645 lines
93 KiB
XML
645 lines
93 KiB
XML
<document id="5BB4EEF32A7EE0AB90AA61F5FD0769B0" ID-CLB-Dataset="45992" ID-DOI="10.11646/zootaxa.3786.4.6" ID-GBIF-Dataset="e8155e8f-5e1a-433d-b55b-bb7cf322f088" ID-ISSN="1175-5326" ID-Zenodo-Dep="229326" ID-ZooBank="EC1A7720-D2C4-4F90-BAFF-786CA6D49C50" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460409342319" checkinUser="plazi" docAuthor="Kluge, Nikita J., Tiunova, Tatiana M. & Novikova, Eugenia A." docDate="2014" docId="038D4E74FFCC30579F92FE1EE4A20923" docLanguage="en" docName="zt03786p491.pdf" docOrigin="Zootaxa 3786 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon monilistylus Kluge, Tiunova & Novikova, 2014, sp. n." docType="treatment" docVersion="8" lastPageNumber="490" masterDocId="FFB4360CFFCD30509F05FF8DE6060E53" masterDocTitle="A new species, Procloeon monilistylus sp. n. (Ephemeroptera, Baetidae), from the Russian Far East" masterLastPageNumber="491" masterPageNumber="483" pageNumber="484" updateTime="1698300259013" updateUser="ExternalLinkService">
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<mods:mods id="9E45C3551A953423888AD265EFDB422F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="28153A9D3C7D39842D2F9D7BD335C320">A new species, Procloeon monilistylus sp. n. (Ephemeroptera, Baetidae), from the Russian Far East</mods:title>
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<mods:name id="7F3810474C2C583F71672509E70475B0" type="personal">
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<mods:namePart id="FB2716851E6AC0BFC16DB6CDC2FB4167">Kluge, Nikita J.</mods:namePart>
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<mods:namePart id="C9D5B463BE1B270B8119407FF22B995B">Tiunova, Tatiana M.</mods:namePart>
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<mods:roleTerm id="145E305DE41A8080B906CDE7FA830392">Author</mods:roleTerm>
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<mods:namePart id="66FB2BB2CC7D1C8428A356B215B7B1DA">Novikova, Eugenia A.</mods:namePart>
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<mods:typeOfResource id="32EF6FFDC799883263E91B907EE2CB6C">text</mods:typeOfResource>
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<mods:title id="E4744F3928AAA0D09EA59A9B4ACB2D49">Zootaxa</mods:title>
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<mods:part id="F544EC16B328AFACBD1D224621B54E6B">
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<mods:date id="E7006724B4872EEBB5720772EC352EDB">2014</mods:date>
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<mods:number id="46AC819B8E0FE704071B7E313D976998">3786</mods:number>
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<treatment id="038D4E74FFCC30579F92FE1EE4A20923" ID-DOI="http://doi.org/10.5281/zenodo.6130475" ID-GBIF-Taxon="119422912" ID-Zenodo-Dep="6130475" LSID="urn:lsid:plazi:treatment:038D4E74FFCC30579F92FE1EE4A20923" httpUri="http://treatment.plazi.org/id/038D4E74FFCC30579F92FE1EE4A20923" lastPageId="7" lastPageNumber="490" pageId="1" pageNumber="484">
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<subSubSection id="C33EACE9FFCC30519F92FE1EE71F0F83" pageId="1" pageNumber="484" type="nomenclature">
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<paragraph id="8B9BFF62FFCC30519F92FE1EE7E10FFE" blockId="1.[151,487,403,464]" box="[151,487,403,429]" pageId="1" pageNumber="484">
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<heading id="D0D3480EFFCC30519F92FE1EE7E10FFE" bold="true" box="[151,487,403,429]" fontSize="10" level="2" pageId="1" pageNumber="484" reason="2">
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<emphasis id="B9502370FFCC30519F92FE1EE7E10FFE" bold="true" box="[151,487,403,429]" pageId="1" pageNumber="484">
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<taxonomicName id="4C2484E1FFCC30519F92FE1EE7A60FFF" box="[151,416,403,428]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="monilistylus" status="sp. nov.">
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<emphasis id="B9502370FFCC30519F92FE1EE7A60FFF" bold="true" box="[151,416,403,428]" italics="true" pageId="1" pageNumber="484">Procloeon monilistylus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2639E0BFFCC30519EA2FE19E7E10FFE" box="[423,487,404,429]" pageId="1" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9BFF62FFCC30519F92FE3AE71F0F83" blockId="1.[151,487,403,464]" box="[151,281,439,464]" pageId="1" pageNumber="484">
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(
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<figureCitation id="131FE3E7FFCC30519F9AFE3AE7160F83" box="[159,272,439,464]" captionStart-0="FIGURES 1 – 15" captionStart-1="FIGURES 16 – 22" captionStart-2="FIGURES 23 – 29" captionStart-3="FIGURES 30 – 31" captionStart-4="FIGURES 32 – 35" captionStartId-0="2.[151,264,1926,1949]" captionStartId-1="3.[151,264,1727,1750]" captionStartId-2="4.[151,264,1654,1677]" captionStartId-3="5.[151,264,1894,1917]" captionStartId-4="6.[151,264,1187,1210]" captionTargetBox-0="[173,1412,362,1896]" captionTargetBox-1="[152,1412,216,1698]" captionTargetBox-2="[151,1431,195,1631]" captionTargetBox-3="[153,1434,220,1850]" captionTargetBox-4="[151,1436,194,1165]" captionTargetId-0="figure@2.[167,1424,345,1909]" captionTargetId-1="figure@3.[151,1436,193,1707]" captionTargetId-2="figure@4.[151,1436,193,1633]" captionTargetId-3="figure@5.[151,1436,193,1874]" captionTargetId-4="figure@6.[151,1436,193,1166]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." captionText-1="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." captionText-2="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." captionText-3="FIGURES 30 – 31. Procloeon monilistylus sp. n., styliger and penis of male imago, ventral view (penis dotted, its hidden part shown by interrupted lines). 30, specimen from Ganukan, in projected condition (muscles not shown); 31, specimen from Kyia, penis in re 4 tracted condition (penial muscles and right gonostylar muscle shown by interrupted lines)." captionText-4="FIGURES 32 – 35. Procloeon monilistylus sp. n., eggs and enlarged micropilar area." httpUri-0="https://zenodo.org/record/229327/files/figure.png" httpUri-1="https://zenodo.org/record/229328/files/figure.png" httpUri-2="https://zenodo.org/record/229329/files/figure.png" httpUri-3="https://zenodo.org/record/229330/files/figure.png" httpUri-4="https://zenodo.org/record/229331/files/figure.png" pageId="1" pageNumber="484">Figs 1–35</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33EACE9FFCC30519F92FD8DE7F80C00" pageId="1" pageNumber="484" type="reference_group">
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<paragraph id="8B9BFF62FFCC30519F92FD8DE7F80C00" blockId="1.[151,819,512,595]" pageId="1" pageNumber="484">
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<treatmentCitationGroup id="AB34D84CFFCC30519F92FD8DE7F80C00" pageId="1" pageNumber="484">
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||
<taxonomicName id="4C2484E1FFCC30519F92FD8DE77B0C45" ID-CoL="3R5Y" authority="(Centroptilum)" baseAuthorityName="Centroptilum" box="[151,381,512,534]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="484" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9502370FFCC30519F92FD8DE6E60C45" box="[151,224,512,534]" italics="true" pageId="1" pageNumber="484">Cloeon</emphasis>
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(
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<emphasis id="B9502370FFCC30519FEAFD8DE7730C45" box="[239,373,512,534]" italics="true" pageId="1" pageNumber="484">Centroptilum</emphasis>
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)
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</taxonomicName>
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sp. 1:
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<treatmentCitation id="0A85D973FFCC30519EC4FD8DE4E40C45" author="Kluge" box="[449,738,512,534]" page="64" pageId="1" pageNumber="484" year="1992">
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<bibRefCitation id="EFB58293FFCC30519EC4FD8DE4B90C45" author="Kluge" box="[449,703,512,534]" pageId="1" pageNumber="484" refString="Kluge, N. J. & Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie, 71 (1), 38 - 55. [in Russian]" type="journal article" year="1992">Kluge & Novikova 1992</bibRefCitation>
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: 64
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</treatmentCitation>
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(larva).
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||
<taxonomicName id="4C2484E1FFCC30519F92FD93E4130C67" authority="Tiunova 2003: 9" authorityName="Tiunova" authorityPageNumber="9" authorityYear="2003" box="[151,533,542,564]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="monilistylus">
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<emphasis id="B9502370FFCC30519F92FD93E7670C67" box="[151,353,542,564]" italics="true" pageId="1" pageNumber="484">Cloeon monilistylus</emphasis>
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||
:
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<treatmentCitation id="0A85D973FFCC30519E6BFD93E4130C67" author="Tiunova" box="[366,533,542,564]" page="9" pageId="1" pageNumber="484" year="2003">
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<bibRefCitation id="EFB58293FFCC30519E6BFD93E7F80C67" author="Tiunova" box="[366,510,542,564]" pageId="1" pageNumber="484" refString="Tiunova, T. M. (2003) Podenki (Ephemeroptera) juga Dalnego Vostoka (fauna, biologia, funktsionanlaya ekologia). Autoreferat of Doctoral Thesis, Vladivostok, 47 pp. [in Russian]" type="book" year="2003">Tiunova 2003</bibRefCitation>
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: 9
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</treatmentCitation>
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||
</taxonomicName>
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||
(nomen nudum).
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<taxonomicName id="4C2484E1FFCC30519F92FDB0E7050C00" ID-CoL="63PRK" box="[151,259,573,595]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="490" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
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/g
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<specimenCount id="9D2234EBFFCC30519E13FDB0E7420C00" box="[278,324,573,595]" pageId="1" pageNumber="484" type="generic">1 sp.</specimenCount>
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||
:
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||
<treatmentCitation id="0A85D973FFCC30519E4BFDB0E7FC0C00" author="Kluge" box="[334,506,573,595]" page="375" pageId="1" pageNumber="484" year="2011">
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<bibRefCitation id="EFB58293FFCC30519E4BFDB0E7C10C00" author="Kluge" box="[334,455,573,595]" pageId="1" pageNumber="484" refString="Kluge, N. J. (2011) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2011">Kluge 2011</bibRefCitation>
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||
: 375
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C33EACE9FFCC30529F92FDF2E4E90F13" lastPageId="2" lastPageNumber="485" pageId="1" pageNumber="484" type="materials_examined">
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<paragraph id="8B9BFF62FFCC30519F92FDF2E5080CEF" blockId="1.[151,1436,639,2033]" pageId="1" pageNumber="484">
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<emphasis id="B9502370FFCC30519F92FDF2E7000CCB" bold="true" box="[151,262,639,664]" pageId="1" pageNumber="484">Material.</emphasis>
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<typeStatus id="549F41C0FFCC30519E15FD0DE7AE0CCB" box="[272,424,640,664]" pageId="1" pageNumber="484" type="holotype">HOLOTYPE</typeStatus>
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||
: L-S-I
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||
<emphasis id="B9502370FFCC30519EF6FDF2E40E0CCB" bold="true" box="[499,520,639,664]" pageId="1" pageNumber="484">♂</emphasis>
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,
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||
<collectingCountry id="F333BFF2FFCC30519D1CFD0DE4790CCB" box="[537,639,640,664]" name="Russia" pageId="1" pageNumber="484">RUSSIA</collectingCountry>
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,
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||
<emphasis id="B9502370FFCC30519D8BFDF2E5580CCB" bold="true" box="[654,862,639,664]" pageId="1" pageNumber="484">Primorskiy Kray</emphasis>
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||
(Primorskiy Territory), Spassk region, Hvalynka, river Odarka,
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||
<date id="FF9AD9A2FFCC30519E36FD29E7BF0CEF" box="[307,441,676,700]" pageId="1" pageNumber="484" value="1998-08-04">4.VIII.1998</date>
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||
, coll. D. Apanaskevich (
|
||
<collectionCode id="ED3567A7FFCC30519DC8FD29E5070CEF" box="[717,769,676,700]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="484">ZIN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCC30519FC2FD45E7140A77" blockId="1.[151,1436,639,2033]" pageId="1" pageNumber="484">
|
||
<typeStatus id="549F41C0FFCC30519FC2FD45E7600CB3" box="[199,358,712,736]" pageId="1" pageNumber="484" type="paratype">PARATYPES</typeStatus>
|
||
: L-S-I♂, reared together with
|
||
<typeStatus id="549F41C0FFCC30519DB8FD4AE5240CB3" box="[701,802,711,736]" pageId="1" pageNumber="484" type="holotype">holotype</typeStatus>
|
||
; the same locality, 1 S-I♀ (larval exuviae lost),
|
||
<specimenCount id="9D2234EBFFCC30519A47FD45E39D0CB3" box="[1346,1435,711,736]" pageId="1" pageNumber="484" type="larva">3 larvae</specimenCount>
|
||
(
|
||
<collectionCode id="ED3567A7FFCC30519F9AFD61E6D50D57" box="[159,211,748,772]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="484">ZIN</collectionCode>
|
||
); Hasan Region, river Sidime (= Narva) near natural reserve Kerdrovaya Pad',
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||
<date id="FF9AD9A2FFCC30519B66FD61E2CD0D57" box="[1123,1227,748,772]" pageId="1" pageNumber="484" value="1980-07">VII.1980</date>
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||
, coll. N. Kluge: 1 larva (
|
||
<collectionCode id="ED3567A7FFCC30519FDAFC9DE7150D7B" box="[223,275,784,808]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="484">ZIN</collectionCode>
|
||
); Hasan Region, near Perevoznaya,
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||
<date id="FF9AD9A2FFCC30519DAEFC9DE52B0D7B" box="[683,813,784,808]" pageId="1" pageNumber="484" value="2013-06-21">21.VI.2013</date>
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||
, coll. I. Tiunov:
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||
<date id="FF9AD9A2FFCC30519CEFFC9DE2100D74" box="[1002,1046,784,808]" pageId="1" pageNumber="484" value="2011-01">11 I</date>
|
||
♂ (
|
||
<collectionCode id="ED3567A7FFCC30519B3EFC9DE27D0D7B" box="[1083,1147,784,808]" pageId="1" pageNumber="484">IBSS</collectionCode>
|
||
).
|
||
<emphasis id="B9502370FFCC30519B8AFC82E39D0D7B" bold="true" box="[1167,1435,783,808]" pageId="1" pageNumber="484">
|
||
KHABAROVSKIY
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||
<collectionCode id="ED3567A7FFCC30519A50FC82E39D0D7B" box="[1365,1435,783,808]" pageId="1" pageNumber="484">KRAY</collectionCode>
|
||
</emphasis>
|
||
(Khabarovsk Territory): Pereyaslavka Region, river Kiya,
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||
<quantity id="4CDC5287FFCC30519C27FCB9E55D0D1E" box="[802,859,820,845]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="484" unit="km" value="2.0">2 km</quantity>
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||
lower Petrovichi,
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||
<date id="FF9AD9A2FFCC30519B2CFCB9E29B0D1E" box="[1065,1181,820,845]" pageId="1" pageNumber="484" value="1996-06-03">3.VI.1996</date>
|
||
, coll. T. Tiunova:
|
||
<date id="FF9AD9A2FFCC30519A6DFCB9E3810D1F" box="[1384,1415,820,844]" pageId="1" pageNumber="484">1 I</date>
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||
♂ (
|
||
<collectionCode id="ED3567A7FFCC30519F9AFCD5E6D50D23" box="[159,211,856,880]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="484">ZIN</collectionCode>
|
||
); Nikolaevsk Region, left bank of river Amur near Innokentievka,
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||
<date id="FF9AD9A2FFCC30519CC8FCDAE24C0D23" box="[973,1098,855,880]" pageId="1" pageNumber="484" value="2000-07-06">6.VII.2000</date>
|
||
, coll. T. Tiunova:
|
||
<date id="FF9AD9A2FFCC30519A19FCD5E33B0D3C" box="[1308,1341,856,880]" pageId="1" pageNumber="484">1 I</date>
|
||
♂, 1 S♂ (
|
||
<collectionCode id="ED3567A7FFCC30519F9AFCF1E6D50DC7" box="[159,211,892,916]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="484">ZIN</collectionCode>
|
||
);
|
||
<date id="FF9AD9A2FFCC30519FE9FCF1E7170DC7" box="[236,273,892,917]" pageId="1" pageNumber="484">5 I</date>
|
||
♂,1 S♀ (
|
||
<collectionCode id="ED3567A7FFCC30519E7AFCF1E7B90DC7" box="[383,447,892,916]" pageId="1" pageNumber="484">IBSS</collectionCode>
|
||
); bassin of river Khor, anabranch Potopilikha,
|
||
<date id="FF9AD9A2FFCC30519CF5FCF1E27D0DC6" box="[1008,1147,892,917]" pageId="1" pageNumber="484" value="2005-07-14">14.VII.2005</date>
|
||
, coll. T. Tiunova:
|
||
<date id="FF9AD9A2FFCC30519A59FCF1E3870DC7" box="[1372,1409,892,916]" pageId="1" pageNumber="484">1 I</date>
|
||
♂.
|
||
<emphasis id="B9502370FFCC30519F92FC12E7A70DEB" bold="true" box="[151,417,927,952]" pageId="1" pageNumber="484">AMURSKAYA OBLAST'</emphasis>
|
||
(Amur Province): river Ganukan near Kusatkino,
|
||
<date id="FF9AD9A2FFCC30519CF6FC2DE2810DEB" box="[1011,1159,928,952]" pageId="1" pageNumber="484" value="2003-08-13">13.VIII.2003</date>
|
||
, coll. T. Tiunova:
|
||
<date id="FF9AD9A2FFCC30519A61FC2DE3810DE4" box="[1380,1415,928,952]" pageId="1" pageNumber="484">2 I</date>
|
||
♂ (
|
||
<collectionCode id="ED3567A7FFCC30519F9AFC49E6D50D8F" box="[159,211,964,988]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="484">ZIN</collectionCode>
|
||
);
|
||
<date id="FF9AD9A2FFCC30519FE9FC49E7180D8F" box="[236,286,964,988]" pageId="1" pageNumber="484" value="2013-01">13 I</date>
|
||
♂,
|
||
<date id="FF9AD9A2FFCC30519E43FC49E76A0D8F" box="[326,364,964,988]" pageId="1" pageNumber="484">8 I</date>
|
||
♀ (
|
||
<collectionCode id="ED3567A7FFCC30519E96FC49E7D50D8F" box="[403,467,964,988]" pageId="1" pageNumber="484">IBSS</collectionCode>
|
||
); river Ignashikha (left tributary of river Amur) near frontier post Ignashino,
|
||
<date id="FF9AD9A2FFCC30519A77FC49E72A0A53" pageId="1" pageNumber="484" value="2004-08-28" valueMax="2004-08-29" valueMin="2004-08-28">28– 29.VIII.2004</date>
|
||
, coll. T. Tiunova: 1 larva (
|
||
<collectionCode id="ED3567A7FFCC30519D52FC65E4910A53" box="[599,663,1000,1024]" pageId="1" pageNumber="484">IBSS</collectionCode>
|
||
); river Zeya upstream of Mazanovo,
|
||
<date id="FF9AD9A2FFCC30519B3CFC6AE2C60A53" box="[1081,1216,999,1024]" pageId="1" pageNumber="484" value="2006-08-05">5.VIII.2006</date>
|
||
, coll. T. Tiunova:
|
||
<date id="FF9AD9A2FFCC30519A8BFC65E6A70A77" pageId="1" pageNumber="484">1 I</date>
|
||
♂ (
|
||
<collectionCode id="ED3567A7FFCC30519FC0FB81E7000A77" box="[197,262,1036,1060]" pageId="1" pageNumber="484">IBSS</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCC30519FC2FBA2E25B0AE6" blockId="1.[151,1436,639,2033]" pageId="1" pageNumber="484">
|
||
<emphasis id="B9502370FFCC30519FC2FBA2E7650A1B" bold="true" box="[199,355,1071,1096]" pageId="1" pageNumber="484">Descriptions.</emphasis>
|
||
<emphasis id="B9502370FFCC30519E77FBBCE7BA0A1B" box="[370,444,1073,1096]" italics="true" pageId="1" pageNumber="484">Larva.</emphasis>
|
||
CUTICULAR COLORATION: Thoracic terga light, with brownish maculae, fore protoptera unicolorous. Legs light, femur with contrasting brown band near apex (
|
||
<figureCitation id="131FE3E7FFCC30519B31FBD9E28F0A3F" box="[1076,1161,1108,1133]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="1" pageNumber="484">Fig. 22</figureCitation>
|
||
). Abdominal terga light, with brown maculae similar on all segments II–IX (
|
||
<figureCitation id="131FE3E7FFCC30519DD2FBFAE52C0AC3" box="[727,810,1143,1168]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="1" pageNumber="484">Fig. 17</figureCitation>
|
||
). Each abdominal sternum I–VIII colorless, with pair of brownish sublateral maculae; sternum IX brownish. Caudalii light, without dark band.
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCC30519FC2FB4DE7060B73" blockId="1.[151,1436,639,2033]" pageId="1" pageNumber="484">
|
||
HYPODERMAL COLORATION: Each abdominal tergum II–IX with pair of brown submedian marking adjacent to posterior margin (as in imago,
|
||
<figureCitation id="131FE3E7FFCC30519D86FB69E4D30AAF" box="[643,725,1252,1277]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="1" pageNumber="484">Fig. 18</figureCitation>
|
||
) and visible through blanks located on cuticle at these places (
|
||
<figureCitation id="131FE3E7FFCC30519F9AFA8AE6F50B73" box="[159,243,1287,1312]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="1" pageNumber="484">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCC30519FC2FAA1E7B10BAB" blockId="1.[151,1436,639,2033]" pageId="1" pageNumber="484">
|
||
<collectionCode id="ED3567A7FFCC30519FC2FAA1E7260B17" box="[199,288,1324,1348]" pageId="1" pageNumber="484">SHAPE</collectionCode>
|
||
AND SETATION: Labrum wide, its distal margin concave; setae on distal margin non-branched; outer surface without constant stout setae, with sparsely and irregularly situated thin setae (not shown in
|
||
<figureCitation id="131FE3E7FFCC30519A4DFAC2E3890B3B" box="[1352,1423,1359,1384]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="1" pageNumber="484">Fig. 4</figureCitation>
|
||
). Mandibles relatively long, with convex outer margin; on both mandibles incisor and kinetodontium separated beginning from base (
|
||
<figureCitation id="131FE3E7FFCC30519E89FA1AE7EA0BE3" box="[396,492,1431,1456]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="1" pageNumber="484">Fig. 5, 6</figureCitation>
|
||
). Maxillary palp 3-segmented, with short 2nd segment and long 3rd segment, length of 3rd segment more than twice exceeds length of 2nd segment (
|
||
<figureCitation id="131FE3E7FFCC30519C6FFA31E5A80B87" box="[874,942,1468,1493]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="1" pageNumber="484">Fig. 7</figureCitation>
|
||
). Labial palp with 3rd segment truncate and widened apically (
|
||
<figureCitation id="131FE3E7FFCC30519E61FA52E7AF0BAB" box="[356,425,1503,1528]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="1" pageNumber="484">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCC30519FC2F989E365096F" blockId="1.[151,1436,639,2033]" pageId="1" pageNumber="484">
|
||
<collectingCountry id="F333BFF2FFCC30519FC2F989E6F9084E" box="[199,255,1540,1565]" name="India" pageId="1" pageNumber="484">Hind</collectingCountry>
|
||
protoptera absent; margin of metanotum smooth, without any vestiges of hind protoptera. Fore femur shorter than middle and hind femora; other segments subequal in length on all legs; in
|
||
<typeStatus id="549F41C0FFCC30519BB1F9A4E3130812" box="[1204,1301,1577,1601]" pageId="1" pageNumber="484" type="paratype">paratype</typeStatus>
|
||
male larva proportion of femur/tibia/tarsus/claw on fore leg 90:50:60:35; on middle leg 102:50:60:35; on hind leg 105:50:60:35. Femur with outer side smooth, lacking setae; apex serrate, lacking setae; inner side with sparse, small (7–10 times shorter than femur width), pointed, spine-like setae; on fore leg these setae more densely distributed than on middle and hind legs (
|
||
<figureCitation id="131FE3E7FFCC30519D63F93AE4BC0883" box="[614,698,1719,1744]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="1" pageNumber="484">Fig. 22</figureCitation>
|
||
). Tibia and tarsus with outer side smooth, lacking setae; inner side with pointed, spine-like setae of moderate size and density (2–6 times shorter than tibia width, with spaces subequal to their length) (
|
||
<figureCitation id="131FE3E7FFCC30519EB0F972E40F094B" box="[437,521,1791,1816]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="1" pageNumber="484">Fig. 22</figureCitation>
|
||
). Patella-tibial suture equally developed on all legs. Claw long and slender; with 2 rows of denticles occupying less than 1/3 of claw length; denticles small, very thin, almost seta-like (
|
||
<figureCitation id="131FE3E7FFCC30519A07F8A9E353096F" box="[1282,1365,1828,1853]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="1" pageNumber="484">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCC30529FD6F8CAE4E90F13" blockId="1.[151,1436,639,2033]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" lastPageNumber="485" pageId="1" pageNumber="484">
|
||
Surface of abdominal terga and sterna with translucent scales in semilunar nests. Tergum I with minute irregular denticles on median part of posterior margin, without posterolateral spines. Terga II–IX with larger and smaller pointed spines on median part of posterior margin. Terga II–VII with pointed posterolateral spines (
|
||
<figureCitation id="131FE3E7FFCC30519A6AF802E6B0099F" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="1" pageNumber="484">Fig. 17</figureCitation>
|
||
). Row of spines along lateral margin well-developed on segments VIII and IX; few lateral spines sometimes present on segment VII. Tergum X with long pointed spines on posterior margin (
|
||
<figureCitation id="131FE3E7FFCC30519B53F855E29B09A3" box="[1110,1181,2008,2033]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="1" pageNumber="484">Fig. 9</figureCitation>
|
||
). Sterna I–III or I–IV without spines or denticles on posterior margin. Posterior margins of sterna V–IX with pointed spines smaller than spines on posterior margins of terga. Paraproct with long pointed spines (similar to spines on terga) (
|
||
<figureCitation id="131FE3E7FFCF30529A1EFF31E3670E87" box="[1307,1377,188,213]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="2" pageNumber="485">Fig. 9</figureCitation>
|
||
). All tergalii I–VII pointed apically; tergalii I–V sharply asymmetric, with prominent proximal-anal lobe (
|
||
<figureCitation id="131FE3E7FFCF30529A09FF52E3960EAB" box="[1292,1424,223,248]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="2" pageNumber="485">Figs 10–15</figureCitation>
|
||
). Caudalii with posterior margin of each 4th segment darkened, with large spines; in distal part of cercus lateral spines very large, equal to segment length (
|
||
<figureCitation id="131FE3E7FFCF30529D79FEAAE4E70F13" box="[636,737,295,320]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="2" pageNumber="485">Figs 1–3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5BAFEAFFCF30529F92F80BE7D40989" httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="2" pageNumber="485" targetBox="[173,1412,362,1896]" targetPageId="2">
|
||
<paragraph id="8B9BFF62FFCF30529F92F80BE7D40989" blockId="2.[151,1436,1926,2010]" pageId="2" pageNumber="485">
|
||
<emphasis id="B9502370FFCF30529F92F80BE74D09CF" bold="true" box="[151,331,1926,1949]" pageId="2" pageNumber="485">FIGURES 1–15.</emphasis>
|
||
<taxonomicName id="4C2484E1FFCF30529E50F80AE44409CE" box="[341,578,1927,1949]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="monilistylus" status="sp. nov.">
|
||
<emphasis id="B9502370FFCF30529E50F80AE44409CE" box="[341,578,1927,1949]" italics="true" pageId="2" pageNumber="485">Procloeon monilistylus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2639E0BFFCF30529D48F80AE48C09CF" box="[589,650,1927,1948]" pageId="2" pageNumber="485" rank="species">
|
||
<emphasis id="B9502370FFCF30529D48F80AE48C09CF" bold="true" box="[589,650,1927,1948]" pageId="2" pageNumber="485">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, larva. 1–3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10–15, tergalii of pairs I, II, IV, V, VI and VII.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5BAFEAFFCE30539F92F932E3050940" httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="3" pageNumber="486" targetBox="[152,1412,216,1698]" targetPageId="3">
|
||
<paragraph id="8B9BFF62FFCE30539F92F932E3050940" blockId="3.[151,1435,1727,1811]" pageId="3" pageNumber="486">
|
||
<emphasis id="B9502370FFCE30539F92F932E7550886" bold="true" box="[151,339,1727,1750]" pageId="3" pageNumber="486">FIGURES 16–22.</emphasis>
|
||
<taxonomicName id="4C2484E1FFCE30539E5CF94DE4470885" box="[345,577,1728,1750]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="monilistylus" status="sp. nov.">
|
||
<emphasis id="B9502370FFCE30539E5CF94DE4470885" box="[345,577,1728,1750]" italics="true" pageId="3" pageNumber="486">Procloeon monilistylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9502370FFCE30539D42F94DE4790886" bold="true" box="[583,639,1728,1749]" pageId="3" pageNumber="486">
|
||
<taxonomicNameLabel id="A2639E0BFFCE30539D42F94DE4790886" box="[583,639,1728,1749]" pageId="3" pageNumber="486" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5BAFEAFFC930549F92F9FBE2330975" httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="4" pageNumber="487" targetBox="[151,1431,195,1631]" targetPageId="4">
|
||
<paragraph id="8B9BFF62FFC930549F92F9FBE2330975" blockId="4.[151,1436,1654,1830]" pageId="4" pageNumber="487">
|
||
<emphasis id="B9502370FFC930549F92F9FBE75008DF" bold="true" box="[151,342,1654,1677]" pageId="4" pageNumber="487">FIGURES 23–29.</emphasis>
|
||
<taxonomicName id="4C2484E1FFC930549E58F9FBE44E08DF" box="[349,584,1654,1676]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="monilistylus" status="sp. nov.">
|
||
<emphasis id="B9502370FFC930549E58F9FBE44E08DF" box="[349,584,1654,1676]" italics="true" pageId="4" pageNumber="487">Procloeon monilistylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9502370FFC930549D55F9FAE48D08DF" bold="true" box="[592,651,1655,1676]" pageId="4" pageNumber="487">
|
||
<taxonomicNameLabel id="A2639E0BFFC930549D55F9FAE48D08DF" box="[592,651,1655,1676]" pageId="4" pageNumber="487" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5BAFEAFFC830559F92F8EBE29909E9" httpUri="https://zenodo.org/record/229330/files/figure.png" pageId="5" pageNumber="488" targetBox="[153,1434,220,1850]" targetPageId="5">
|
||
<paragraph id="8B9BFF62FFC830559F92F8EBE29909E9" blockId="5.[151,1436,1894,1978]" pageId="5" pageNumber="488">
|
||
<emphasis id="B9502370FFC830559F92F8EBE753092F" bold="true" box="[151,341,1894,1917]" pageId="5" pageNumber="488">FIGURES 30–31.</emphasis>
|
||
<taxonomicName id="4C2484E1FFC830559E5EF8EAE443092E" box="[347,581,1895,1917]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="monilistylus" status="sp. nov.">
|
||
<emphasis id="B9502370FFC830559E5EF8EAE443092E" box="[347,581,1895,1917]" italics="true" pageId="5" pageNumber="488">Procloeon monilistylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9502370FFC830559D49F8EAE480092F" bold="true" box="[588,646,1895,1916]" pageId="5" pageNumber="488">
|
||
<taxonomicNameLabel id="A2639E0BFFC830559D49F8EAE480092F" box="[588,646,1895,1916]" pageId="5" pageNumber="488" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, styliger and penis of male imago, ventral view (penis dotted, its hidden part shown by interrupted lines). 30, specimen from Ganukan, in projected condition (muscles not shown); 31, specimen from Kyia, penis in re4tracted condition (penial muscles and right gonostylar muscle shown by interrupted lines).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5BAFEAFFCB30569F92FB2EE5EA0AEA" box="[151,1004,1187,1210]" httpUri="https://zenodo.org/record/229331/files/figure.png" pageId="6" pageNumber="489" targetBox="[151,1436,194,1165]" targetPageId="6">
|
||
<paragraph id="8B9BFF62FFCB30569F92FB2EE5EA0AEA" blockId="6.[151,1004,1187,1210]" box="[151,1004,1187,1210]" pageId="6" pageNumber="489">
|
||
<emphasis id="B9502370FFCB30569F92FB2EE7550AEB" bold="true" box="[151,339,1187,1210]" pageId="6" pageNumber="489">FIGURES 32–35.</emphasis>
|
||
<taxonomicName id="4C2484E1FFCB30569E5FFB2EE4450AEA" box="[346,579,1187,1209]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="489" phylum="Arthropoda" rank="species" species="monilistylus" status="sp. nov.">
|
||
<emphasis id="B9502370FFCB30569E5FFB2EE4450AEA" box="[346,579,1187,1209]" italics="true" pageId="6" pageNumber="489">Procloeon monilistylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9502370FFCB30569D4CFB29E4840AEA" bold="true" box="[585,642,1188,1209]" pageId="6" pageNumber="489">
|
||
<taxonomicNameLabel id="A2639E0BFFCB30569D4CFB29E4840AEA" box="[585,642,1188,1209]" pageId="6" pageNumber="489" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, eggs and enlarged micropilar area.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33EACE9FFCB30579FC2FB6AE47F0FA7" lastPageId="7" lastPageNumber="490" pageId="6" pageNumber="489" type="description">
|
||
<paragraph id="8B9BFF62FFCB30569FC2FB6AE2FE0B71" blockId="6.[151,1437,1254,2035]" pageId="6" pageNumber="489">
|
||
MALE STRUCTURE: Sibimaginal gonostyli developing under larval cuticle, folded such that 1st segments approximate, 2nd segments directed laterally-caudally and 3rd segments directed caudally (
|
||
<figureCitation id="131FE3E7FFCB30569B92FA87E2EA0B71" box="[1175,1260,1290,1315]" captionStart="FIGURES 23 – 29" captionStartId="4.[151,264,1654,1677]" captionTargetBox="[151,1431,195,1631]" captionTargetId="figure@4.[151,1436,193,1633]" captionTargetPageId="4" captionText="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="6" pageNumber="489">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCB30569FC2FAA3E7040BDC" blockId="6.[151,1437,1254,2035]" pageId="6" pageNumber="489">
|
||
<emphasis id="B9502370FFCB30569FC2FAA3E7380B15" box="[199,318,1326,1350]" italics="true" pageId="6" pageNumber="489">Subimago.</emphasis>
|
||
CUTICULAR COLORATION: Lateral part of mesonotum with brown markings (
|
||
<figureCitation id="131FE3E7FFCB30569A37FAA3E3890B14" box="[1330,1423,1326,1351]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="6" pageNumber="489">Fig. 20</figureCitation>
|
||
); mesothoracic anepimeron brown; posterior part of postsubalar sclerite brown (
|
||
<figureCitation id="131FE3E7FFCB30569B16FADFE2610B39" box="[1043,1127,1362,1387]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="6" pageNumber="489">Fig. 19</figureCitation>
|
||
); other subimaginal cuticle colorless.
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCB30569FC2FA17E4E30BE0" blockId="6.[151,1437,1254,2035]" box="[199,741,1434,1459]" pageId="6" pageNumber="489">
|
||
HYPODERMAL COLORATION:
|
||
<collectingCountry id="F333BFF2FFCB30569D50FA17E4720BE1" box="[597,628,1434,1458]" name="American Samoa" pageId="6" pageNumber="489">As</collectingCountry>
|
||
in imago.
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCB30569FC2FA32E50F08D9" blockId="6.[151,1437,1254,2035]" pageId="6" pageNumber="489">
|
||
SHAPE AND TEXTURE: On fore leg 1st tarsal segment (shortened) covered by microtrichiae (as tibia and most of rest of body); 2nd–5th segments covered by pointed microlepides. On middle and hind legs, area corresponding to 1st tarsal segment (fused with 2nd segment) covered by microtrichiae (as tibia and most of rest of body); 2nd–5th segments covered by pointed microlepides. Gonostylus with 2nd segment thicker than in imago, without subapical projection and without grooved concavity of inner margin; unlike imago, 3rd (apical) segment attached not ventrally, but apically or dorsally (
|
||
<figureCitation id="131FE3E7FFCB30569DADF9FFE4FA08D9" box="[680,764,1650,1675]" captionStart="FIGURES 23 – 29" captionStartId="4.[151,264,1654,1677]" captionTargetBox="[151,1431,195,1631]" captionTargetId="figure@4.[151,1436,193,1633]" captionTargetPageId="4" captionText="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="6" pageNumber="489">Fig. 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCB30569FC2F91AE2FC09F8" blockId="6.[151,1437,1254,2035]" pageId="6" pageNumber="489">
|
||
<emphasis id="B9502370FFCB30569FC2F91AE75808FD" box="[199,350,1686,1710]" italics="true" pageId="6" pageNumber="489">Imago, male.</emphasis>
|
||
Head and thorax light-yellowish. Turbinate eyes unicolorous, yellow, high, widened apically; facetted surfaces nearly symmetrically-oval (
|
||
<figureCitation id="131FE3E7FFCB30569D95F937E51C0881" box="[656,794,1722,1747]" captionStart="FIGURES 23 – 29" captionStartId="4.[151,264,1654,1677]" captionTargetBox="[151,1431,195,1631]" captionTargetId="figure@4.[151,1436,193,1633]" captionTargetPageId="4" captionText="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="6" pageNumber="489">Figs. 28, 29</figureCitation>
|
||
). Medial longitudinal suture of mesonotum convex (as in
|
||
<figureCitation id="131FE3E7FFCB30569F92F953E6EC08A4" box="[151,234,1758,1783]" captionStart="FIGURES 23 – 29" captionStartId="4.[151,264,1654,1677]" captionTargetBox="[151,1431,195,1631]" captionTargetId="figure@4.[151,1436,193,1633]" captionTargetPageId="4" captionText="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="6" pageNumber="489">Fig. 26</figureCitation>
|
||
). Fore wings colorless, veins colorless, pterostigma with 3–6 non-branched oblique veins, sometimes some incomplete (
|
||
<figureCitation id="131FE3E7FFCB30569E20F88FE77D0948" box="[293,379,1794,1819]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="6" pageNumber="489">Fig. 16</figureCitation>
|
||
).
|
||
<collectingCountry id="F333BFF2FFCB30569E97F88FE7CC0948" box="[402,458,1794,1819]" name="India" pageId="6" pageNumber="489">Hind</collectingCountry>
|
||
wings absent. Legs light yellowish, unicolorous. In
|
||
<typeStatus id="549F41C0FFCB30569B22F88FE28A0948" box="[1063,1164,1794,1819]" pageId="6" pageNumber="489" type="holotype">holotype</typeStatus>
|
||
, proportion of femur to tibia to tarsal segments 1st+2nd, 3rd, 4th and 5th on fore leg 120:150:70:60:35:20; on middle and hind leg 110:85:40:14:6:16. On middle and hind legs tarsus with 2 apical thorns: 1st+2nd and 3rd tarsomeres thorn-baring. Abdomen light yellowish, each tergum with pair of submedian brown maculae near posterior margin; terga IV and VII with pair of indistinct sublateral brownish maculae (
|
||
<figureCitation id="131FE3E7FFCB30569C08F81FE56709F9" box="[781,865,1938,1963]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="6" pageNumber="489">Fig. 18</figureCitation>
|
||
). Cerci light yellowish, unicolorous.
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCB30579FC2F83AE79E0F13" blockId="6.[151,1437,1254,2035]" lastBlockId="7.[151,1437,151,1904]" lastPageId="7" lastPageNumber="490" pageId="6" pageNumber="489">
|
||
STRUCTURE OF MALE GENITALS (
|
||
<figureCitation id="131FE3E7FFCB30569D8AF83BE514099C" box="[655,786,1974,1999]" captionStart="FIGURES 23 – 29" captionStartId="4.[151,264,1654,1677]" captionTargetBox="[151,1431,195,1631]" captionTargetId="figure@4.[151,1436,193,1633]" captionTargetPageId="4" captionText="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="6" pageNumber="489">Figs 23–25</figureCitation>
|
||
,
|
||
<figureCitation id="131FE3E7FFCB30569C1AF83AE56D099C" box="[799,875,1975,1999]" captionStart="FIGURES 30 – 31" captionStartId="5.[151,264,1894,1917]" captionTargetBox="[153,1434,220,1850]" captionTargetId="figure@5.[151,1436,193,1874]" captionTargetPageId="5" captionText="FIGURES 30 – 31. Procloeon monilistylus sp. n., styliger and penis of male imago, ventral view (penis dotted, its hidden part shown by interrupted lines). 30, specimen from Ganukan, in projected condition (muscles not shown); 31, specimen from Kyia, penis in re 4 tracted condition (penial muscles and right gonostylar muscle shown by interrupted lines)." httpUri="https://zenodo.org/record/229330/files/figure.png" pageId="6" pageNumber="489">30, 31</figureCitation>
|
||
): Styliger with median finger-like process. Shape of gonostylus composite: proximal part of 1st segment sharply projected medially; 2nd segment widened apically, with subapical median projection and grooved concave median margin proximad of this projection; 3rd (apical) segment long, attached on ventral side of 2nd segment (
|
||
<figureCitation id="131FE3E7FFCA30579C19FF31E5740E86" box="[796,882,188,213]" captionStart="FIGURES 23 – 29" captionStartId="4.[151,264,1654,1677]" captionTargetBox="[151,1431,195,1631]" captionTargetId="figure@4.[151,1436,193,1633]" captionTargetPageId="4" captionText="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="7" pageNumber="490">Fig. 25</figureCitation>
|
||
). Penis with unusual 2-stepped shape of median external part (
|
||
<figureCitation id="131FE3E7FFCA30579E37FF52E7B50EAB" box="[306,435,223,248]" captionStart="FIGURES 30 – 31" captionStartId="5.[151,264,1894,1917]" captionTargetBox="[153,1434,220,1850]" captionTargetId="figure@5.[151,1436,193,1874]" captionTargetPageId="5" captionText="FIGURES 30 – 31. Procloeon monilistylus sp. n., styliger and penis of male imago, ventral view (penis dotted, its hidden part shown by interrupted lines). 30, specimen from Ganukan, in projected condition (muscles not shown); 31, specimen from Kyia, penis in re 4 tracted condition (penial muscles and right gonostylar muscle shown by interrupted lines)." httpUri="https://zenodo.org/record/229330/files/figure.png" pageId="7" pageNumber="490">Figs 30, 31</figureCitation>
|
||
). Ventral-proximal margin of penis (hidden by styliger) with pair of sclerotized sublateral thickenings at points of muscles attachment; apices of gonovectes fused with penial bridge, so that gonovectes resemble pair of loops.
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCA30579FC2FEC0E2D30FFE" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="490">
|
||
<emphasis id="B9502370FFCA30579FC2FEC0E7740F37" box="[199,370,332,356]" italics="true" pageId="7" pageNumber="490">Imago, female.</emphasis>
|
||
Coloration as in male. Eyes large, elevated above head, approximate, with parallel median margins (
|
||
<figureCitation id="131FE3E7FFCA30579FFAFEE2E7870FDB" box="[255,385,367,392]" captionStart="FIGURES 23 – 29" captionStartId="4.[151,264,1654,1677]" captionTargetBox="[151,1431,195,1631]" captionTargetId="figure@4.[151,1436,193,1633]" captionTargetPageId="4" captionText="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="7" pageNumber="490">Figs 26, 27</figureCitation>
|
||
). On fore leg tarsus with 2 apical thorns: 2nd and 3rd tarsomeres thorn-bearing; on middle and hind legs tarsus also with 2 apical thorns: 1st+2nd and 3rd tarsomeres thorn-bearing (as in male).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCA30579FC2FE34E31A0F83" blockId="7.[151,1437,151,1904]" box="[199,1308,439,464]" pageId="7" pageNumber="490">
|
||
<emphasis id="B9502370FFCA30579FC2FE34E6FD0F83" box="[199,251,441,464]" italics="true" pageId="7" pageNumber="490">Egg.</emphasis>
|
||
Oval, 0.16 mm long. Chorion with even net-like relief with irregular small cells (
|
||
<figureCitation id="131FE3E7FFCA30579B88FE3AE3160F83" box="[1165,1296,439,464]" captionStart="FIGURES 32 – 35" captionStartId="6.[151,264,1187,1210]" captionTargetBox="[151,1436,194,1165]" captionTargetId="figure@6.[151,1436,193,1166]" captionTargetPageId="6" captionText="FIGURES 32 – 35. Procloeon monilistylus sp. n., eggs and enlarged micropilar area." httpUri="https://zenodo.org/record/229331/files/figure.png" pageId="7" pageNumber="490">Figs 32–35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCA30579FC2FE50E47F0FA7" blockId="7.[151,1437,151,1904]" box="[199,633,476,501]" pageId="7" pageNumber="490">
|
||
<emphasis id="B9502370FFCA30579FC2FE50E74E0FA7" box="[199,328,477,500]" italics="true" pageId="7" pageNumber="490">Dimension.</emphasis>
|
||
Fore wing length
|
||
<quantity id="4CDC5287FFCA30579D11FE51E4720FA7" box="[532,628,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="7" pageNumber="490" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33EACE9FFCA30579FC2FE72E58B0C6E" pageId="7" pageNumber="490" type="etymology">
|
||
<paragraph id="8B9BFF62FFCA30579FC2FE72E58B0C6E" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="490">
|
||
<emphasis id="B9502370FFCA30579FC2FE72E7480C4B" bold="true" box="[199,334,511,536]" pageId="7" pageNumber="490">Etymology.</emphasis>
|
||
The specific epithet is a reference to the appearance of the gonostyli. They appear moniliform because of the presence of a median projection on the 2nd segment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33EACE9FFCA30579FC2FDCAE7010CD6" pageId="7" pageNumber="490" type="diagnosis">
|
||
<paragraph id="8B9BFF62FFCA30579FC2FDCAE7010CD6" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="490">
|
||
<emphasis id="B9502370FFCA30579FC2FDCAE7470C33" bold="true" box="[199,321,583,608]" pageId="7" pageNumber="490">Diagnosis.</emphasis>
|
||
The male imago of
|
||
<taxonomicName id="4C2484E1FFCA30579D3BFDC4E4E10C33" box="[574,743,584,608]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="monilistylus">
|
||
<emphasis id="B9502370FFCA30579D3BFDC4E44A0C33" box="[574,588,585,608]" italics="true" pageId="7" pageNumber="490">P</emphasis>
|
||
.
|
||
<emphasis id="B9502370FFCA30579D65FDC5E4E10C33" box="[608,743,584,608]" italics="true" pageId="7" pageNumber="490">monilistylus</emphasis>
|
||
</taxonomicName>
|
||
differs from all other mayfly species by the shape of the gonostyli.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33EACE9FFCA30579FC2FD02E39B0D46" pageId="7" pageNumber="490" type="reference_group">
|
||
<paragraph id="8B9BFF62FFCA30579FC2FD02E39B0D46" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="490">
|
||
Besides
|
||
<taxonomicName id="4C2484E1FFCA30579E20FD1CE7CF0CFB" box="[293,457,656,680]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="monilistylus">
|
||
<emphasis id="B9502370FFCA30579E20FD1CE7CF0CFB" box="[293,457,656,680]" italics="true" pageId="7" pageNumber="490">P. monilistylus</emphasis>
|
||
</taxonomicName>
|
||
, only two Palaearctic species of
|
||
<taxonomicName id="4C2484E1FFCA30579C39FD1DE5B60CFB" ID-CoL="63PRK" box="[828,944,656,680]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579C39FD1DE5B60CFB" box="[828,944,656,680]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
lack hind wings in the adult stage and hind protoptera in the larval stage:
|
||
<taxonomicName id="4C2484E1FFCA30579EFEFD38E5320C9F" ID-CoL="4MNVT" authority="Bengtsson 1912" authorityName="Bengtsson" authorityYear="1912" box="[507,820,692,717]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="bifidum">
|
||
<emphasis id="B9502370FFCA30579EFEFD38E40F0C9F" box="[507,521,693,716]" italics="true" pageId="7" pageNumber="490">P</emphasis>
|
||
.
|
||
<emphasis id="B9502370FFCA30579D19FD39E4770C9F" box="[540,625,692,716]" italics="true" pageId="7" pageNumber="490">bifidum</emphasis>
|
||
Bengtsson 1912
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C2484E1FFCA30579C61FD38E2AF0C9F" authority="Tshernova 1928" authorityName="Tshernova" authorityYear="1928" box="[868,1193,692,717]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="ornatum">
|
||
<emphasis id="B9502370FFCA30579C61FD38E5740C9F" box="[868,882,693,716]" italics="true" pageId="7" pageNumber="490">P</emphasis>
|
||
.
|
||
<emphasis id="B9502370FFCA30579C81FD38E5E50C9F" box="[900,995,693,716]" italics="true" pageId="7" pageNumber="490">ornatum</emphasis>
|
||
Tshernova 1928
|
||
</taxonomicName>
|
||
; =
|
||
<taxonomicName id="4C2484E1FFCA30579BD1FD38E7440CA3" authority="Kimmins 1957" authorityName="Kimmins" authorityYear="1957" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="pseudorufulum">
|
||
<emphasis id="B9502370FFCA30579BD1FD38E39A0C9F" box="[1236,1436,692,716]" italics="true" pageId="7" pageNumber="490">P. pseudorufulum</emphasis>
|
||
Kimmins 1957
|
||
</taxonomicName>
|
||
; =
|
||
<taxonomicName id="4C2484E1FFCA30579E6DFD54E4C10CA3" authority="Ikonomov 1962" authorityName="Ikonomov" authorityYear="1962" box="[360,711,727,752]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="lychnidense">
|
||
<emphasis id="B9502370FFCA30579E6DFD54E7700CA3" box="[360,374,729,752]" italics="true" pageId="7" pageNumber="490">P</emphasis>
|
||
.
|
||
<emphasis id="B9502370FFCA30579E80FD55E40F0CA3" box="[389,521,728,752]" italics="true" pageId="7" pageNumber="490">lychnidense</emphasis>
|
||
Ikonomov 1962
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C2484E1FFCA30579C0DFD54E2EE0CA3" authority="Kluge & Novikova 1992" authorityName="Kluge & Novikova" authorityYear="1992" box="[776,1256,727,752]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="heterophillum">
|
||
<emphasis id="B9502370FFCA30579C0DFD54E5100CA3" box="[776,790,729,752]" italics="true" pageId="7" pageNumber="490">P</emphasis>
|
||
.
|
||
<emphasis id="B9502370FFCA30579C20FD55E5C70CA3" box="[805,961,728,752]" italics="true" pageId="7" pageNumber="490">heterophillum</emphasis>
|
||
<bibRefCitation id="EFB58293FFCA30579CCFFD5AE2EE0CA3" author="Kluge" box="[970,1256,727,752]" pageId="7" pageNumber="490" refString="Kluge, N. J. & Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie, 71 (1), 38 - 55. [in Russian]" type="journal article" year="1992">Kluge & Novikova 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
. The larva of
|
||
<taxonomicName id="4C2484E1FFCA30579A82FD54E7180D47" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="monilistylus">
|
||
<emphasis id="B9502370FFCA30579A82FD54E3930CA3" box="[1415,1429,729,752]" italics="true" pageId="7" pageNumber="490">P</emphasis>
|
||
.
|
||
<emphasis id="B9502370FFCA30579F92FD71E7180D47" box="[151,286,764,788]" italics="true" pageId="7" pageNumber="490">monilistylus</emphasis>
|
||
</taxonomicName>
|
||
differs from these two species by having a deeply divided incisor and kinetodontium of the mandibles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33EACE9FFCA30579FC2FC92E4A20923" pageId="7" pageNumber="490" type="discussion">
|
||
<paragraph id="8B9BFF62FFCA30579FC2FC92E7530DD3" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="490">
|
||
<emphasis id="B9502370FFCA30579FC2FC92E74A0D6B" bold="true" box="[199,332,799,824]" pageId="7" pageNumber="490">Discussion.</emphasis>
|
||
Originally, the species described here was wrongly attributed not to
|
||
<taxonomicName id="4C2484E1FFCA30579BC1FCADE33E0D6B" box="[1220,1336,800,824]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579BC1FCADE33E0D6B" box="[1220,1336,800,824]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
, but to
|
||
<taxonomicName id="4C2484E1FFCA30579F92FCC9E72D0D0F" box="[151,299,836,860]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579F92FCC9E72D0D0F" box="[151,299,836,860]" italics="true" pageId="7" pageNumber="490">Centroptilum</emphasis>
|
||
</taxonomicName>
|
||
, because it was known only from a single larval specimen lacking tergalii and caudalii (
|
||
<bibRefCitation id="EFB58293FFCA30579A30FCC9E74E0DD3" author="Kluge" pageId="7" pageNumber="490" refString="Kluge, N. J. & Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie, 71 (1), 38 - 55. [in Russian]" type="journal article" year="1992">Kluge & Novikova 1992</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCA30579FC2FC01E5930B5E" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="490">
|
||
The taxon
|
||
<taxonomicName id="4C2484E1FFCA30579E44FC01E7B30DF7" box="[321,437,908,932]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B9502370FFCA30579E44FC01E7B30DF7" box="[321,437,908,932]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2639E0BFFCA30579EB9FC01E7DB0DF6" box="[444,477,908,933]" pageId="7" pageNumber="490" sensu="lato">s.l.</taxonomicNameLabel>
|
||
(or
|
||
<taxonomicName id="4C2484E1FFCA30579D08FC01E4800DF6" box="[525,646,908,933]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
|
||
/g1, according to the rank-free hierarchical nomenclature) corresponds to the phylogenetic branch
|
||
<quantity id="4CDC5287FFCA30579EDDFC3DE4150D9B" box="[472,531,943,968]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="7" pageNumber="490" unit="in" value="11.0">11 in</quantity>
|
||
our earlier paper (
|
||
<bibRefCitation id="EFB58293FFCA30579DF7FC22E27C0D9B" author="Kluge" box="[754,1146,943,968]" pageId="7" pageNumber="490" refString="Kluge, N. J. & Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie, 71 (1), 38 - 55. [in Russian]" type="journal article" year="1992">
|
||
Kluge & Novikova 1992:
|
||
<figureCitation id="131FE3E7FFCA30579B2DFC22E27C0D9B" box="[1064,1146,943,968]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="7" pageNumber="490">Fig. 11</figureCitation>
|
||
</bibRefCitation>
|
||
) and is characterized by autapomorphies: (1) denticles on outer side of larval cerci are enlarged and transformed to heavy spines in such a manner, that in distal part of cercus each segment bears a single spine as long as the segment; (2) eyes of female imago are enlarged, elongate and elevated, with parallel inner margins. Besides this,
|
||
<taxonomicName id="4C2484E1FFCA30579B79FB91E2F60A67" box="[1148,1264,1052,1076]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579B79FB91E2F60A67" box="[1148,1264,1052,1076]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
has characters common with
|
||
<taxonomicName id="4C2484E1FFCA30579E32FBCDE78E0A0B" box="[311,392,1088,1112]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B9502370FFCA30579E32FBCDE78E0A0B" box="[311,392,1088,1112]" italics="true" pageId="7" pageNumber="490">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2639E0BFFCA30579E8BFBCDE7C30A04" box="[398,453,1088,1111]" pageId="7" pageNumber="490" sensu="stricto">s.str.</taxonomicNameLabel>
|
||
: (3) lateral sides of larval abdominal segments VIII and IX bear denticles; (4) in genitals of the male imago gonovectes are fused by their apices with the penial bridge. All these characters are present in
|
||
<taxonomicName id="4C2484E1FFCA30579F92FB05E79C0AF3" box="[151,410,1160,1184]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="monilistylus" status="sp. nov.">
|
||
<emphasis id="B9502370FFCA30579F92FB05E79C0AF3" box="[151,410,1160,1184]" italics="true" pageId="7" pageNumber="490">Procloeon monilistylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9502370FFCA30579EA7FB0AE7E40AF3" bold="true" box="[418,482,1159,1184]" pageId="7" pageNumber="490">
|
||
<taxonomicNameLabel id="A2639E0BFFCA30579EA7FB0AE7E40AF3" box="[418,482,1159,1184]" pageId="7" pageNumber="490" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="131FE3E7FFCA30579EF7FB0AE43D0AF3" box="[498,571,1159,1184]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,264,1926,1949]" captionTargetBox="[173,1412,362,1896]" captionTargetId="figure@2.[167,1424,345,1909]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Procloeon monilistylus sp. n., larva. 1 – 3, cercus and two portions of its distal part; 4, labrum; 5, right mandible; 6, left mandible; 7, maxillary palp; 8, labial palp; 9, paraproct and margin of tenth abdominal tergum; 10 – 15, tergalii of pairs I, II, IV, V, VI and VII." httpUri="https://zenodo.org/record/229327/files/figure.png" pageId="7" pageNumber="490">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="131FE3E7FFCA30579D4CFB05E4610AF3" box="[585,615,1160,1184]" captionStart="FIGURES 16 – 22" captionStartId="3.[151,264,1727,1750]" captionTargetBox="[152,1412,216,1698]" captionTargetId="figure@3.[151,1436,193,1707]" captionTargetPageId="3" captionText="FIGURES 16 – 22. Procloeon monilistylus sp. n. 16, wing; 17, cuticle of larval terga, spread on slide (hypodermal pigmentation not shown); 18, abdominal terga of male imago, spread on slide; 19, cuticle of subimaginal postsubalar sclerite and lateropostnotal crest; 20, cuticle of subimaginal mesonotum, spread on slide; 21, larval claw; 22, larval fore leg." httpUri="https://zenodo.org/record/229328/files/figure.png" pageId="7" pageNumber="490">17</figureCitation>
|
||
,
|
||
<figureCitation id="131FE3E7FFCA30579D71FB05E4EE0AF3" box="[628,744,1159,1184]" captionStart="FIGURES 23 – 29" captionStartId="4.[151,264,1654,1677]" captionTargetBox="[151,1431,195,1631]" captionTargetId="figure@4.[151,1436,193,1633]" captionTargetPageId="4" captionText="FIGURES 23 – 29. Procloeon monilistylus sp. n. 23, posterior margin of male larval abdominal sternum IX and subimaginal genitals developing under larval cuticle (right gonostylar muscle shown by interrupted lines); 24, genitals of male subimago, ventral view (imaginal gonostyli and gonostyli pedestals developing under subimaginal cuticle, shown by interrupted lines); 25, male imaginal abdominal sternum IX with styliger and gonostyli (unpaired sterno-styligeral muscle, pair of penial muscles and right gonostylar muscle shown by interrupted lines); 26, had and thorax of female imago, frontal view; 27, head of female imago, dorsal view; 28, head of male imago, dorsal view; head of male imago, lateral view." httpUri="https://zenodo.org/record/229329/files/figure.png" pageId="7" pageNumber="490">23, 26, 27</figureCitation>
|
||
,
|
||
<figureCitation id="131FE3E7FFCA30579DF0FB05E5390AF3" box="[757,831,1160,1184]" captionStart="FIGURES 30 – 31" captionStartId="5.[151,264,1894,1917]" captionTargetBox="[153,1434,220,1850]" captionTargetId="figure@5.[151,1436,193,1874]" captionTargetPageId="5" captionText="FIGURES 30 – 31. Procloeon monilistylus sp. n., styliger and penis of male imago, ventral view (penis dotted, its hidden part shown by interrupted lines). 30, specimen from Ganukan, in projected condition (muscles not shown); 31, specimen from Kyia, penis in re 4 tracted condition (penial muscles and right gonostylar muscle shown by interrupted lines)." httpUri="https://zenodo.org/record/229330/files/figure.png" pageId="7" pageNumber="490">30, 31</figureCitation>
|
||
). The taxon
|
||
<taxonomicName id="4C2484E1FFCA30579CCAFB05E2650AF3" box="[975,1123,1160,1184]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579CCAFB05E2650AF3" box="[975,1123,1160,1184]" italics="true" pageId="7" pageNumber="490">Centroptilum</emphasis>
|
||
</taxonomicName>
|
||
is monotypic (
|
||
<bibRefCitation id="EFB58293FFCA30579A08FB0AE3930AF3" author="Kluge" box="[1293,1429,1159,1184]" pageId="7" pageNumber="490" refString="Kluge, N. J. (2011) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376." type="journal article" year="2011">Kluge 2011</bibRefCitation>
|
||
) and differs from
|
||
<taxonomicName id="4C2484E1FFCA30579E57FB21E7C00A97" box="[338,454,1196,1220]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579E57FB21E7C00A97" box="[338,454,1196,1220]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
by the following characters: (1) larval cerci lack heavy spines; (2) eyes of female imago are small, widely separated and located in the same plane with dorsal surface of head; (3) larval abdominal segments without lateral denticles; (4) gonovectes retain free apices.
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCA30579FC2FA9AE2F80BB7" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="490">
|
||
The taxon
|
||
<taxonomicName id="4C2484E1FFCA30579E53FA9AE7CB0B63" box="[342,461,1303,1328]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
|
||
/g1 includes a subordinated taxon
|
||
<taxonomicName id="4C2484E1FFCA30579C87FA95E31D0B63" authority="Jacob 1987" authorityName="Jacob" authorityYear="1987" box="[898,1307,1303,1328]" class="Insecta" family="Baetidae" genus="Pseudocentroptiloides" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579C87FA95E27D0B63" box="[898,1147,1304,1328]" italics="true" pageId="7" pageNumber="490">Pseudocentroptiloides</emphasis>
|
||
Jacob 1987
|
||
</taxonomicName>
|
||
, which is characterized by autapomorphies in the larval mouth apparatus and can be treated either as a subgenus of the genus
|
||
<taxonomicName id="4C2484E1FFCA30579F92FAEDE70D0B2B" box="[151,267,1376,1400]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579F92FAEDE70D0B2B" box="[151,267,1376,1400]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
, or as a genus in the
|
||
<taxonomicName id="4C2484E1FFCA30579D15FAEDE4820B2B" box="[528,644,1376,1400]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579D15FAEDE4820B2B" box="[528,644,1376,1400]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
group of genera. Other representatives of
|
||
<taxonomicName id="4C2484E1FFCA30579B7DFAD2E2E90B2B" box="[1144,1263,1375,1400]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
|
||
/g1 constitute a plesiomorphon
|
||
<taxonomicName id="4C2484E1FFCA30579E40FA09E7BD0BCE" box="[325,443,1412,1437]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
|
||
/g2, which in rank-based nomenclature can be treated either as a genus
|
||
<taxonomicName id="4C2484E1FFCA30579BD7FA09E3400BCF" box="[1234,1350,1412,1436]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579BD7FA09E3400BCF" box="[1234,1350,1412,1436]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
, or as a subgenus
|
||
<taxonomicName id="4C2484E1FFCA30579E02FA25E77D0B93" box="[263,379,1448,1472]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579E02FA25E77D0B93" box="[263,379,1448,1472]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
(e.g.,
|
||
<bibRefCitation id="EFB58293FFCA30579EC1FA2AE4E70B93" author="Kluge" box="[452,737,1447,1472]" pageId="7" pageNumber="490" refString="Kluge, N. J. & Novikova, E. A. (1992) [Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR]. Entomologicheskoe Obozrenie, 71 (1), 38 - 55. [in Russian]" type="journal article" year="1992">Kluge & Novikova 1992</bibRefCitation>
|
||
); it includes both two-winged species originally placed to the genus
|
||
<taxonomicName id="4C2484E1FFCA30579FDAFA41E7550BB7" box="[223,339,1484,1508]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579FDAFA41E7550BB7" box="[223,339,1484,1508]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
, and species with hind wings, originally placed to the artificial genus
|
||
<taxonomicName id="4C2484E1FFCA30579B66FA41E2F10BB7" box="[1123,1271,1484,1508]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579B66FA41E2F10BB7" box="[1123,1271,1484,1508]" italics="true" pageId="7" pageNumber="490">Centroptilum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9BFF62FFCA30579FC2FA7DE4A20923" blockId="7.[151,1437,151,1904]" pageId="7" pageNumber="490">
|
||
Some authors (e.g.,
|
||
<bibRefCitation id="EFB58293FFCA30579EAFFA62E42D085B" author="Jacob" box="[426,555,1519,1544]" pageId="7" pageNumber="490" refString="Jacob, U. (1991) Ephemeroptera: Zur Systematic der europaischen Baetidae aus Gattungsbene. Verhandlungen der Westdeutschen Entomologischen Tagung, 1990, 271 - 290." type="journal article" year="1991">Jacob 1991</bibRefCitation>
|
||
) attribute to the genus
|
||
<taxonomicName id="4C2484E1FFCA30579C37FA7DE5A0085B" box="[818,934,1520,1544]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579C37FA7DE5A0085B" box="[818,934,1520,1544]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
, besides other species with hind wings, also
|
||
<taxonomicName id="4C2484E1FFCA30579F92F999E475087F" authority="Tshernova 1941" authorityName="Tshernova" authorityYear="1941" box="[151,627,1556,1581]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="unguiculatum">
|
||
<emphasis id="B9502370FFCA30579F92F999E79F087F" box="[151,409,1556,1580]" italics="true" pageId="7" pageNumber="490">Cloeon unguiculatum</emphasis>
|
||
Tshernova 1941
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C2484E1FFCA30579DBEF999E2C0087F" authority="Bogoescu 1947" authorityName="Bogoescu" authorityYear="1947" box="[699,1222,1556,1581]" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="motasi">
|
||
<emphasis id="B9502370FFCA30579DBEF999E5F8087F" box="[699,1022,1556,1580]" italics="true" pageId="7" pageNumber="490">Pseudocentroptilum motasi</emphasis>
|
||
Bogoescu 1947
|
||
</taxonomicName>
|
||
; =
|
||
<taxonomicName id="4C2484E1FFCA30579A0DF999E4000803" authority="Braasch & Soldan 1983" authorityName="Braasch & Soldan" authorityYear="1983" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="species" species="limnale">
|
||
<emphasis id="B9502370FFCA30579A0DF999E6ED0803" italics="true" pageId="7" pageNumber="490">Centroptilum limnale</emphasis>
|
||
Braasch & Soldan 1983
|
||
</taxonomicName>
|
||
), which is the
|
||
<typeStatus id="549F41C0FFCA30579DB7F9B5E4E40803" box="[690,738,1592,1616]" pageId="7" pageNumber="490">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C2484E1FFCA30579C63F9B5E3050803" authority="Bogoescu 1947" authorityName="Bogoescu" authorityYear="1947" box="[870,1283,1592,1616]" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579C63F9B5E2400803" box="[870,1094,1592,1616]" italics="true" pageId="7" pageNumber="490">Pseudocentroptilum</emphasis>
|
||
Bogoescu 1947
|
||
</taxonomicName>
|
||
. This species has no characters (1) and (2) and its relationship with true
|
||
<taxonomicName id="4C2484E1FFCA30579C30F9D1E5AF0827" box="[821,937,1628,1652]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579C30F9D1E5AF0827" box="[821,937,1628,1652]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
is questionable. They also divide the genus
|
||
<taxonomicName id="4C2484E1FFCA30579F92F90DE70D08CB" box="[151,267,1664,1688]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579F92F90DE70D08CB" box="[151,267,1664,1688]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
into two subgenera—
|
||
<taxonomicName id="4C2484E1FFCA30579D06F90DE47108CB" box="[515,631,1664,1688]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579D06F90DE47108CB" box="[515,631,1664,1688]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2484E1FFCA30579DA8F90DE58B08CB" box="[685,909,1664,1688]" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579DA8F90DE58B08CB" box="[685,909,1664,1688]" italics="true" pageId="7" pageNumber="490">Pseudocentroptilum</emphasis>
|
||
</taxonomicName>
|
||
, based on a single character: absence/presence of hind wings. However, absence of hind wings is not enough to prove holophyly of the subgenus
|
||
<taxonomicName id="4C2484E1FFCA30579A24F929E39308EF" box="[1313,1429,1700,1724]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579A24F929E39308EF" box="[1313,1429,1700,1724]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
</taxonomicName>
|
||
, because loss of hind wings independently took place in many taxa of
|
||
<taxonomicName id="4C2484E1FFCA30579CA6F94AE20F08B3" box="[931,1033,1735,1760]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
. Besides the genus
|
||
<taxonomicName id="4C2484E1FFCA30579BE6F945E7260957" authority="Jacob 1991" authorityName="Jacob" authorityYear="1991" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579BE6F945E35108B3" box="[1251,1367,1736,1760]" italics="true" pageId="7" pageNumber="490">Procloeon</emphasis>
|
||
sensu
|
||
<bibRefCitation id="EFB58293FFCA30579F92F961E7260957" author="Jacob" box="[151,288,1772,1797]" pageId="7" pageNumber="490" refString="Jacob, U. (1991) Ephemeroptera: Zur Systematic der europaischen Baetidae aus Gattungsbene. Verhandlungen der Westdeutschen Entomologischen Tagung, 1990, 271 - 290." type="journal article" year="1991">Jacob 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
, in many other recently accepted genera of
|
||
<taxonomicName id="4C2484E1FFCA30579C4EF961E5AB0956" box="[843,941,1772,1797]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
(e.g.,
|
||
<taxonomicName id="4C2484E1FFCA30579B00F961E39E0957" authority="Jacob 1987" authorityName="Jacob" authorityYear="1987" box="[1029,1432,1772,1797]" class="Insecta" family="Baetidae" genus="Pseudocentroptiloides" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579B00F961E2F80957" box="[1029,1278,1772,1796]" italics="true" pageId="7" pageNumber="490">Pseudocentroptiloides</emphasis>
|
||
Jacob 1987
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2484E1FFCA30579F92F89DE44F097B" authority="Novikova & Kluge 1987" authorityName="Novikova & Kluge" authorityYear="1987" box="[151,585,1807,1832]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579F92F89DE71D097B" box="[151,283,1808,1832]" italics="true" pageId="7" pageNumber="490">Labiobaetis</emphasis>
|
||
Novikova & Kluge 1987
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2484E1FFCA30579D52F89DE5AF097B" authority="Kazlauskas 1987" authorityName="Kazlauskas" authorityYear="1987" box="[599,937,1807,1832]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579D52F89DE4DC097B" box="[599,730,1808,1832]" italics="true" pageId="7" pageNumber="490">Nigrobaetis</emphasis>
|
||
Kazlauskas 1987
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2484E1FFCA30579CBDF89DE2BC097B" authority="Traver 1938" authorityName="Traver" authorityYear="1938" box="[952,1210,1808,1832]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579CBDF89DE227097B" box="[952,1057,1808,1832]" italics="true" pageId="7" pageNumber="490">Cloeodes</emphasis>
|
||
Traver 1938
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2484E1FFCA30579BCCF89DE6D4091E" authority="Day 1955" authorityName="Day" authorityYear="1955" class="Insecta" family="Baetidae" genus="Paracloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579BCCF89DE365097B" box="[1225,1379,1808,1832]" italics="true" pageId="7" pageNumber="490">Paracloeodes</emphasis>
|
||
Day 1955
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2484E1FFCA30579FD9F8B9E40D091F" authority="Bengtsson 1912" authorityName="Bengtsson" authorityYear="1912" box="[220,523,1844,1869]" class="Insecta" family="Baetidae" genus="Acentrella" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="490" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502370FFCA30579FD9F8B9E756091F" box="[220,336,1844,1868]" italics="true" pageId="7" pageNumber="490">Acentrella</emphasis>
|
||
Bengtsson 1912
|
||
</taxonomicName>
|
||
) hind wings are retained in some species, being lost in others. In many other nonrelated baetid taxa the hind wings also are lost.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |