272 lines
33 KiB
XML
272 lines
33 KiB
XML
<document id="5E246259D498E04442520D7109E780E3" ID-CLB-Dataset="86219" ID-DOI="10.11646/zootaxa.5159.3.3" ID-GBIF-Dataset="1f624c93-2d62-44b0-8867-ae3d15ee5239" ID-ISSN="1175-5326" ID-Zenodo-Dep="6781621" ID-ZooBank="08C32EB5-66E5-4949-AFF3-7F66B7F42E14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656566258139" checkinUser="plazi" docAuthor="Momtazi, Farzaneh" docDate="2022" docId="038D47195B185F1F0498FA2B9C9E91E0" docLanguage="en" docName="zootaxa.5159.3.3.pdf" docOrigin="Zootaxa 5159 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cymadusa makranica Momtazi 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="373" masterDocId="FFB43F615B1D5F19040FFF829809963D" masterDocTitle="Ampithoidae (Crustacea: Amphipoda) from the Persian Gulf and the Gulf of Oman" masterLastPageNumber="382" masterPageNumber="367" pageNumber="372" updateTime="1699353675599" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="8FF31BC733D14E70607CDB2E4638E1A6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="565E5433F29132F6EB7254EA5A383DFF">
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<mods:title id="010F19402C97FFC2D03BDCB894A72773">Ampithoidae (Crustacea: Amphipoda) from the Persian Gulf and the Gulf of Oman</mods:title>
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</mods:titleInfo>
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<mods:name id="2314AF833D09E9FBB34430073FCE172C" type="personal">
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<mods:role id="F98982B0EF98D12A23019A83AEC0891F">
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<mods:roleTerm id="1FDDBDB927944BD4568F20681A290C3A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DD267D3A092FC475BA071E408C1BB717">Momtazi, Farzaneh</mods:namePart>
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<mods:nameIdentifier id="8444BFBA299C45025C5DE14DD6861809" type="ORCID">0000-0002-3074-0865</mods:nameIdentifier>
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<mods:affiliation id="16E450B38AC66A682CF0290D945D068C">Iranian National Institute for Oceanography and Atmospheric Science (INIOAS), Marine Bioscience Dept. Tehran, Iran. Corresponding author. momtazi. f @ gmail. com; momtazi. f @ inio. ac. ir; https: // orcid. org / 0000 - 0002 - 3074 - 0865</mods:affiliation>
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<mods:nameIdentifier id="9998EC692FC2FC9CD928579C8183B35E" type="email">momtazi.f@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="4D84C3EFE742D33B04C953E90A03F3CC">text</mods:typeOfResource>
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<mods:title id="F4BC2F31B49337A507DD71E2B52D64EE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A252E01554E3EB65940696A7DE522E70">
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<mods:date id="C796B942853DE2F416B32707A80254F0">2022</mods:date>
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<mods:number id="F59C43C4AFBB814A4CE74580E1ABD834">2022-06-30</mods:number>
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<mods:detail id="D0A8E88550A37308B6AD7B1F7819CC14" type="volume">
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<mods:number id="14FFCF4F20894C0C988B4843DD676A33">5159</mods:number>
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</mods:detail>
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<mods:detail id="07278697EDCCA047D52CC6B3877E447C" type="issue">
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<mods:number id="063203F516808DCE60BAF09B098A9CCC">3</mods:number>
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<mods:start id="1A3AFC970F1B2EA2450183B225C3E18C">367</mods:start>
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<mods:end id="4ED0E72ED64B7B8DE1F2510281BCB6B3">382</mods:end>
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<mods:classification id="B1A4C80A96081C48BBA2C738F4D97A2D">journal article</mods:classification>
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<mods:identifier id="14789AE8658CAECD254560929517F14C" type="CLB-Dataset">86219</mods:identifier>
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<mods:identifier id="C521BD0E50F873610FBFAA152AFEC545" type="DOI">10.11646/zootaxa.5159.3.3</mods:identifier>
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<mods:identifier id="5BEA16C12AEB7D1079CFCCAD04E97965" type="GBIF-Dataset">1f624c93-2d62-44b0-8867-ae3d15ee5239</mods:identifier>
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<mods:identifier id="08C1FE1DE0B08E8D806CC633C1955C32" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A098839F435C7C5DA70C6E4D974DB112" type="Zenodo-Dep">6781621</mods:identifier>
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</mods:mods>
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<treatment id="038D47195B185F1F0498FA2B9C9E91E0" ID-DOI="http://doi.org/10.5281/zenodo.6781633" ID-GBIF-Taxon="196373219" ID-Zenodo-Dep="6781633" LSID="urn:lsid:plazi:treatment:038D47195B185F1F0498FA2B9C9E91E0" httpUri="http://treatment.plazi.org/id/038D47195B185F1F0498FA2B9C9E91E0" lastPageId="6" lastPageNumber="373" pageId="5" pageNumber="372">
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<subSubSection id="C33EA5845B185F1C0498FA2B9A0C93FE" box="[151,517,1449,1476]" pageId="5" pageNumber="372" type="nomenclature">
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<paragraph id="8B9BF60F5B185F1C0498FA2B9A0C93FE" blockId="5.[151,517,1449,1512]" box="[151,517,1449,1476]" pageId="5" pageNumber="372">
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<heading id="D0D341635B185F1C0498FA2B9A0C93FE" bold="true" box="[151,517,1449,1476]" fontSize="11" level="1" pageId="5" pageNumber="372" reason="1">
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<emphasis id="B9502A1D5B185F1C0498FA2B9A0C93FE" bold="true" box="[151,517,1449,1476]" pageId="5" pageNumber="372">
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<taxonomicName id="4C248D8C5B185F1C0498FA2B99A993F9" ID-CoL="ea629cff-9a6c-47f9-bee9-555f5f7202ca" authority="Momtazi, 2022" authorityName="Momtazi" authorityYear="2022" box="[151,416,1449,1476]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="372" phylum="Arthropoda" rank="species" species="makranica" status="sp. nov.">
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<emphasis id="B9502A1D5B185F1C0498FA2B99A993F9" bold="true" box="[151,416,1449,1476]" italics="true" pageId="5" pageNumber="372">Cymadusa makranica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26397665B185F1C05A8FA2B9A0C93FE" box="[423,517,1449,1475]" pageId="5" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33EA5845B185F1C0498FA4F992693D5" box="[151,303,1485,1512]" pageId="5" pageNumber="372" type="description">
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<paragraph id="8B9BF60F5B185F1C0498FA4F992693D5" blockId="5.[151,517,1449,1512]" box="[151,303,1485,1512]" pageId="5" pageNumber="372">
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||
(
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<figureCitation id="131FEA8A5B185F1C0490FA4F992E93D5" box="[159,295,1485,1512]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[151,250,1943,1967]" captionStartId-1="8.[151,250,1943,1967]" captionTargetBox-0="[165,1423,181,1919]" captionTargetBox-1="[230,1358,181,1913]" captionTargetId-0="figure-15@7.[165,1423,181,1920]" captionTargetId-1="figure-29@8.[230,1358,181,1920]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Cymadusa makranica sp. nov. male, 10.8 mm, (INIOC2–14S), Djod, Gulf of Oman." captionText-1="FIGURE 6. Cymadusa makranica sp. nov. male, 10.8 mm, (INIOC2–14S), Djod, Gulf of Oman." figureDoi-0="http://doi.org/10.5281/zenodo.6781639" figureDoi-1="http://doi.org/10.5281/zenodo.6781641" httpUri-0="https://zenodo.org/record/6781639/files/figure.png" httpUri-1="https://zenodo.org/record/6781641/files/figure.png" pageId="5" pageNumber="372">Figures 5–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33EA5845B185F1C0498F9979A0290A6" pageId="5" pageNumber="372" type="materials_examined">
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<paragraph id="8B9BF60F5B185F1C0498F9979A8F904A" blockId="5.[151,1437,1557,2016]" pageId="5" pageNumber="372">
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<emphasis id="B9502A1D5B185F1C0498F99799719012" bold="true" box="[151,376,1557,1583]" pageId="5" pageNumber="372">Material examined</emphasis>
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.
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<materialsCitation id="3B4CFC525B185F1C058BF99799D89069" ID-GBIF-Occurrence="3821896303" collectingDate="2015-05" collectionCode="INIOC" country="Oman" latitude="25.45" location="Djod" longLatPrecision="1" longitude="59.51" municipality="Gulf" pageId="5" pageNumber="372" specimenCode="INIOC2-14" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<typeStatus id="549F48AD5B185F1C058BF99799E69012" box="[388,495,1557,1583]" pageId="5" pageNumber="372" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="9D223D865B185F1C05F6F9979A4C9012" box="[505,581,1557,1583]" pageId="5" pageNumber="372" type="male">1 male</specimenCount>
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||
,
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||
<quantity id="4CDC5BEA5B185F1C0640F9979AB8900D" box="[591,689,1557,1584]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="5" pageNumber="372" unit="mm" value="10.8">10.8 mm</quantity>
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||
(
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<specimenCode id="DB825E745B185F1C06B1F9979B4D9012" box="[702,836,1557,1583]" collectionCode="INIOC" pageId="5" pageNumber="372">INIOC2-14</specimenCode>
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S) intertidal zone,
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<location id="8EFBA0D45B185F1C0005F9979C4C9012" LSID="urn:lsid:plazi:treatment:038D47195B185F1F0498FA2B9C9E91E0:8EFBA0D45B185F1C0005F9979C4C9012" box="[1034,1093,1557,1583]" country="Oman" latitude="25.45" longLatPrecision="1" longitude="59.51" municipality="Gulf" name="Djod" pageId="5" pageNumber="372">Djod</location>
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,
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||
<collectingMunicipality id="6BFF6C755B185F1C0040F9979C8B9012" box="[1103,1154,1557,1583]" pageId="5" pageNumber="372">Gulf</collectingMunicipality>
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||
of
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||
<collectingCountry id="F333B69F5B185F1C00ABF9979CE19012" box="[1188,1256,1557,1583]" name="Oman" pageId="5" pageNumber="372">Oman</collectingCountry>
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(
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<geoCoordinate id="EE1090C85B185F1C00F9F9979D91900D" box="[1270,1432,1557,1584]" degrees="25" direction="north" minutes="27" orientation="latitude" pageId="5" pageNumber="372" precision="1" seconds="0.00" value="25.45">25°27’0.00”N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE1090C85B185F1C0498F9BB994D9069" box="[151,324,1593,1620]" degrees="59" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="372" precision="1" seconds="36.00" value="59.51">59°30’36.00”E</geoCoordinate>
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||
),
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||
<date id="FF9AD0CF5B185F1C0556F9B899D89069" box="[345,465,1593,1620]" pageId="5" pageNumber="372" value="2015-05">
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||
<collectingDate id="EFDE29275B185F1C0556F9B899D89069" box="[345,465,1593,1620]" pageId="5" pageNumber="372" value="2015-05">May 2015</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B4CFC525B185F1C05D3F9B89A88904A" ID-GBIF-Occurrence="3821896310" collectingDate="2011-05" collectionCode="ZUTC" country="Oman" latitude="25.45" location="Djod" longLatPrecision="1" longitude="59.51" municipality="Gulf" pageId="5" pageNumber="372" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<typeStatus id="549F48AD5B185F1C05D3F9B89A489069" box="[476,577,1594,1620]" pageId="5" pageNumber="372" type="paratype">Paratype</typeStatus>
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||
,
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||
<specimenCount id="9D223D865B185F1C0641F9BB9A94906E" box="[590,669,1593,1619]" pageId="5" pageNumber="372" type="male">1 male</specimenCount>
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||
,
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||
<quantity id="4CDC5BEA5B185F1C06A5F9BB9AF29069" box="[682,763,1593,1620]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="372" unit="mm" value="10.0">10 mm</quantity>
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||
(
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||
<collectionCode id="ED356ECA5B185F1C0705F9BB9B5C906E" box="[778,853,1593,1619]" pageId="5" pageNumber="372">ZUTC</collectionCode>
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||
Amph. 2365), intertidal zone,
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||
<location id="8EFBA0D45B185F1C00BAF9BB9CF9906E" LSID="urn:lsid:plazi:treatment:038D47195B185F1F0498FA2B9C9E91E0:8EFBA0D45B185F1C00BAF9BB9CF9906E" box="[1205,1264,1593,1619]" country="Oman" latitude="25.45" longLatPrecision="1" longitude="59.51" municipality="Gulf" name="Djod" pageId="5" pageNumber="372">Djod</location>
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||
,
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||
<collectingMunicipality id="6BFF6C755B185F1C00F3F9BB9D26906E" box="[1276,1327,1593,1619]" pageId="5" pageNumber="372">Gulf</collectingMunicipality>
|
||
of
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||
<collectingCountry id="F333B69F5B185F1C0157F9BB9D95906E" box="[1368,1436,1593,1619]" name="Oman" pageId="5" pageNumber="372">Oman</collectingCountry>
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||
(
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||
<geoCoordinate id="EE1090C85B185F1C0490F9DF994B9045" box="[159,322,1629,1656]" degrees="25" direction="north" minutes="27" orientation="latitude" pageId="5" pageNumber="372" precision="1" seconds="0.00" value="25.45">25°27’0.00”N</geoCoordinate>
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,
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||
<geoCoordinate id="EE1090C85B185F1C0542F9DF99F39045" box="[333,506,1629,1656]" degrees="59" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="372" precision="1" seconds="36.00" value="59.51">59°30’36.00”E</geoCoordinate>
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||
),
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||
<date id="FF9AD0CF5B185F1C0602F9DC9A88904A" box="[525,641,1629,1656]" pageId="5" pageNumber="372" value="2011-05">
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||
<collectingDate id="EFDE29275B185F1C0602F9DC9A88904A" box="[525,641,1629,1656]" pageId="5" pageNumber="372" value="2011-05">May 2011</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="8B9BF60F5B185F1C04C8F9039A0290A6" blockId="5.[151,1437,1557,2016]" box="[199,523,1665,1692]" pageId="5" pageNumber="372">
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||
<materialsCitation id="3B4CFC525B185F1C04C8F9039A0290A6" ID-GBIF-Occurrence="3821896311" box="[199,523,1665,1692]" country="Oman" location="Gulf" pageId="5" pageNumber="372" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B9502A1D5B185F1C04C8F903996990A6" bold="true" box="[199,352,1665,1691]" pageId="5" pageNumber="372">
|
||
<typeStatus id="549F48AD5B185F1C04C8F903990B90A6" box="[199,258,1665,1691]" pageId="5" pageNumber="372">Type</typeStatus>
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||
locality
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||
</emphasis>
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||
.
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||
<location id="8EFBA0D45B185F1C0561F90399A890A6" LSID="urn:lsid:plazi:treatment:038D47195B185F1F0498FA2B9C9E91E0:8EFBA0D45B185F1C0561F90399A890A6" box="[366,417,1665,1691]" country="Oman" name="Gulf" pageId="5" pageNumber="372">Gulf</location>
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||
of
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||
<collectingCountry id="F333B69F5B185F1C05C8F9039A0290A6" box="[455,523,1665,1691]" name="Oman" pageId="5" pageNumber="372">Oman</collectingCountry>
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||
</materialsCitation>
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C33EA5845B185F1C04C8F9279C169149" pageId="5" pageNumber="372" type="diagnosis">
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||
<paragraph id="8B9BF60F5B185F1C04C8F9279C169149" blockId="5.[151,1437,1557,2016]" pageId="5" pageNumber="372">
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||
<emphasis id="B9502A1D5B185F1C04C8F927994B9082" bold="true" box="[199,322,1701,1727]" pageId="5" pageNumber="372">Diagnosis.</emphasis>
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Antenna 1 accessory flagellum with two article. Maxilla 1 inner plate with 4 long and three short slender setae. Coxa 1 not anteriorly round, gnathopod 1 with sparsely setose margins; merus produced to form a small, subacute distoventral lobe; carpus as long as propodus; palm acute, entire, with 1 defining robust seta. Gnathopod 2 palm defining with large tooth and small tooth in the middle that separated each other with cleft; dactylus shorter than palm. Epimeron 3 posteroventral corner rounded. Uropod 3 peduncle with 6 distal robust setae. Telson distally truncated, apical cusps small, with multi apical and medial slender setae.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C33EA5845B185F1F04C8F8FF9991930C" lastPageId="6" lastPageNumber="373" pageId="5" pageNumber="372" type="description">
|
||
<paragraph id="8B9BF60F5B185F1C04C8F8FF9B8591AA" blockId="5.[151,1437,1557,2016]" box="[199,908,1917,1944]" pageId="5" pageNumber="372">
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||
<emphasis id="B9502A1D5B185F1C04C8F8FF995891AA" bold="true" box="[199,337,1917,1943]" pageId="5" pageNumber="372">Description</emphasis>
|
||
. Based on
|
||
<typeStatus id="549F48AD5B185F1C05C1F8FF9A3A91AA" box="[462,563,1917,1943]" pageId="5" pageNumber="372" type="holotype">holotype</typeStatus>
|
||
, male,
|
||
<quantity id="4CDC5BEA5B185F1C068DF8FF9AE191A5" box="[642,744,1917,1944]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="5" pageNumber="372" unit="mm" value="10.8">10.8 mm</quantity>
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, INIOC2-14S.
|
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</paragraph>
|
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<paragraph id="8B9BF60F5B185F1F04C8F8239D7A97B4" blockId="5.[151,1437,1557,2016]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="373" pageId="5" pageNumber="372">
|
||
<emphasis id="B9502A1D5B185F1C04C8F823990E9186" bold="true" box="[199,263,1953,1979]" pageId="5" pageNumber="372">Head</emphasis>
|
||
. Head as long as deep; eyes poorly developed.
|
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<emphasis id="B9502A1D5B185F1C0701F8239B8F9186" box="[782,902,1953,1979]" italics="true" pageId="5" pageNumber="372">Antenna 1,</emphasis>
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slightly longer than second antenna; peduncular article 1 subequal to article 2, second article longer than third one; flagellum with 42 articles; accessory flagellum 2–articulate.
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<emphasis id="B9502A1D5B1B5F1F0525FF1599AD968C" box="[298,420,151,177]" italics="true" pageId="6" pageNumber="373">Antenna 2,</emphasis>
|
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peduncle without brush setae; fourth article longest; flagellum with 36 articles.
|
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<emphasis id="B9502A1D5B1B5F1F0120FF159D94968C" box="[1327,1437,151,177]" italics="true" pageId="6" pageNumber="373">Upper lip</emphasis>
|
||
distally setose. Lower lip, outer plates notched forming a deep distal cleft.
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<emphasis id="B9502A1D5B1B5F1F07DFFF399C3296E8" box="[976,1083,187,213]" italics="true" pageId="6" pageNumber="373">Mandible</emphasis>
|
||
, molar well developed, triturating, accessory setal row with 7 serrate setae; palp 3–articulate, article 1 shorter than article 2; article 2 with 1 distal seta; article 3 the longest, with 9 apical and two marginal setae.
|
||
<emphasis id="B9502A1D5B1B5F1F076EFE819BDC9720" box="[865,981,259,285]" italics="true" pageId="6" pageNumber="373">Maxilla 1,</emphasis>
|
||
inner plate with 4 distal and three proximal setae; outer plate with 9 dentate setae; palp well developed, two articulate, distal article apically with 10 short robust and 4 long setae.
|
||
<emphasis id="B9502A1D5B1B5F1F05A4FEC99A179758" box="[427,542,331,357]" italics="true" pageId="6" pageNumber="373">Maxilla 2,</emphasis>
|
||
inner plate narrower than outer plate. Maxilliped, outer plate with row of large robust setae along medial margin, inner plate developed, not reach to end of first palp article, palp 4–articulate.
|
||
</paragraph>
|
||
<paragraph id="8B9BF60F5B1B5F1F04C8FE1199A4922C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="373">
|
||
<emphasis id="B9502A1D5B1B5F1F04C8FE1199139790" bold="true" box="[199,282,403,429]" pageId="6" pageNumber="373">Pereon</emphasis>
|
||
.
|
||
<emphasis id="B9502A1D5B1B5F1F0526FE1199B49790" box="[297,445,403,429]" italics="true" pageId="6" pageNumber="373">Gnathopod 1</emphasis>
|
||
; smaller than gnathopod 2; coxa 1 not anteriorly produced; basis longer than propodus, with 6 long proximal setae on the posterior margin, anterodistal lobe present with one seta; ischium small, trapezoidshaped with multi setae on the posterior margin; merus anterodistal lobe acute; carpus as long as propodus with one defining large robust seta; dactylus subequal in length to palm, inner margin crenate; propodus subquadrate, length 1.9 × width, palm acute.
|
||
<emphasis id="B9502A1D5B1B5F1F05A3FDA19A369400" box="[428,575,547,573]" italics="true" pageId="6" pageNumber="373">Gnathopod 2</emphasis>
|
||
; coxa as long as deep; basis without anterodistal lobe, shorter than propodus, with 5 long proximal setae on the posterior margin; ischium trapezoid with one posterodistal seta; merus subrectangular, anterior margin naked; carpus as long as merus, shorter than propodus, subtriangular, with small posterodistal lobe, anterior margin with one seta; propodus 1.5 × width, subrectangular, anterior margin without seta, posterior margin straight, palm defining with large tooth and small tooth in the middle that separated each other with cleft; dactylus shorter than palm.
|
||
<emphasis id="B9502A1D5B1B5F1F05DCFD559A7894CC" box="[467,625,727,753]" italics="true" pageId="6" pageNumber="373">Pereopod 3–4</emphasis>
|
||
similar, basis slightly expanded with multi long setae on both margins, ischium trapezoid; merus subequal to carpus; propodus with multi slender setae on the posterior margin.
|
||
<emphasis id="B9502A1D5B1B5F1F013FFD7998AC9504" italics="true" pageId="6" pageNumber="373">Pereopod 5</emphasis>
|
||
, coxa with multi surface setae; basis expanded; merus slightly longer than carpus; carpus subquadrate; propodus subrectangular with four robust setae on posterior margin; dactylus strongly curved.
|
||
<emphasis id="B9502A1D5B1B5F1F0030FCC19CB69561" box="[1087,1215,834,861]" italics="true" pageId="6" pageNumber="373">Pereopod 6</emphasis>
|
||
similar to pereopod 7, slightly shorter than P7, basis slightly expanded with two posterodistal and three anterodistal robust setae; merus slightly longer than carpus; carpus subquadrate; propodus subrectangular with five robust setae on posterior margin; dactylus curved.
|
||
<emphasis id="B9502A1D5B1B5F1F0554FC2D99D695F4" box="[347,479,943,969]" italics="true" pageId="6" pageNumber="373">Pereopod 7</emphasis>
|
||
, basis slightly expanded with two posterodistal and three anterodistal robust setae; merus slightly longer than carpus; carpus subquadrate; propodus subrectangular with six robust setae on posterior margin; dactylus curved.
|
||
</paragraph>
|
||
<paragraph id="8B9BF60F5B1B5F1F04C8FB999D1D9330" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="373">
|
||
<emphasis id="B9502A1D5B1B5F1F04C8FB9999039208" bold="true" box="[199,266,1051,1077]" pageId="6" pageNumber="373">Pleon</emphasis>
|
||
.
|
||
<emphasis id="B9502A1D5B1B5F1F0517FB99998A9208" box="[280,387,1051,1077]" italics="true" pageId="6" pageNumber="373">Uropod 1</emphasis>
|
||
; peduncle with nine robust setae, with large interannal spur, setal fringe absent; rami unequal, shorter than peduncle, outer ramus larger with 11 marginal robust setae and four apical setae; inner ramus with 13 marginal and four apical robust setae.
|
||
<emphasis id="B9502A1D5B1B5F1F0664FBE19ADE9240" box="[619,727,1123,1149]" italics="true" pageId="6" pageNumber="373">Uropod 2</emphasis>
|
||
, peduncle larger than outer ramus, with five robust setae, setal fringe absent; rami unequal, outer ramus shorter than inner ramus with eight marginal and three apical robust setae; inner ramus with nine marginal and five apical robust setae.
|
||
<emphasis id="B9502A1D5B1B5F1F074EFB299BA492F8" box="[833,941,1195,1221]" italics="true" pageId="6" pageNumber="373">Uropod 3</emphasis>
|
||
, peduncle longer than broad; peduncle with six distal robust setae, inner ramus 0.6 × peduncle, with four apical and one marginal robust setae and some long simple setae; outer ramus with 2 apical large curved robust setae. Telson, trapezoid, apically truncated.
|
||
</paragraph>
|
||
<paragraph id="8B9BF60F5B1B5F1F04C8FA959991930C" blockId="6.[151,1437,151,2013]" box="[199,408,1303,1329]" pageId="6" pageNumber="373">Female, unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33EA5845B1B5F1F04C8FAB9992C9344" pageId="6" pageNumber="373" type="etymology">
|
||
<paragraph id="8B9BF60F5B1B5F1F04C8FAB9992C9344" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="373">
|
||
<emphasis id="B9502A1D5B1B5F1F04C8FAB999419368" bold="true" box="[199,328,1339,1365]" pageId="6" pageNumber="373">Etymology</emphasis>
|
||
. The term “makranica” is referred to the ancient name of the
|
||
<typeStatus id="549F48AD5B1B5F1F002AFAB99C5C9368" box="[1061,1109,1339,1365]" pageId="6" pageNumber="373">type</typeStatus>
|
||
locality, the Gulf of
|
||
<collectingCountry id="F333B69F5B1B5F1F015EFAB99D919368" box="[1361,1432,1339,1365]" name="Oman" pageId="6" pageNumber="373">Oman</collectingCountry>
|
||
, Makran Sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33EA5845B1B5F1F04C8FA019C9E91E0" pageId="6" pageNumber="373" type="discussion">
|
||
<paragraph id="8B9BF60F5B1B5F1F04C8FA019942906C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="373">
|
||
<emphasis id="B9502A1D5B1B5F1F04C8FA01993D93A0" bold="true" box="[199,308,1411,1437]" pageId="6" pageNumber="373">Remarks</emphasis>
|
||
. The genus
|
||
<taxonomicName id="4C248D8C5B1B5F1F05CEFA019AEB93A0" authority="Savigny, 1816" authorityName="Savigny" authorityYear="1816" box="[449,738,1411,1437]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="373" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502A1D5B1B5F1F05CEFA019A3E93A0" box="[449,567,1411,1437]" italics="true" pageId="6" pageNumber="373">Cymadusa</emphasis>
|
||
<bibRefCitation id="EFB58BFE5B1B5F1F064FFA019AEB93A0" author="Savigny, J. C." box="[576,738,1411,1437]" pageId="6" pageNumber="373" pagination="39 - 117" refId="ref6244" refString="Savigny, J. C. (1816) Observations generales sur la bouche des arachnides, des crustaces et des entomostraces. Memories sur les Animaux sans Vertebres, 1816, 39 - 117. https: // doi. org / 10.5962 / bhl. title. 125538" type="journal article" year="1816">Savigny, 1816</bibRefCitation>
|
||
</taxonomicName>
|
||
was described from the
|
||
<collectingCountry id="F333B69F5B1B5F1F07F3FA019C4A93A0" box="[1020,1091,1411,1437]" name="Egypt" pageId="6" pageNumber="373">Egypt</collectingCountry>
|
||
.
|
||
<bibRefCitation id="EFB58BFE5B1B5F1F0040FA019D9593A0" author="Barnard, J. L. & Karaman, G. S." box="[1103,1436,1411,1437]" pageId="6" pageNumber="373" pagination="866" refId="ref4490" refString="Barnard, J. L. & Karaman, G. S. (1991) The families and genera of marine gammaridean Amphipoda (except marine gammaroids). Records of the Australian Museum, Supplement 13, 1 D 866. https: // doi. org / 10.3853 / j. 0812 - 7387.13.1991.367" type="journal article" year="1991">Barnard and Karaman (1991)</bibRefCitation>
|
||
diagnose
|
||
<taxonomicName id="4C248D8C5B1B5F1F050DFA25997193FC" authorityName="Savigny" authorityYear="1816" box="[258,376,1447,1473]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="373" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502A1D5B1B5F1F050DFA25997193FC" box="[258,376,1447,1473]" italics="true" pageId="6" pageNumber="373">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
as having a 1–2-articulate accessory flagellum.
|
||
<bibRefCitation id="EFB58BFE5B1B5F1F0799FA259C1593FC" author="Peart, R. A." box="[918,1052,1447,1473]" pageId="6" pageNumber="373" pagination="301 - 336" refId="ref5828" refString="Peart, R. A. (2004) A revision of the Cymadusa filosa complex (Crustacea: Amphipoda: Corophioidea: Ampithoidae). Journal of Natural History, 38, 301 - 336. https: // doi. org / 10.1080 / 0022293021000055441" type="journal article" year="2004">Peart (2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB58BFE5B1B5F1F0028FA259C6893FC" author="Peart, R. A." box="[1063,1121,1447,1473]" pageId="6" pageNumber="373" pagination="1 - 95" refId="ref5995" refString="Peart, R. A. (2007) A review of the Australian species of Ampithoe Leach, 1814 (Crustacea: Amphipoda: mpithoidae) with descriptions of seventeen new species. Zootaxa, 1566 (1), 1 - 95. https: // doi. org / 10.11646 / zootaxa. 1566.1.1" type="journal article" year="2007">2007</bibRefCitation>
|
||
) re-evaluated the genus and prepared a detailed description differentiating it from alliance genera based on the presence of 1–3 articulated accessory flagellum and the shape of third uropod.
|
||
<taxonomicName id="4C248D8C5B1B5F1F0694FA6D9B189034" authorityName="Savigny" authorityYear="1816" box="[667,785,1519,1545]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="373" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502A1D5B1B5F1F0694FA6D9B189034" box="[667,785,1519,1545]" italics="true" pageId="6" pageNumber="373">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
was known as the tropical genus (
|
||
<bibRefCitation id="EFB58BFE5B1B5F1F0099FA6D9D1B9034" author="Peart, R. A." box="[1174,1298,1519,1545]" pageId="6" pageNumber="373" pagination="1 - 95" refId="ref5995" refString="Peart, R. A. (2007) A review of the Australian species of Ampithoe Leach, 1814 (Crustacea: Amphipoda: mpithoidae) with descriptions of seventeen new species. Zootaxa, 1566 (1), 1 - 95. https: // doi. org / 10.11646 / zootaxa. 1566.1.1" type="journal article" year="2007">Peart 2007</bibRefCitation>
|
||
) composing of 52 species (Horton
|
||
<emphasis id="B9502A1D5B1B5F1F0598F99199C39010" box="[407,458,1555,1581]" italics="true" pageId="6" pageNumber="373">et al</emphasis>
|
||
., 2020). However, the assumption of being monophyletic was debunked by
|
||
<bibRefCitation id="EFB58BFE5B1B5F1F013EF991994F906C" author="Peart, R. A. & Ahyong, S. T." pageId="6" pageNumber="373" pagination="456 - 474" refId="ref5934" refString="Peart, R. A. & Ahyong, S. T. (2016) Phylogenetic analysis of the Family Ampithoidae (Crustacea: Amphipoda), with a synopsis of the genera. Journal of Crustacean Biology, 36 (4), 456 - 474. https: // doi. org / 10.1163 / 1937240 X- 00002449" type="journal article" year="2016">Peart and Ahyong (2016)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9BF60F5B1B5F1F04C8F9D999619184" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="373">
|
||
The new species
|
||
<taxonomicName id="4C248D8C5B1B5F1F0587F9D99A779048" authorityName="Momtazi" authorityYear="2022" box="[392,638,1627,1653]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="373" phylum="Arthropoda" rank="species" species="makranica">
|
||
<emphasis id="B9502A1D5B1B5F1F0587F9D99A779048" box="[392,638,1627,1653]" italics="true" pageId="6" pageNumber="373">Cymadusa makranica</emphasis>
|
||
</taxonomicName>
|
||
is clearly distinguishable from other species with unique shape of propodus of second male gnathopod, presence of two set of setae on inner plate of first maxilla, bearing two articulated accessory flagellum, the first gnathopod with carpus as long as propodus and less setae on the both antennae. This species could be characterized from other species of
|
||
<taxonomicName id="4C248D8C5B1B5F1F06FEF9459B6E90DC" authorityName="Savigny" authorityYear="1816" box="[753,871,1735,1761]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="373" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9502A1D5B1B5F1F06FEF9459B6E90DC" box="[753,871,1735,1761]" italics="true" pageId="6" pageNumber="373">Cymadusa</emphasis>
|
||
</taxonomicName>
|
||
in the studied area with less setation in antennae, gnathopods and coxa1–4 and bearing first gnathopods with carpus as long as propodus. The new species is most similar to
|
||
<taxonomicName id="4C248D8C5B1B5F1F0500F88D9AE29114" authority="Rabindranath, 1972" authorityName="Rabindranath" authorityYear="1972" box="[271,747,1807,1833]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="373" phylum="Arthropoda" rank="species" species="imbroglio">
|
||
<emphasis id="B9502A1D5B1B5F1F0500F88D99F49114" box="[271,509,1807,1833]" italics="true" pageId="6" pageNumber="373">Cymadusa imbroglio</emphasis>
|
||
<bibRefCitation id="EFB58BFE5B1B5F1F0608F88D9AE29114" author="Rabindranath, P." box="[519,747,1807,1833]" pageId="6" pageNumber="373" pagination="161 - 178" refId="ref6052" refString="Rabindranath, P. (1972) Marine Gammaridea (Crustacea: Amphipoda) from the Indian region. Family Ampithoidae. Marine Biology, 14 (2), 161 - 178. https: // doi. org / 10.1007 / BF 00373215" type="journal article" year="1972">Rabindranath, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="4C248D8C5B1B5F1F0760F88D9C029114" authorityName="Momtazi" authorityYear="2022" box="[879,1035,1807,1833]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="makranica">
|
||
C.
|
||
<emphasis id="B9502A1D5B1B5F1F079CF88D9C029114" box="[915,1035,1807,1833]" italics="true" pageId="6" pageNumber="373">makranica</emphasis>
|
||
</taxonomicName>
|
||
could be differentiated by two articulated accessory flagellum in the first antennae (one articulate in
|
||
<taxonomicName id="4C248D8C5B1B5F1F0795F8B19C209170" authorityName="Rabindranath" authorityYear="1972" box="[922,1065,1843,1869]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="373" phylum="Arthropoda" rank="species" species="imbroglio">
|
||
<emphasis id="B9502A1D5B1B5F1F0795F8B19C209170" box="[922,1065,1843,1869]" italics="true" pageId="6" pageNumber="373">C. imbroglio</emphasis>
|
||
</taxonomicName>
|
||
), 4+3 setae on inner plate of the first maxilla (5 setae in
|
||
<emphasis id="B9502A1D5B1B5F1F05AAF8D59A4D914C" box="[421,580,1879,1905]" italics="true" pageId="6" pageNumber="373">
|
||
<taxonomicName id="4C248D8C5B1B5F1F05AAF8D59A31914C" authorityName="Rabindranath" authorityYear="1972" box="[421,568,1879,1905]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="373" phylum="Arthropoda" rank="species" species="imbroglio">C. imbroglio</taxonomicName>
|
||
),
|
||
</emphasis>
|
||
the shape of propodus in the male gnathopod 2, presence of fewer setae on uropods, having longer third article than second one in the mandibular palp, and absence of protuberance on palm of first gnathopod.
|
||
</paragraph>
|
||
<paragraph id="8B9BF60F5B1B5F1F04C8F8419C9E91E0" blockId="6.[151,1437,151,2013]" box="[199,1175,1987,2013]" pageId="6" pageNumber="373">
|
||
In spite of multi sampling in Djod station, only 2 samples of
|
||
<taxonomicName id="4C248D8C5B1B5F1F077EF8419C0391E0" authorityName="Momtazi" authorityYear="2022" box="[881,1034,1987,2013]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="380" phylum="Arthropoda" rank="species" species="makranica">
|
||
C.
|
||
<emphasis id="B9502A1D5B1B5F1F079DF8419C0391E0" box="[914,1034,1987,2013]" italics="true" pageId="6" pageNumber="373">makranica</emphasis>
|
||
</taxonomicName>
|
||
were found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |