" type="journal article" year="1876">Holmberg, 1876</bibRefCitation>
)
</taxonomicName>
<emphasis id="B950DB1EFF83BE04055EF87AF3FBF836" bold="true" box="[1305,1436,1965,1991]" pageId="1">
<taxonomicNameLabel id="A2636665FF83BE04055EF87AF3FBF836" box="[1305,1436,1965,1991]" pageId="1" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131F1B89FF83BE0400D8F806F76CF81A" box="[159,267,2001,2027]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1917,1941]" captionStartId-1="4.[151,250,1518,1542]" captionTargetBox-0="[153,1434,765,1889]" captionTargetBox-1="[163,1428,184,1466]" captionTargetId-0="figure-298@3.[151,1435,762,1890]" captionTargetId-1="figure-233@4.[151,1436,161,1486]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Live males of Acanthopachylus aculeatus (A–B: El Caracol, Departamento Rocha, Uruguay; photos by Javier A. Canteros), and Acanthopachylus robustus comb. nov. (C–D: Lago Lugano, Buenos Aires, Argentina; photos by Ricardo A. Palonsky—RAP).All photos used with permission of the authors." captionText-1="FIGURE 2. Diagnostic characters to distinguish male Acanthopachylus aculeatus (A–B) (MHNM, Departamento Lavalleja, Uruguay) and Acanthopachylus robustus comb. nov. (C–D) (LEA 000.428, Sierra de la Ventana, Buenos Aires Province, Argentina). A, C: dorsal scutum, coxae, trochanter and femur IV, dorsal view; B, D: coxa (with rear end of body), trochanter, femur and patella IV, prolateral view. Scale bars: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5531540" figureDoi-1="http://doi.org/10.5281/zenodo.5531542" httpUri-0="https://zenodo.org/record/5531540/files/figure.png" httpUri-1="https://zenodo.org/record/5531542/files/figure.png" pageId="1">Figs. 1–2</figureCitation>
).
</paragraph>
<paragraph id="8B9B070CFF80BE070080FF40F531FF08" blockId="2.[151,1437,150,969]" pageId="2">
<emphasis id="B950DB1EFF80BE070080FF40F782FF41" bold="true" box="[199,485,150,177]" pageId="2">Excluded combinations.</emphasis>
<taxonomicName id="4C247C8FFF80BE0701ACFF40F5D5FF40" ID-CoL="9X9QL" authority="(Thorell, 1877)" baseAuthorityName="Thorell" baseAuthorityYear="1877" box="[491,946,151,177]" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="butleri">
<emphasis id="B950DB1EFF80BE0701ACFF40F499FF40" box="[491,766,151,177]" italics="true" pageId="2">Acanthopachylus butleri</emphasis>
(
<bibRefCitation id="EFB57AFDFF80BE07034CFF40F5CDFF40" author="Thorell, T." box="[779,938,151,177]" pageId="2" pagination="201 - 218" refId="ref7298" refString="Thorell, T. (1877) Sobre algunos aracnidos de la Republica Argentina. Periodico Zoologico, 2 (4), 201 - 218." type="journal article" year="1877">Thorell, 1877</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="EFB57AFDFF80BE0703FBFF40F235FF40" author="Roewer, C. F." box="[956,1106,151,177]" pageId="2" pagination="4 - 5" refId="ref7023" refString="Roewer, C. F. (1913) Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, 79 A (4 - 5), 1 - 472, Pl. Ia, b." type="journal article" year="1913">Roewer 1913</bibRefCitation>
(nowadays assigned to genus
<taxonomicName id="4C247C8FFF80BE0700D0FF6CF7BEFF24" ID-CoL="7PRBS" authority="Muller, 1918" authorityName="Muller" authorityYear="1918" box="[151,473,187,213]" class="Arachnida" family="Gonyleptidae" genus="Pachyloidellus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF80BE0700D0FF6CF75AFF24" box="[151,317,187,213]" italics="true" pageId="2">Pachyloidellus</emphasis>
<bibRefCitation id="EFB57AFDFF80BE070103FF6CF7BEFF24" author="Muller, A." box="[324,473,187,213]" pageId="2" pagination="89 - 94" refId="ref6549" refString="Muller, A. (1918) Einige neue Gonyleptiden. Zoologischer Anzeiger, 49 (3 - 4), 89 - 94." type="journal article" year="1918">Müller, 1918</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="EFB57AFDFF80BE0701A6FF6CF416FF24" author="Acosta, L. E." box="[481,625,187,213]" pageId="2" pagination="1 - 18" refId="ref5303" refString="Acosta, L. E. (1993) El genero Pachyloidellus Muller, 1918 (Opiliones, Gonyleptidae, Pachylinae). Bonner zoologische Beitrage, 44 (1 - 2), 1 - 18." type="journal article" year="1993">Acosta 1993</bibRefCitation>
);
<taxonomicName id="4C247C8FFF80BE0702C2FF6CF23EFF24" authority="(Roewer, 1943)" baseAuthorityName="Roewer" baseAuthorityYear="1943" box="[645,1113,187,213]" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B950DB1EFF80BE0702C2FF6CF5C7FF25" box="[645,928,187,213]" italics="true" pageId="2">Acanthopachylus crassus</emphasis>
(
<bibRefCitation id="EFB57AFDFF80BE0703E9FF6CF237FF24" author="Roewer, C. F." box="[942,1104,187,213]" pageId="2" pagination="12 - 68" refId="ref7138" refString="Roewer, C. F. (1943) Uber Gonyleptiden. Weitere Weberknechte (Arachn., Opil.) XI. Senckenbergiana, 26 (1 - 3), 12 - 68." type="journal article" year="1943">Roewer, 1943</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="EFB57AFDFF80BE070424FF6CF6A8FF08" author="Capocasale, R. &amp; Bruno Trezza, L." pageId="2" pagination="19 - 32" refId="ref5751" refString="Capocasale, R. &amp; Bruno Trezza, L. (1964) Biologia de Acanthopachylus aculeatus (Kirby, 1819), (Opiliones; Pachylinae). Revista de la Sociedad Uruguaya de Entomologia, 6, 19 - 32." type="journal article" year="1964">Capocasale &amp; Bruno Trezza 1964</bibRefCitation>
(currently in
<taxonomicName id="4C247C8FFF80BE07012FFF08F40EFF08" ID-CoL="7PRBX" authority="Koch, 1839 a" authorityName="Koch" authorityYear="1839" box="[360,617,223,249]" class="Arachnida" family="Gonyleptidae" genus="Pachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF80BE07012FFF08F7A8FF08" box="[360,463,223,249]" italics="true" pageId="2">Pachylus</emphasis>
<bibRefCitation id="EFB57AFDFF80BE070191FF08F40EFF08" author="Koch, C. L." box="[470,617,223,249]" pageId="2" pagination="217 - 247" refId="ref6193" refString="Koch, C. L. (1839 a) Die Arachniden getreu nach der Natur abgebildet und beschrieben. Siebenter Band. C. H. Zeh'schen Buchhandlung, Nurnberg, 106 pp., pls. 217 - 247." type="book chapter" year="1839">Koch, 1839a</bibRefCitation>
</taxonomicName>
:
<bibRefCitation id="EFB57AFDFF80BE070235FF08F565FF08" author="Acosta, L. E." box="[626,770,223,249]" pageId="2" pagination="1 - 18" refId="ref5303" refString="Acosta, L. E. (1993) El genero Pachyloidellus Muller, 1918 (Opiliones, Gonyleptidae, Pachylinae). Bonner zoologische Beitrage, 44 (1 - 2), 1 - 18." type="journal article" year="1993">Acosta 1993</bibRefCitation>
,
<bibRefCitation id="EFB57AFDFF80BE07034AFF08F52EFF08" author="Acosta, L. E." box="[781,841,223,249]" pageId="2" pagination="134 - 147" refId="ref5373" refString="Acosta, L. E. (2021) The identity of an elusive Chilean harvestman, Pachylus crassus (Roewer, 1943) (Opiliones, Gonyleptidae, Pachylinae), with taxonomic and distribution notes. In: Jager, P., Schwendinger, P. &amp; Shear, B. (Eds.), A Festschrift honouring Prof. Dr. Jochen Martens on occasion of his 80 th birthday. Zootaxa, 4984 (1), pp. 134 - 147. https: // doi. org / 10.11646 / zootaxa. 4984.1.13" type="journal article" year="2021">2021</bibRefCitation>
).
</paragraph>
<paragraph id="8B9B070CFF80BE070080FED4F2DBFDCC" blockId="2.[151,1437,150,969]" pageId="2">
<emphasis id="B950DB1EFF80BE070080FED4F462FEED" bold="true" box="[199,517,258,285]" pageId="2">Note on the genus spelling.</emphasis>
In the original publication,
<bibRefCitation id="EFB57AFDFF80BE07037AFED4F581FEEC" author="Roewer, C. F." box="[829,998,259,285]" pageId="2" pagination="4 - 5" refId="ref7023" refString="Roewer, C. F. (1913) Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte, 79 A (4 - 5), 1 - 472, Pl. Ia, b." type="journal article" year="1913">Roewer (1913)</bibRefCitation>
consistently spelled the genus name as
<taxonomicName id="4C247C8FFF80BE0700D0FEF0F72DFEB0" authorityName="Roewer" authorityYear="1913" box="[151,330,295,321]" class="Arachnida" family="Gonyleptidae" genus="Acanthpachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF80BE0700D0FEF0F72DFEB0" box="[151,330,295,321]" italics="true" pageId="2">Acanthpachylus</emphasis>
</taxonomicName>
[
<collectingCountry id="F333479CFF80BE070110FEF0F7E6FEB0" box="[343,385,295,321]" name="New Zealand" pageId="2">NZ</collectingCountry>
, 1: 22]; 10 years later he emended it by inserting a missing
<emphasis id="B950DB1EFF80BE07045FFEF0F223FEB0" box="[1048,1092,295,321]" italics="true" pageId="2">“o”</emphasis>
, to get the current spelling
<taxonomicName id="4C247C8FFF80BE070532FEF0F75DFE94" ID-CoL="7NMM8" authorityName=": Roewer" authorityYear="1923" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF80BE070532FEF0F75DFE94" italics="true" pageId="2">Acanthopachylus</emphasis>
</taxonomicName>
. Although
<bibRefCitation id="EFB57AFDFF80BE0701F2FE9CF43BFE94" author="Roewer, C. F." box="[437,604,331,357]" pageId="2" refId="ref7061" refString="Roewer, C. F. (1923) Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. G. Fischer Verlag, Jena, 1116 pp." type="book" year="1923">Roewer (1923)</bibRefCitation>
does not explain the reasons for the change, according to ICZN Art. 33.2.1. this action is clearly intentional (
<emphasis id="B950DB1EFF80BE07024EFEB8F44BFE79" box="[521,556,367,392]" italics="true" pageId="2">i.e.</emphasis>
, the original and the modified spellings are both cited by
<bibRefCitation id="EFB57AFDFF80BE0704F0FEB8F336FE78" author="Roewer, C. F." box="[1207,1361,367,393]" pageId="2" refId="ref7061" refString="Roewer, C. F. (1923) Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. G. Fischer Verlag, Jena, 1116 pp." type="book" year="1923">Roewer 1923</bibRefCitation>
), what constitutes primarily an unjustified emendation [
<emphasis id="B950DB1EFF80BE0702F4FE44F4A8FE5C" box="[691,719,403,429]" italics="true" pageId="2">cf.</emphasis>
<collectingCountry id="F333479CFF80BE070293FE44F49BFE5C" box="[724,764,403,429]" name="New Zealand" pageId="2">NZ</collectingCountry>
, 1: 17]. Moreover, the original spelling
<taxonomicName id="4C247C8FFF80BE0704F4FE44F301FE5C" authorityName="Roewer" authorityYear="1913" box="[1203,1382,403,429]" class="Arachnida" family="Gonyleptidae" genus="Acanthpachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF80BE0704F4FE44F301FE5C" box="[1203,1382,403,429]" italics="true" pageId="2">Acanthpachylus</emphasis>
</taxonomicName>
cannot be interpreted as an inadvertent error, since this category is not met by the incorrect use of connecting vowels in forming a name (ICZN, Art. 32.5.1.). In any case, the emended spelling
<taxonomicName id="4C247C8FFF80BE07038FFE0CF2EEFE04" ID-CoL="7NMM8" authorityName=": Roewer" authorityYear="1923" box="[968,1161,475,501]" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF80BE07038FFE0CF2EEFE04" box="[968,1161,475,501]" italics="true" pageId="2">Acanthopachylus</emphasis>
</taxonomicName>
gained prevailing usage (
<emphasis id="B950DB1EFF80BE0700E6FE28F6DAFDE8" box="[161,189,511,537]" italics="true" pageId="2">cf.</emphasis>
synonymy in
<bibRefCitation id="EFB57AFDFF80BE07011DFE28F7B1FDE8" author="Kury, A. B." box="[346,470,511,537]" pageId="2" pagination="5 - 337" refId="ref6264" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, Volumen especial monografico, 1, 5 - 337." type="journal article" year="2003">Kury 2003</bibRefCitation>
), so it became by force of use a “justified emendation” (ICZN Art. 33.2.3.1.),
<emphasis id="B950DB1EFF80BE07057CFE28F33AFDE9" box="[1339,1373,511,536]" italics="true" pageId="2">i.e.</emphasis>
, with no separate authorship and date, rendering
<taxonomicName id="4C247C8FFF80BE07023FFDF4F54CFDCC" authorityName="Roewer" authorityYear="1913" box="[632,811,547,573]" class="Arachnida" family="Gonyleptidae" genus="Acanthpachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF80BE07023FFDF4F54CFDCC" box="[632,811,547,573]" italics="true" pageId="2">Acanthpachylus</emphasis>
</taxonomicName>
as the “incorrect original spelling”.
</paragraph>
</subSubSection>
<subSubSection id="C33E5487FF80BE070080FD91F4FFFC38" pageId="2" type="distribution">
<paragraph id="8B9B070CFF80BE070080FD91F4FFFC38" blockId="2.[151,1437,150,969]" pageId="2">
<emphasis id="B950DB1EFF80BE070080FD91F706FD91" bold="true" box="[199,353,582,608]" pageId="2">Distribution.</emphasis>
<taxonomicName id="4C247C8FFF80BE070120FD90F4FDFD90" authorityName=": Roewer" authorityYear="1923" baseAuthorityName="Kirby" baseAuthorityYear="1819" box="[359,666,583,609]" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="aculeatus">
<emphasis id="B950DB1EFF80BE070120FD90F4FDFD90" box="[359,666,583,609]" italics="true" pageId="2">Acanthopachylus aculeatus</emphasis>
</taxonomicName>
auct. has been deemed to be one of the most widespread harvestmen in South America. It was cited from Cayenne in the
<collectingCountry id="F333479CFF80BE0702AEFDBCF5E8FD74" box="[745,911,619,645]" name="French Guiana" pageId="2">French Guiana</collectingCountry>
(presumed “
<typeStatus id="549FB9AEFF80BE070467FDBCF207FD74" box="[1056,1120,619,645]" pageId="2">types</typeStatus>
” of
<taxonomicName id="4C247C8FFF80BE0704D6FDBCF3F2FD74" authorityName="Perty" authorityYear="1833" box="[1169,1429,619,645]" class="Arachnida" family="Gonyleptidae" genus="Gonyleptes" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="acanthurus">
<emphasis id="B950DB1EFF80BE0704D6FDBCF3F2FD74" box="[1169,1429,619,645]" italics="true" pageId="2">Gonyleptes acanthurus</emphasis>
</taxonomicName>
, according to
<bibRefCitation id="EFB57AFDFF80BE070160FD58F792FD58" author="Roewer, C. F." box="[295,501,655,681]" pageId="2" refId="ref7061" refString="Roewer, C. F. (1923) Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. G. Fischer Verlag, Jena, 1116 pp." type="book" year="1923">Roewer 1923: 413</bibRefCitation>
) up to the Strait of Magellan, in the southernmost tip of the continent (
<bibRefCitation id="EFB57AFDFF80BE0704B9FD58F6A4FD3C" author="Roewer, C. F." pageId="2" pagination="179 - 284" refId="ref7092" refString="Roewer, C. F. (1929) Weitere Weberknechte III. III. Erganzung der &quot; Weberknechte der Erde &quot;, 1923. Abhandlungen herausgegeben vom Naturwissenschaftlichen Verein zu Bremen, 27 (2), 179 - 284, pl. I." type="journal article" year="1929">Roewer 1929: 190</bibRefCitation>
,
<specimenCount id="9D22CC85FF80BE070089FD64F72AFD3F" box="[206,333,691,718]" pageId="2" type="female">one female</specimenCount>
).
<bibRefCitation id="EFB57AFDFF80BE070118FD64F424FD3C" author="Mello-Leitao, C. de" box="[351,579,691,717]" pageId="2" pagination="601 - 630" refId="ref6522" refString="Mello-Leitao, C. de (1939) Les arachnides et la zoogeographie de l'Argentine. Physis, 17 (49), 601 - 630." type="journal article" year="1939">Mello-Leitão (1939)</bibRefCitation>
argued that the species spreads over “toute l’Amérique du Sud, du coté atlantique”, an exaggerated assertion repeated by B. Soares &amp; H.
<bibRefCitation id="EFB57AFDFF80BE070305FD00F585FD00" author="Soares, B. A. M. &amp; Soares, H. E. M." box="[834,994,726,753]" pageId="2" pagination="225 - 302" refId="ref7174" refString="Soares, B. A. M. &amp; Soares, H. E. M. (1954) Monografia dos generos de opilioes neotropicos III. Arquivos de Zoologia do Estado de Sao Paulo, 8 (9), 225 - 302." type="journal article" year="1954">Soares (1954)</bibRefCitation>
.
<bibRefCitation id="EFB57AFDFF80BE0703A9FD00F2ADFD00" author="Eisner, T. &amp; Rossini, C. &amp; Gonzalez, A. &amp; Eisner, M." box="[1006,1226,727,753]" pageId="2" pagination="1313 - 1321" refId="ref5928" refString="Eisner, T., Rossini, C., Gonzalez, A. &amp; Eisner, M. (2004) Chemical defense of an opilionid (Acanthopachylus aculeatus). The Journal of Experimental Biology, 207, 1313 - 1321. https: // doi. org / 10.1242 / jeb. 00849" type="journal article" year="2004">
Eisner
<emphasis id="B950DB1EFF80BE07047AFD00F212FD00" box="[1085,1141,727,753]" italics="true" pageId="2">et al.</emphasis>
(2004)
</bibRefCitation>
, citing
<bibRefCitation id="EFB57AFDFF80BE07055AFD00F7C5FCE7" author="Capocasale, R. &amp; Bruno Trezza, L." pageId="2" pagination="19 - 32" refId="ref5751" refString="Capocasale, R. &amp; Bruno Trezza, L. (1964) Biologia de Acanthopachylus aculeatus (Kirby, 1819), (Opiliones; Pachylinae). Revista de la Sociedad Uruguaya de Entomologia, 6, 19 - 32." type="journal article" year="1964">Capocasale &amp; Bruno Trezza (1964)</bibRefCitation>
, still claimed that the species extends from the Guianas to
<collectingCountry id="F333479CFF80BE07047DFD2CF2CAFCE4" box="[1082,1197,763,789]" name="Argentina" pageId="2">Argentina</collectingCountry>
. The only well documented records of
<taxonomicName id="4C247C8FFF80BE07012EFCC8F44DFCC8" authorityName=": Roewer" authorityYear="1923" box="[361,554,799,825]" class="Arachnida" family="Gonyleptidae" genus="Acanthopachylus" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B950DB1EFF80BE07012EFCC8F44DFCC8" box="[361,554,799,825]" italics="true" pageId="2">Acanthopachylus</emphasis>
</taxonomicName>
, however, come from southern
<collectingCountry id="F333479CFF80BE0703C9FCC8F5B5FCC8" box="[910,978,799,825]" name="Brazil" pageId="2">Brazil</collectingCountry>
(
<collectingRegion id="49E0C9EEFF80BE0703A6FCC8F290FCC8" box="[993,1271,799,825]" country="Brazil" name="Rio Grande do Sul" pageId="2">Rio Grande do Sul State</collectingRegion>
),
<collectingCountry id="F333479CFF80BE07054CFCC8F30AFCC8" box="[1291,1389,799,825]" name="Uruguay" pageId="2">Uruguay</collectingCountry>
and
<collectingCountry id="F333479CFF80BE0700D0FC94F761FCAC" box="[151,262,835,861]" name="Argentina" pageId="2">Argentina</collectingCountry>
(
<bibRefCitation id="EFB57AFDFF80BE070153FC94F7A2FCAC" author="Ringuelet, R. A." box="[276,453,834,861]" pageId="2" pagination="127 - 439" refId="ref6977" refString="Ringuelet, R. A. (1959) Los aracnidos argentinos del orden Opiliones. Revista del Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Cs. Zool., 5 (2), 127 - 439, pls. I - XX." type="journal article" year="1959">Ringuelet 1959</bibRefCitation>
;
<bibRefCitation id="EFB57AFDFF80BE070197FC94F4F6FCAC" author="Capocasale, R." box="[464,657,835,861]" pageId="2" pagination="65 - 71" refId="ref5718" refString="Capocasale, R. (1968) Nuevos aportes para el conocimiento de la distribucion geografica de los opiliones de Uruguay. Neotropica, 14 (44), 65 - 71." type="journal article" year="1968">Capocasale 1968</bibRefCitation>
; H. Soares &amp; B.
<bibRefCitation id="EFB57AFDFF80BE07030CFC94F5B0FCAC" author="Soares, H. E. M. &amp; Soares, B. A. M." box="[843,983,835,861]" pageId="2" pagination="87 - 100" refId="ref7220" refString="Soares, H. E. M. &amp; Soares, B. A. M. (1986) Opera opiliologica varia. XXVI. (Opiliones, Gonyleptidae). Revista brasileira de Entomologia, 30 (1), 87 - 100." type="journal article" year="1986">Soares 1986</bibRefCitation>
;
<bibRefCitation id="EFB57AFDFF80BE0703A5FC94F238FCAC" author="Kury, A. B." box="[994,1119,835,861]" pageId="2" pagination="5 - 337" refId="ref6264" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, Volumen especial monografico, 1, 5 - 337." type="journal article" year="2003">Kury 2003</bibRefCitation>
). Despite this restriction, the genus distributes over an extensive range on grassland-type biomes (Pampas in
<collectingCountry id="F333479CFF80BE07044BFCB0F219FC70" box="[1036,1150,871,897]" name="Argentina" pageId="2">Argentina</collectingCountry>
, similar savanna environments in
<collectingCountry id="F333479CFF80BE0700BAFC5CF738FC54" box="[253,351,907,933]" name="Uruguay" pageId="2">Uruguay</collectingCountry>
and
<collectingCountry id="F333479CFF80BE0701D3FC5CF7BBFC54" box="[404,476,907,933]" name="Brazil" pageId="2">Brazil</collectingCountry>
;
<bibRefCitation id="EFB57AFDFF80BE0701A2FC5CF411FC54" author="Acosta, L. E." box="[485,630,907,933]" pageId="2" pagination="69 - 84" refId="ref5341" refString="Acosta, L. E. (2002) Patrones zoogeograficos de los opiliones argentinos (Arachnida: Opiliones). Revista Iberica de Aracnologia, 6, 69 - 84." type="journal article" year="2002">Acosta 2002</bibRefCitation>
), along with some peripheral records in neighboring ecoregions (
<figureCitation id="131F1B89FF80BE07050DFC5CF3E9FC54" box="[1354,1422,907,933]" captionStart="FIGURE 3" captionStartId="6.[151,250,1232,1256]" captionTargetBox="[151,1436,152,1209]" captionTargetId="figure-366@6.[151,1436,151,1209]" captionTargetPageId="6" captionText="FIGURE 3. Geographical distribution of Acanthopachylus aculeatus (Kirby) (blue dots) and A. robustus (Holmberg) comb. nov. (red dots) in Argentina, Uruguay and southern Brazil. Records with verified taxonomic identity are denoted with full color symbols, small white dots represent literature citations not checked as yet. Ecoregions according to Olson et al. (2001); both grassland-type ecoregions (Humid Pampas on the Argentinean side, Uruguayan Savanna in Uruguay and southern Brazil) are given the same color (light yellow). APAF: Alto Parana Atlantic Forests; HCH: Humid Chaco; MS: Southern Cone Mesopotamian Savanna; ESP: Espinal. Inset (bottom right): location of the area represented in South America." figureDoi="http://doi.org/10.5281/zenodo.5531544" httpUri="https://zenodo.org/record/5531544/files/figure.png" pageId="2">Fig. 3</figureCitation>
). Both species have marked synantropic habits.
</paragraph>
</subSubSection>
</treatment>
</document>
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