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<document id="20FC122B0154AC7C159B4744D999A5DB" ID-CLB-Dataset="5931" ID-DOI="10.11646/zootaxa.4981.1.1" ID-GBIF-Dataset="a6f5f409-8f2f-442a-915a-bbe8129d625a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4897374" ID-ZooBank="C98FB7A5-59AD-4D8D-9900-3EAFA339C537" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1622727723354" checkinUser="plazi" docAuthor="Barros, Lurdiana D., Barão, Kim R. &amp; Grazia, Jocelia" docDate="2021" docId="038D720BFF9BFFE6C2FCC3A4B7E9F869" docLanguage="en" docName="zootaxa.4981.1.1.pdf" docOrigin="Zootaxa 4981 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stysiana pardidens Grazia, Fernandes &amp; Schwertner 1999" docType="treatment" docVersion="10" lastPageNumber="33" masterDocId="FFB40A73FF84FFC6C26BC53AB279FFC0" masterDocTitle="Taxonomic updates on the Mecocephala group (Hemiptera: Pentatomidae) Redescription of ‘ Hypatropis complex’, with a review of genitalic terminology and new records" masterLastPageNumber="46" masterPageNumber="1" pageNumber="32" updateTime="1698999355226" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CA9ED838D3F66694C9788CDE46D7A864">Taxonomic updates on the Mecocephala group (Hemiptera: Pentatomidae) Redescription of ‘ Hypatropis complex’, with a review of genitalic terminology and new records</mods:title>
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<mods:namePart id="CF123696C930D6958CEE027986C6CB72">Barros, Lurdiana D.</mods:namePart>
<mods:affiliation id="643A0E778F3A0D79B02793960D062E63">Laboratory of Systematic Entomology, Department of Zoology, Institute of Biosciences, Campus do Vale, Federal University of Rio Grande do Sul, Avenue Bento Gonçalves, 9500, Porto Alegre 91501 - 970, Brazil.</mods:affiliation>
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<mods:namePart id="4C764310E8659498896B498D7E5AF9D2">Barão, Kim R.</mods:namePart>
<mods:affiliation id="1B393AD2A07F3996E19F846CFDC8603D">Laboratory of Systematics and Diversity of Arthropods, Penedo Educational Unit, Arapiraca Campus, Federal University of Alagoas, Avenue Beira Rio, s / n, Penedo 57200 - 000, Brazil.</mods:affiliation>
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<mods:namePart id="2B76E85321579EB13A97258F0A67F2E8">Grazia, Jocelia</mods:namePart>
<mods:affiliation id="BFF8A329335F3134713FC7341DD6FEC9">Laboratory of Systematic Entomology, Department of Zoology, Institute of Biosciences, Campus do Vale, Federal University of Rio Grande do Sul, Avenue Bento Gonçalves, 9500, Porto Alegre 91501 - 970, Brazil.</mods:affiliation>
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<treatment id="038D720BFF9BFFE6C2FCC3A4B7E9F869" ID-DOI="http://doi.org/10.5281/zenodo.5042738" ID-GBIF-Taxon="183217582" ID-Zenodo-Dep="5042738" LSID="urn:lsid:plazi:treatment:038D720BFF9BFFE6C2FCC3A4B7E9F869" httpUri="http://treatment.plazi.org/id/038D720BFF9BFFE6C2FCC3A4B7E9F869" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<subSubSection id="C33E9096FF9BFFD9C2FCC3A4B115F978" box="[151,876,1693,1720]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8B9BC31DFF9BFFD9C2FCC3A4B115F978" blockId="31.[151,876,1693,1757]" box="[151,876,1693,1720]" pageId="31" pageNumber="32">
<heading id="D0D37471FF9BFFD9C2FCC3A4B115F978" bold="true" box="[151,876,1693,1720]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<taxonomicName id="4C24B89EFF9BFFD9C2FCC3A4B115F978" authority="Grazia, Fernandes &amp; Schwertner, 1999" authorityName="Grazia, Fernandes &amp; Schwertner" authorityYear="1999" box="[151,876,1693,1720]" class="Insecta" family="Pentatomidae" genus="Stysiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pardidens">
<emphasis id="B9501F0FFF9BFFD9C2FCC3A4B115F978" bold="true" box="[151,876,1693,1720]" pageId="31" pageNumber="32">
<emphasis id="B9501F0FFF9BFFD9C2FCC3A4B305F978" bold="true" box="[151,380,1693,1720]" italics="true" pageId="31" pageNumber="32">Stysiana pardidens</emphasis>
<bibRefCitation id="EFB5BEECFF9BFFD9C3EFC3A7B115F978" author="Grazia, J. &amp; Fernandes, J. A. M. &amp; Schwertner, C. F." box="[388,876,1693,1720]" pageId="31" pageNumber="32" pagination="71 - 83" refId="ref23063" refString="Grazia, J., Fernandes, J. A. M. &amp; Schwertner, C. F. (1999) Stysiana, a new genus and four new species of Pentatomini (Heteroptera: Pentatomidae) of the Neotropical region. Acta Societatis Zoologicae Bohemicae, 63 (1 / 2), 71 - 83." type="journal article" year="1999">Grazia, Fernandes &amp; Schwertner, 1999</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C33E9096FF9BFFD9C2FCC3F8B00BF91C" box="[151,626,1730,1757]" pageId="31" pageNumber="32" type="description">
<paragraph id="8B9BC31DFF9BFFD9C2FCC3F8B00BF91C" blockId="31.[151,876,1693,1757]" box="[151,626,1730,1757]" pageId="31" pageNumber="32">
<figureCitation id="131FDF98FF9BFFD9C2FCC3F8B36EF91C" box="[151,279,1730,1756]" captionStart="FIGURES 13–20" captionStartId="3.[151,264,1922,1946]" captionTargetBox="[171,1420,319,1883]" captionTargetId="figure-83@3.[151,1436,309,1898]" captionTargetPageId="3" captionText="FIGURES 13–20. Head and pronotum in ventral view and habitus lateral. 13: Hypatropis inermis; 14: Hypatropis rolstoni; 15–16: Amauromelpia ussu; 17–18: Luridocimex wygodzinskyi; 19–20: Stysiana pardidens. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4897382" httpUri="https://zenodo.org/record/4897382/files/figure.png" pageId="31" pageNumber="32">Figs 19–20</figureCitation>
,
<figureCitation id="131FDF98FF9BFFD9C349C3F8B312F91C" box="[290,363,1730,1756]" captionStart="FIGURES 27–38" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[241,1346,204,1852]" captionTargetId="figure-22@6.[220,1367,181,1878]" captionTargetPageId="6" captionText="FIGURES 27–38. Male genitalia. Pygophore in dorsal, posterior and ventral view, respectively. 27–29: Hypatropis inermis; 30–32: Amauromelpia ussu; 33–35: Luridocimex wygodzinskyi: 36-38: Stysiana pardidens. Abbreviations: dr, dorsal rim; ilvr, inferior layer of ventral rim; slvr, superior layer of ventral rim; spp, superior process of pygophore; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4897390" httpUri="https://zenodo.org/record/4897390/files/figure.png" pageId="31" pageNumber="32">36–38</figureCitation>
,
<figureCitation id="131FDF98FF9BFFD9C31DC3F8B3EDF91D" box="[374,404,1730,1757]" captionStart="FIGURES 48–51" captionStartId="8.[151,264,1206,1230]" captionTargetBox="[175,1414,205,1164]" captionTargetId="figure-22@8.[151,1436,181,1182]" captionTargetPageId="8" captionText="FIGURES 48–51. Female genitalia. Genital plates in posteroventral view. 48: Hypatropis australis; 49: Amauromelpia miri; 50: Luridocimex wygodzinskyi: 51: Stysiana pardidens. Abbreviations: la8, laterotergites 8; la9, laterotergites 9; vf8, valvifers 8; vf9, valvifers 9. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4897396" httpUri="https://zenodo.org/record/4897396/files/figure.png" pageId="31" pageNumber="32">51</figureCitation>
,
<figureCitation id="131FDF98FF9BFFD9C3CBC3F8B3B5F91D" box="[416,460,1730,1757]" captionStart="FIGURES 143–146" captionStartId="29.[151,264,1935,1959]" captionTargetBox="[172,1407,193,1894]" captionTargetId="figure-19@29.[151,1436,181,1911]" captionTargetPageId="29" captionText="FIGURES 143–146. Stysiana Grazia, Fernandes &amp; Schwertner, 1999. Holotype, habitus dorsal. 143: S. acarinatis; 144: S. meridionalis; 145: S. pardidens; 146: S. tropicalis. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4897438" httpUri="https://zenodo.org/record/4897438/files/figure.png" pageId="31" pageNumber="32">145</figureCitation>
,
<figureCitation id="131FDF98FF9BFFD9C3B3C3F8B044F91C" box="[472,573,1730,1756]" captionStart="FIGURES 166–170" captionStartId="32.[151,264,1684,1708]" captionTargetBox="[174,1414,474,1640]" captionTargetId="figure-134@32.[151,1436,453,1661]" captionTargetPageId="32" captionText="FIGURES 166–170. Stysiana pardidens Grazia, Fernandes &amp; Schwertner, 1999. 166–167: Habitus dorsal and ventral; 168: Head in dorsal view; 169: Evaporatorium; 170: Female receptaculum seminis and aussenwand. Abbreviations: st, secondary thickenings of valvulae 9. Scale bars = 166–167, 170: 1.0 mm; 168–169: 0.5 mm. Ductus not preserved." figureDoi="http://doi.org/10.5281/zenodo.4897446" httpUri="https://zenodo.org/record/4897446/files/figure.png" pageId="31" pageNumber="32">166–170</figureCitation>
,
<figureCitation id="131FDF98FF9BFFD9C023C3F8B00BF91C" box="[584,626,1730,1756]" captionStart="FIGURE 184" captionStartId="35.[151,250,1449,1473]" captionTargetBox="[173,1414,545,1408]" captionTargetId="figure-171@35.[151,1436,525,1425]" captionTargetPageId="35" captionText="FIGURE 184. Occurrence records of the species of Stysiana, Luridocimex and Pedinonotus." figureDoi="http://doi.org/10.5281/zenodo.4897450" httpUri="https://zenodo.org/record/4897450/files/figure.png" pageId="31" pageNumber="32">184</figureCitation>
</paragraph>
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<paragraph id="8B9BC31DFF9BFFD9C2FCC23DB6B3F8DE" blockId="31.[151,1226,1798,1822]" box="[151,1226,1798,1822]" pageId="31" pageNumber="32">
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<treatmentCitation id="0A85E50CFF9BFFD9C2FCC23DB14EF8DE" author="Grazia, J. &amp; Fernandes, J. A. M. &amp; Schwertner, C. F." box="[151,823,1798,1822]" page="81" pageId="31" pageNumber="32" year="1999">
<taxonomicName id="4C24B89EFF9BFFD9C2FCC23DB14EF8DE" authority="Grazia, Fernandes &amp; Schwertner, 1999: 80 - 81" authorityName="Grazia, Fernandes &amp; Schwertner" authorityPageNumber="80 - 81" authorityYear="1999" box="[151,823,1798,1822]" class="Insecta" family="Pentatomidae" genus="Stysiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pardidens">
<emphasis id="B9501F0FFF9BFFD9C2FCC23DB32EF8DE" box="[151,343,1798,1822]" italics="true" pageId="31" pageNumber="32">Stysiana pardidens</emphasis>
<bibRefCitation id="EFB5BEECFF9BFFD9C335C23CB14EF8DE" author="Grazia, J. &amp; Fernandes, J. A. M. &amp; Schwertner, C. F." box="[350,823,1798,1822]" pageId="31" pageNumber="32" pagination="71 - 83" refId="ref23063" refString="Grazia, J., Fernandes, J. A. M. &amp; Schwertner, C. F. (1999) Stysiana, a new genus and four new species of Pentatomini (Heteroptera: Pentatomidae) of the Neotropical region. Acta Societatis Zoologicae Bohemicae, 63 (1 / 2), 71 - 83." type="journal article" year="1999">Grazia, Fernandes &amp; Schwertner, 1999: 80–81</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1, 7, 11, 15, 19, 22, 25, 28, 32, 36.
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</paragraph>
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<emphasis id="B9501F0FFF9BFFD9C2FCC277B3C6F8A6" bold="true" box="[151,447,1868,1895]" pageId="31" pageNumber="32">Type material examined.</emphasis>
<materialsCitation id="3B4CC940FF9BFFD9C3AEC276B0F7F84A" ID-GBIF-Occurrence="3311790355" collectingDate="1972-04-24" collectionCode="AMNH" collectorName="R. T &amp; J. C. Schuh" country="Peru" location="San Ramon de Pangoa" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549F7DBFFF9BFFD9C3AEC276B054F8A6" box="[453,557,1868,1894]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
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,
<collectingCountry id="F333838DFF9BFFD9C03CC277B0D8F8A7" box="[599,673,1869,1895]" name="Peru" pageId="31" pageNumber="32">PERU</collectingCountry>
, [Satipo],
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,
<quantity id="4CDC6EF8FF9BFFD9C675C277B61CF8A6" box="[1054,1125,1868,1895]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="32" unit="km" value="40.0">40 km</quantity>
SE,
<date id="FF9AE5DDFF9BFFD9C6F2C277B761F8A7" box="[1177,1304,1869,1895]" pageId="31" pageNumber="32" value="1972-04-24">
<collectingDate id="EFDE1C35FF9BFFD9C6F2C277B761F8A7" box="[1177,1304,1869,1895]" pageId="31" pageNumber="32" value="1972-04-24">24.IV.1972</collectingDate>
</date>
,
<collectorName id="26D1A6CBFF9BFFD9C748C277B2A4F84A" pageId="31" pageNumber="32">R.T &amp; J.C. Schuh</collectorName>
leg. (
<collectionCode id="ED355BD8FF9BFFD9C374C24BB301F84B" box="[287,376,1905,1931]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="31" pageNumber="32">AMNH</collectionCode>
) &lt;illustrated specimen&gt;
</materialsCitation>
.
<materialsCitation id="3B4CC940FF9BFFD9C0F1C24BB759F84B" ID-GBIF-Occurrence="3311790339" box="[666,1312,1904,1931]" collectionCode="UFRG" collectorName="Vicosa" country="Brazil" location="Minas Gerais" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<typeStatus id="549F7DBFFF9BFFD9C0F1C24BB172F84B" box="[666,779,1905,1931]" pageId="31" pageNumber="32" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F333838DFF9BFFD9C17CC24BB106F84B" box="[791,895,1905,1931]" name="Brazil" pageId="31" pageNumber="32">BRAZIL</collectingCountry>
,
<specimenCount id="9D220894FF9BFFD9C1E7C24BB1D8F84B" box="[908,929,1905,1931]" pageId="31" pageNumber="32" type="female">♀</specimenCount>
, [
<collectingRegion id="49E00DFFFF9BFFD9C1DCC24AB621F84B" box="[951,1112,1904,1931]" country="Brazil" name="Minas Gerais" pageId="31" pageNumber="32">Minas Gerais</collectingRegion>
],
<collectorName id="26D1A6CBFF9BFFD9C601C24AB6CFF84A" box="[1130,1206,1904,1930]" pageId="31" pageNumber="32">Viçosa</collectorName>
(
<collectionCode id="ED355BD8FF9BFFD9C6ACC24BB761F84B" box="[1223,1304,1905,1931]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="31" pageNumber="32">UFRG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4CC940FF9BFFD9C747C24BB13BF86F" ID-GBIF-Occurrence="3311790336" collectingDate="1941-12-09" collectionCode="UFRG" collectorName="Berla" country="Brazil" location="Pedra Branca" municipality="Parati" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<specimenCount id="9D220894FF9BFFD9C747C24BB738F84B" box="[1324,1345,1905,1931]" pageId="31" pageNumber="32" type="female">♀</specimenCount>
,
<collectingRegion id="49E00DFFFF9BFFD9C73AC24AB292F86E" country="Brazil" name="Rio de Janeiro" pageId="31" pageNumber="32">Rio de Janeiro</collectingRegion>
,
<collectingMunicipality id="6BFF5967FF9BFFD9C29EC2AEB340F86E" box="[245,313,1940,1966]" pageId="31" pageNumber="32">Parati</collectingMunicipality>
,
<location id="8EFB95C6FF9BFFD9C32FC2AEB3A5F86F" LSID="urn:lsid:plazi:treatment:038D720BFF9BFFE6C2FCC3A4B7E9F869:8EFB95C6FF9BFFD9C32FC2AEB3A5F86F" box="[324,476,1940,1967]" country="Brazil" municipality="Parati" name="Pedra Branca" pageId="31" pageNumber="32" stateProvince="Rio de Janeiro">Pedra Branca</location>
,
<date id="FF9AE5DDFF9BFFD9C38CC2AFB01DF86F" box="[487,612,1941,1967]" pageId="31" pageNumber="32" value="1941-12-09">
<collectingDate id="EFDE1C35FF9BFFD9C38CC2AFB01DF86F" box="[487,612,1941,1967]" pageId="31" pageNumber="32" value="1941-12-09">9.XII.1941</collectingDate>
</date>
,
<collectorName id="26D1A6CBFF9BFFD9C004C2AEB0D5F86E" box="[623,684,1940,1966]" pageId="31" pageNumber="32">Berla</collectorName>
leg. (
<collectionCode id="ED355BD8FF9BFFD9C082C2AFB143F86F" box="[745,826,1941,1967]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="31" pageNumber="32">UFRG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4CC940FF9BFFD9C127C2AFB313F813" ID-GBIF-Occurrence="3311790320" collectingDate="1976-01" collectionCode="UFRG" collectorName="Fritz" country="Brazil" location="Nova Teutonia" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="9D220894FF9BFFD9C127C2AFB11AF86F" box="[844,867,1941,1967]" pageId="31" pageNumber="32" type="male">♂</specimenCount>
,
<collectingRegion id="49E00DFFFF9BFFD9C105C2AFB66CF86E" box="[878,1045,1940,1967]" country="Brazil" name="Santa Catarina" pageId="31" pageNumber="32">Santa Catarina</collectingRegion>
,
<location id="8EFB95C6FF9BFFD9C674C2AFB6BFF86E" LSID="urn:lsid:plazi:treatment:038D720BFF9BFFE6C2FCC3A4B7E9F869:8EFB95C6FF9BFFD9C674C2AFB6BFF86E" box="[1055,1222,1940,1967]" country="Brazil" name="Nova Teutonia" pageId="31" pageNumber="32" stateProvince="Santa Catarina">Nova Teutônia</location>
,
<date id="FF9AE5DDFF9BFFD9C6BBC2AEB762F86E" box="[1232,1307,1940,1966]" pageId="31" pageNumber="32" value="1976-01">
<collectingDate id="EFDE1C35FF9BFFD9C6BBC2AEB762F86E" box="[1232,1307,1940,1966]" pageId="31" pageNumber="32" value="1976-01">I.1976</collectingDate>
</date>
,
<collectorName id="26D1A6CBFF9BFFD9C74DC2AEB722F86E" box="[1318,1371,1940,1966]" pageId="31" pageNumber="32">Fritz</collectorName>
Plaumann leg. (
<collectionCode id="ED355BD8FF9BFFD9C37FC283B31BF813" box="[276,354,1977,2003]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="31" pageNumber="32">UFRG</collectionCode>
)
</materialsCitation>
&lt;illustrated specimen&gt;;
<materialsCitation id="3B4CC940FF9BFFD9C0EBC283B63CF813" ID-GBIF-Occurrence="3311790323" box="[640,1093,1976,2003]" collectingDate="1975-10" collectionCode="UFRG" collectorName="Fritz Plaumann" country="Brazil" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D220894FF9BFFD9C0EBC283B0ECF813" box="[640,661,1977,2003]" pageId="31" pageNumber="32" type="female">♀</specimenCount>
,
<date id="FF9AE5DDFF9BFFD9C0C8C282B081F813" box="[675,760,1976,2003]" pageId="31" pageNumber="32" value="1975-10">
<collectingDate id="EFDE1C35FF9BFFD9C0C8C282B081F813" box="[675,760,1976,2003]" pageId="31" pageNumber="32" value="1975-10">X.1975</collectingDate>
</date>
,
<collectorName id="26D1A6CBFF9BFFD9C16FC282B1C9F812" box="[772,944,1976,2002]" pageId="31" pageNumber="32">Fritz Plaumann</collectorName>
leg. (
<collectionCode id="ED355BD8FF9BFFD9C184C283B644F813" box="[1007,1085,1977,2003]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="31" pageNumber="32">UFRG</collectionCode>
)
</materialsCitation>
&lt;illustrated specimen&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C33E9096FF9BFFE6C2ACC2E6B758FF39" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8B9BC31DFF9BFFE6C2ACC2E6B758FF39" blockId="31.[151,1437,1868,2039]" lastBlockId="32.[151,1437,151,429]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="B9501F0FFF9BFFD9C2ACC2E6B33BF836" bold="true" box="[199,322,2012,2038]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
Mesial margins of mandibular plates apically convergent (
<figureCitation id="131FDF98FF9BFFD9C1B6C2E6B638F837" box="[989,1089,2012,2039]" captionStart="FIGURES 166–170" captionStartId="32.[151,264,1684,1708]" captionTargetBox="[174,1414,474,1640]" captionTargetId="figure-134@32.[151,1436,453,1661]" captionTargetPageId="32" captionText="FIGURES 166–170. Stysiana pardidens Grazia, Fernandes &amp; Schwertner, 1999. 166–167: Habitus dorsal and ventral; 168: Head in dorsal view; 169: Evaporatorium; 170: Female receptaculum seminis and aussenwand. Abbreviations: st, secondary thickenings of valvulae 9. Scale bars = 166–167, 170: 1.0 mm; 168–169: 0.5 mm. Ductus not preserved." figureDoi="http://doi.org/10.5281/zenodo.4897446" httpUri="https://zenodo.org/record/4897446/files/figure.png" pageId="31" pageNumber="32">Fig. 168</figureCitation>
). Labiomere 2 sub equal to labiomeres 3 and 4 combined (
<figureCitation id="131FDF98FFA4FFE6C3B4C5ADB04BFF71" box="[479,562,151,177]" captionStart="FIGURES 13–20" captionStartId="3.[151,264,1922,1946]" captionTargetBox="[171,1420,319,1883]" captionTargetId="figure-83@3.[151,1436,309,1898]" captionTargetPageId="3" captionText="FIGURES 13–20. Head and pronotum in ventral view and habitus lateral. 13: Hypatropis inermis; 14: Hypatropis rolstoni; 15–16: Amauromelpia ussu; 17–18: Luridocimex wygodzinskyi; 19–20: Stysiana pardidens. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4897382" httpUri="https://zenodo.org/record/4897382/files/figure.png" pageId="32" pageNumber="33">Fig. 19</figureCitation>
). Mesosternum carinate. Median furrow of the peritreme long, reaching more than half of peritreme. Gyrification of evaporatorium with high wrinkles (
<figureCitation id="131FDF98FFA4FFE6C1A3C581B653FF16" box="[968,1066,187,214]" captionStart="FIGURES 166–170" captionStartId="32.[151,264,1684,1708]" captionTargetBox="[174,1414,474,1640]" captionTargetId="figure-134@32.[151,1436,453,1661]" captionTargetPageId="32" captionText="FIGURES 166–170. Stysiana pardidens Grazia, Fernandes &amp; Schwertner, 1999. 166–167: Habitus dorsal and ventral; 168: Head in dorsal view; 169: Evaporatorium; 170: Female receptaculum seminis and aussenwand. Abbreviations: st, secondary thickenings of valvulae 9. Scale bars = 166–167, 170: 1.0 mm; 168–169: 0.5 mm. Ductus not preserved." figureDoi="http://doi.org/10.5281/zenodo.4897446" httpUri="https://zenodo.org/record/4897446/files/figure.png" pageId="32" pageNumber="33">Fig. 169</figureCitation>
). Anterolateral angles of sternites dark-brown. Sub lateral margins of sternites dark-brown (
<figureCitation id="131FDF98FFA4FFE6C177C5E5B10DFF39" box="[796,884,223,249]" captionStart="FIGURES 13–20" captionStartId="3.[151,264,1922,1946]" captionTargetBox="[171,1420,319,1883]" captionTargetId="figure-83@3.[151,1436,309,1898]" captionTargetPageId="3" captionText="FIGURES 13–20. Head and pronotum in ventral view and habitus lateral. 13: Hypatropis inermis; 14: Hypatropis rolstoni; 15–16: Amauromelpia ussu; 17–18: Luridocimex wygodzinskyi; 19–20: Stysiana pardidens. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4897382" httpUri="https://zenodo.org/record/4897382/files/figure.png" pageId="32" pageNumber="33">Figs 20</figureCitation>
,
<figureCitation id="131FDF98FFA4FFE6C114C5E5B1D4FF3A" box="[895,941,223,250]" captionStart="FIGURES 166–170" captionStartId="32.[151,264,1684,1708]" captionTargetBox="[174,1414,474,1640]" captionTargetId="figure-134@32.[151,1436,453,1661]" captionTargetPageId="32" captionText="FIGURES 166–170. Stysiana pardidens Grazia, Fernandes &amp; Schwertner, 1999. 166–167: Habitus dorsal and ventral; 168: Head in dorsal view; 169: Evaporatorium; 170: Female receptaculum seminis and aussenwand. Abbreviations: st, secondary thickenings of valvulae 9. Scale bars = 166–167, 170: 1.0 mm; 168–169: 0.5 mm. Ductus not preserved." figureDoi="http://doi.org/10.5281/zenodo.4897446" httpUri="https://zenodo.org/record/4897446/files/figure.png" pageId="32" pageNumber="33">167</figureCitation>
). Spiracles dark-brown (
<figureCitation id="131FDF98FFA4FFE6C6ABC5E5B76DFF39" box="[1216,1300,223,249]" captionStart="FIGURES 13–20" captionStartId="3.[151,264,1922,1946]" captionTargetBox="[171,1420,319,1883]" captionTargetId="figure-83@3.[151,1436,309,1898]" captionTargetPageId="3" captionText="FIGURES 13–20. Head and pronotum in ventral view and habitus lateral. 13: Hypatropis inermis; 14: Hypatropis rolstoni; 15–16: Amauromelpia ussu; 17–18: Luridocimex wygodzinskyi; 19–20: Stysiana pardidens. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4897382" httpUri="https://zenodo.org/record/4897382/files/figure.png" pageId="32" pageNumber="33">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E9096FFA4FFE6C2ACC439B7E9F869" pageId="32" pageNumber="33" type="description">
<paragraph id="8B9BC31DFFA4FFE6C2ACC439B074FEDD" blockId="32.[151,1437,151,429]" box="[199,525,259,285]" pageId="32" pageNumber="33">
<emphasis id="B9501F0FFFA4FFE6C2ACC439B307FEDD" bold="true" box="[199,382,259,285]" pageId="32" pageNumber="33">Measurements.</emphasis>
See
<tableCitation id="C6A6F6A6FFA4FFE6C3D8C439B07EFEDD" box="[435,519,259,285]" captionStart="TABLE 1" captionStartId="39.[151,239,491,517]" captionTargetPageId="39" captionText="TABLE 1. Morphometric parameters measured of complex Hypatropis species. Abbreviations: A1, length of antennomere 1; A2, length of antennomere 2; A3, length of antennomere 3; A4, length of antennomere 4; A5, length of antennomere 5; BW, body width; BL, body length; CL, length of clypeus; HL, head length; HLE, length of head at compound eye level; HW, head width; IOD, interocular distance; L1, length of labiomere 1; L2, length of labiomere 2; L3, length of labiomere 3; L4, length of labiomere 4; PL, pronotum length; PW, pronotum width; SL, scutellum length; SW, scutellum width." httpUri="http://table.plazi.org/id/DF5B9395FFA3FFE1C2FCC4D1B1DCFDB1" pageId="32" pageNumber="33" tableUuid="DF5B9395FFA3FFE1C2FCC4D1B1DCFDB1">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8B9BC31DFFA4FFE6C2ACC41DB14CFE6D" blockId="32.[151,1437,151,429]" pageId="32" pageNumber="33">
<emphasis id="B9501F0FFFA4FFE6C2ACC41DB30EFE80" bold="true" box="[199,375,294,321]" pageId="32" pageNumber="33">Male genitalia.</emphasis>
Genital cup occupying more than half of pygophore length. Middle region of dorsal rim notched (
<figureCitation id="131FDF98FFA4FFE6C2F4C471B35CFEA6" box="[159,293,331,358]" captionStart="FIGURES 27–38" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[241,1346,204,1852]" captionTargetId="figure-22@6.[220,1367,181,1878]" captionTargetPageId="6" captionText="FIGURES 27–38. Male genitalia. Pygophore in dorsal, posterior and ventral view, respectively. 27–29: Hypatropis inermis; 30–32: Amauromelpia ussu; 33–35: Luridocimex wygodzinskyi: 36-38: Stysiana pardidens. Abbreviations: dr, dorsal rim; ilvr, inferior layer of ventral rim; slvr, superior layer of ventral rim; spp, superior process of pygophore; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4897390" httpUri="https://zenodo.org/record/4897390/files/figure.png" pageId="32" pageNumber="33">Figs 36–38</figureCitation>
).
<emphasis id="B9501F0FFFA4FFE6C350C471B3E8FEA5" box="[315,401,331,357]" italics="true" pageId="32" pageNumber="33">Phallus</emphasis>
: Dorsal connectives longer, surpassing the distal half of
<emphasis id="B9501F0FFFA4FFE6C64DC471B6D3FEA5" box="[1062,1194,331,357]" italics="true" pageId="32" pageNumber="33">phallotheca</emphasis>
. Dorsal processes of
<emphasis id="B9501F0FFFA4FFE6C2FCC455B362FE49" box="[151,283,367,393]" italics="true" pageId="32" pageNumber="33">phallotheca</emphasis>
medially narrow. Conjunctiva with two pairs of processes, 1+1 acute, and 1+1 sub truncate apically (see
<bibRefCitation id="EFB5BEECFFA4FFE6C2FCC4A9B001FE6D" author="Grazia, J. &amp; Fernandes, J. A. M. &amp; Schwertner, C. F." box="[151,632,403,429]" pageId="32" pageNumber="33" pagination="71 - 83" refId="ref23063" refString="Grazia, J., Fernandes, J. A. M. &amp; Schwertner, C. F. (1999) Stysiana, a new genus and four new species of Pentatomini (Heteroptera: Pentatomidae) of the Neotropical region. Acta Societatis Zoologicae Bohemicae, 63 (1 / 2), 71 - 83." type="journal article" year="1999">Grazia, Fernandes &amp; Schwertner, 1999: 79</bibRefCitation>
, figs 23, 26, 29).
</paragraph>
<caption id="DF5B9395FFA4FFE6C2FCC3AEB6C1F934" ID-DOI="http://doi.org/10.5281/zenodo.4897446" ID-Zenodo-Dep="4897446" httpUri="https://zenodo.org/record/4897446/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,264,1684,1708]" targetBox="[174,1414,474,1640]" targetPageId="32">
<paragraph id="8B9BC31DFFA4FFE6C2FCC3AEB6C1F934" blockId="32.[151,1437,1684,1781]" pageId="32" pageNumber="33">
<emphasis id="B9501F0FFFA4FFE6C2FCC3AEB309F96C" bold="true" box="[151,368,1684,1708]" pageId="32" pageNumber="33">FIGURES 166–170.</emphasis>
<taxonomicName id="4C24B89EFFA4FFE6C313C3AFB1A3F96D" authority="Grazia, Fernandes &amp; Schwertner, 1999" authorityName="Grazia, Fernandes &amp; Schwertner" authorityYear="1999" box="[376,986,1684,1709]" class="Insecta" family="Pentatomidae" genus="Stysiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pardidens">
<emphasis id="B9501F0FFFA4FFE6C313C3AFB043F96C" box="[376,570,1684,1708]" italics="true" pageId="32" pageNumber="33">Stysiana pardidens</emphasis>
<bibRefCitation id="EFB5BEECFFA4FFE6C028C3AEB1A3F96D" author="Grazia, J. &amp; Fernandes, J. A. M. &amp; Schwertner, C. F." box="[579,986,1684,1709]" pageId="32" pageNumber="33" pagination="71 - 83" refId="ref23063" refString="Grazia, J., Fernandes, J. A. M. &amp; Schwertner, C. F. (1999) Stysiana, a new genus and four new species of Pentatomini (Heteroptera: Pentatomidae) of the Neotropical region. Acta Societatis Zoologicae Bohemicae, 63 (1 / 2), 71 - 83." type="journal article" year="1999">Grazia, Fernandes &amp; Schwertner, 1999</bibRefCitation>
</taxonomicName>
. 166–167: Habitus dorsal and ventral; 168: Head in dorsal view; 169: Evaporatorium; 170: Female receptaculum seminis and aussenwand.
<emphasis id="B9501F0FFFA4FFE6C604C382B687F910" bold="true" box="[1135,1278,1720,1744]" pageId="32" pageNumber="33">Abbreviation</emphasis>
s: st, secondary thickenings of valvulae 9. Scale bars = 166–167, 170: 1.0 mm; 168–169: 0.5 mm. Ductus not preserved.
</paragraph>
</caption>
<paragraph id="8B9BC31DFFA4FFE6C2ACC219B1FBF846" blockId="32.[151,1436,1826,1962]" pageId="32" pageNumber="33">
<emphasis id="B9501F0FFFA4FFE6C2ACC219B3EDF8FC" bold="true" box="[199,404,1826,1853]" pageId="32" pageNumber="33">Female genitalia.</emphasis>
Anterior margins of valvifers 9 concave. Posterior margins of laterotergites 8 straight (
<figureCitation id="131FDF98FFA4FFE6C701C219B2CCF8A1" captionStart="FIGURES 27–38" captionStartId="6.[151,264,1902,1926]" captionTargetBox="[241,1346,204,1852]" captionTargetId="figure-22@6.[220,1367,181,1878]" captionTargetPageId="6" captionText="FIGURES 27–38. Male genitalia. Pygophore in dorsal, posterior and ventral view, respectively. 27–29: Hypatropis inermis; 30–32: Amauromelpia ussu; 33–35: Luridocimex wygodzinskyi: 36-38: Stysiana pardidens. Abbreviations: dr, dorsal rim; ilvr, inferior layer of ventral rim; slvr, superior layer of ventral rim; spp, superior process of pygophore; vr, ventral rim. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4897390" httpUri="https://zenodo.org/record/4897390/files/figure.png" pageId="32" pageNumber="33">Figs 38</figureCitation>
,
<figureCitation id="131FDF98FFA4FFE6C2A8C27DB288F8A2" box="[195,241,1863,1890]" captionStart="FIGURES 166–170" captionStartId="32.[151,264,1684,1708]" captionTargetBox="[174,1414,474,1640]" captionTargetId="figure-134@32.[151,1436,453,1661]" captionTargetPageId="32" captionText="FIGURES 166–170. Stysiana pardidens Grazia, Fernandes &amp; Schwertner, 1999. 166–167: Habitus dorsal and ventral; 168: Head in dorsal view; 169: Evaporatorium; 170: Female receptaculum seminis and aussenwand. Abbreviations: st, secondary thickenings of valvulae 9. Scale bars = 166–167, 170: 1.0 mm; 168–169: 0.5 mm. Ductus not preserved." figureDoi="http://doi.org/10.5281/zenodo.4897446" httpUri="https://zenodo.org/record/4897446/files/figure.png" pageId="32" pageNumber="33">166</figureCitation>
). Proximal
<emphasis id="B9501F0FFFA4FFE6C31DC27DB03CF8A1" box="[374,581,1863,1889]" italics="true" pageId="32" pageNumber="33">ductus receptaculi</emphasis>
with equal diameter than the ductus inside vesicular area. Annular flanges divergent (see
<bibRefCitation id="EFB5BEECFFA4FFE6C357C251B165F845" author="Grazia, J. &amp; Fernandes, J. A. M. &amp; Schwertner, C. F." box="[316,796,1899,1925]" pageId="32" pageNumber="33" pagination="71 - 83" refId="ref23063" refString="Grazia, J., Fernandes, J. A. M. &amp; Schwertner, C. F. (1999) Stysiana, a new genus and four new species of Pentatomini (Heteroptera: Pentatomidae) of the Neotropical region. Acta Societatis Zoologicae Bohemicae, 63 (1 / 2), 71 - 83." type="journal article" year="1999">Grazia, Fernandes &amp; Schwertner, 1999: 81</bibRefCitation>
, fig. 36).
</paragraph>
<paragraph id="8B9BC31DFFA4FFE6C2ACC2B4B7E9F869" blockId="32.[151,1436,1826,1962]" box="[199,1424,1934,1962]" pageId="32" pageNumber="33">
<emphasis id="B9501F0FFFA4FFE6C2ACC2B4B318F868" bold="true" box="[199,353,1934,1960]" pageId="32" pageNumber="33">Distribution.</emphasis>
<collectingCountry id="F333838DFFA4FFE6C30CC2B5B3A0F869" box="[359,473,1935,1961]" name="Colombia" pageId="32" pageNumber="33">Colombia</collectingCountry>
;
<collectingCountry id="F333838DFFA4FFE6C38EC2B5B039F869" box="[485,576,1935,1961]" name="Guyana" pageId="32" pageNumber="33">Guyana</collectingCountry>
;
<collectingCountry id="F333838DFFA4FFE6C020C2B5B0F6F869" box="[587,655,1935,1961]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
(
<collectingRegion id="49E00DFFFFA4FFE6C0F5C2B5B141F869" box="[670,824,1935,1961]" country="Brazil" name="Minas Gerais" pageId="32" pageNumber="33">Minas Gerais</collectingRegion>
,
<collectingRegion id="49E00DFFFFA4FFE6C128C2B5B191F869" box="[835,1000,1935,1961]" country="Brazil" name="Rio de Janeiro" pageId="32" pageNumber="33">Rio de Janeiro</collectingRegion>
, and
<collectingRegion id="49E00DFFFFA4FFE6C648C2B5B6B5F86A" box="[1059,1228,1935,1962]" country="Brazil" name="Santa Catarina" pageId="32" pageNumber="33">Santa Catarina</collectingRegion>
);
<collectingCountry id="F333838DFFA4FFE6C6B4C2B5B76BF869" box="[1247,1298,1935,1961]" name="Peru" pageId="32" pageNumber="33">Peru</collectingCountry>
(
<figureCitation id="131FDF98FFA4FFE6C74AC2B5B7FDF869" box="[1313,1412,1935,1961]" captionStart="FIGURE 184" captionStartId="35.[151,250,1449,1473]" captionTargetBox="[173,1414,545,1408]" captionTargetId="figure-171@35.[151,1436,525,1425]" captionTargetPageId="35" captionText="FIGURE 184. Occurrence records of the species of Stysiana, Luridocimex and Pedinonotus." figureDoi="http://doi.org/10.5281/zenodo.4897450" httpUri="https://zenodo.org/record/4897450/files/figure.png" pageId="32" pageNumber="33">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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