140 lines
17 KiB
XML
140 lines
17 KiB
XML
<document id="AB175812C9A898E1D6B3F2AC9CA3845E" ID="10.11646/zootaxa.4178.3.3" ID-DOI="10.11646/zootaxa.4178.3.3" ID-GBIF-Dataset="e86d5f4d-f965-4329-91c5-cfcd7c8ed33b" ID-ISSN="1175-5326" ID-Zenodo-Dep="163367" ID-ZooBank="70509AD2-640A-497C-83EA-85B3EBEA35A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477466785637" checkinUser="plazi" docAuthor="Gilligan, Todd, Huemer, Peter & Wiesmair, Benjamin" docDate="2016" docId="7A0E87C6E97DFFA516B415DBF6693E74" docLanguage="en" docName="zootaxa.4178.3.3.pdf" docOrigin="Zootaxa 4178 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ancylis geminana" docType="treatment" docVersion="8" lastPageNumber="363" masterDocId="8637FFBEE972FFB51623102FF7133D29" masterDocTitle="Different continents, same species? Resolving the taxonomy of some Holarctic Ancylis Hübner (Lepidoptera: Tortricidae)" masterLastPageNumber="370" masterPageNumber="347" pageNumber="362" updateTime="1698390823962" updateUser="plazi">
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<mods:title id="2A86C96B16FD06F3522A22E7CFCDB75B">Different continents, same species? Resolving the taxonomy of some Holarctic Ancylis Hübner (Lepidoptera: Tortricidae)</mods:title>
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<mods:name id="A6DFCF38D484966463B68EB60ED3E91A" type="personal">
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<mods:namePart id="AD5AD3FBC75DBC6DC3104A749E09504F">Gilligan, Todd</mods:namePart>
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<mods:namePart id="0F6CE1AE4E4FB2377A0EDC9AC8F75CAF">Huemer, Peter</mods:namePart>
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<mods:namePart id="FC87F8A2A2FAC4DED2D6E37BE02C05BA">Wiesmair, Benjamin</mods:namePart>
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<mods:typeOfResource id="9DF9BD864FF17A74BC3D9DC6D4FD3DCB">text</mods:typeOfResource>
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<mods:title id="E79FA387AE2E3DF375BDB1E3E022956E">Zootaxa</mods:title>
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<mods:date id="23269AF0906B4067E174EF6F419346BA">2016</mods:date>
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<mods:number id="65D14A0A5EF28EC6285910E0C565DB65">4178</mods:number>
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<mods:classification id="476A648840324D6FC670A026595FA044">journal article</mods:classification>
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<mods:identifier id="7226EFBC7B88F0BA863665DC8341424A" type="DOI">10.11646/zootaxa.4178.3.3</mods:identifier>
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<treatment id="7A0E87C6E97DFFA516B415DBF6693E74" ID-DOI="http://doi.org/10.5281/zenodo.6087644" ID-GBIF-Taxon="124558266" ID-Zenodo-Dep="6087644" LSID="urn:lsid:plazi:treatment:7A0E87C6E97DFFA516B415DBF6693E74" httpUri="http://treatment.plazi.org/id/7A0E87C6E97DFFA516B415DBF6693E74" lastPageId="16" lastPageNumber="363" pageId="15" pageNumber="362">
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<subSubSection id="BABD655BE97DFFBA16B415DBF5EF3B96" pageId="15" pageNumber="362" type="nomenclature">
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<paragraph id="F21836D0E97DFFBA16B415DBF6D13B27" blockId="15.[151,450,1524,1550]" box="[151,450,1524,1550]" pageId="15" pageNumber="362">
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<heading id="A95081BCE97DFFBA16B415DBF6D13B27" bold="true" box="[151,450,1524,1550]" fontSize="11" level="1" pageId="15" pageNumber="362" reason="1">
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<emphasis id="C0D3EAC2E97DFFBA16B415DBF6D13B27" bold="true" box="[151,450,1524,1550]" pageId="15" pageNumber="362">
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<taxonomicName id="35A74D53E97DFFBA16B415DBF6623B27" ID-CoL="DNHC" box="[151,369,1524,1550]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="geminana">
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<emphasis id="C0D3EAC2E97DFFBA16B415DBF6623B27" bold="true" box="[151,369,1524,1550]" italics="true" pageId="15" pageNumber="362">Ancylis geminana</emphasis>
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</taxonomicName>
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group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="F21836D0E97DFFBA16B41614F5EF3B96" blockId="15.[151,1437,1595,2015]" pageId="15" pageNumber="362">
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The
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<taxonomicName id="35A74D53E97DFFBA16E81613F6493B7A" box="[203,346,1596,1619]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="geminana">
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<emphasis id="C0D3EAC2E97DFFBA16E81613F6493B7A" box="[203,346,1596,1619]" italics="true" pageId="15" pageNumber="362">A. geminana</emphasis>
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</taxonomicName>
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group contains
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<taxonomicName id="35A74D53E97DFFBA14311613F5B23B7A" box="[530,673,1596,1619]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="geminana">
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<emphasis id="C0D3EAC2E97DFFBA14311613F5B23B7A" box="[530,673,1596,1619]" italics="true" pageId="15" pageNumber="362">A. geminana</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="35A74D53E97DFFBA14931613F4673B7D" box="[688,884,1595,1620]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="christiandiana">
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<emphasis id="C0D3EAC2E97DFFBA14931613F4673B7D" box="[688,884,1595,1620]" italics="true" pageId="15" pageNumber="362">A. christiandiana</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="35A74D53E97DFFBA15A71613F3303B7D" box="[900,1059,1595,1620]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="diminutana">
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<emphasis id="C0D3EAC2E97DFFBA15A71613F3303B7D" box="[900,1059,1595,1620]" italics="true" pageId="15" pageNumber="362">A. diminutana</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="35A74D53E97DFFBA12171613F3F23B7D" box="[1076,1249,1595,1620]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="diminuatana">
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<emphasis id="C0D3EAC2E97DFFBA12171613F3F23B7D" box="[1076,1249,1595,1620]" italics="true" pageId="15" pageNumber="362">A. diminuatana</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="35A74D53E97DFFBA12D21613F2863B7D" box="[1265,1429,1595,1620]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="subarcuana">
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<emphasis id="C0D3EAC2E97DFFBA12D21613F2863B7D" box="[1265,1429,1595,1620]" italics="true" pageId="15" pageNumber="362">A. subarcuana</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="35A74D53E97DFFBA16EA1670F62D3B5E" box="[201,318,1630,1655]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="saliana">
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<emphasis id="C0D3EAC2E97DFFBA16EA1670F62D3B5E" box="[201,318,1630,1655]" italics="true" pageId="15" pageNumber="362">A. saliana</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="35A74D53E97DFFBA176D1671F52A3B5E" box="[334,569,1630,1655]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="diminuatana">
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<emphasis id="C0D3EAC2E97DFFBA176D1671F52A3B5E" box="[334,569,1630,1655]" italics="true" pageId="15" pageNumber="362">Ancylis diminuatana</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="35A74D53E97DFFBA14571670F40A3B5E" box="[628,793,1630,1655]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="subarcuana">
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<emphasis id="C0D3EAC2E97DFFBA14571670F40A3B5E" box="[628,793,1630,1655]" italics="true" pageId="15" pageNumber="362">A. subarcuana</emphasis>
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</taxonomicName>
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are elevated from synonymy with
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<taxonomicName id="35A74D53E97DFFBA12931670F2533B5E" box="[1200,1344,1631,1655]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="geminana">
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<emphasis id="C0D3EAC2E97DFFBA12931670F2533B5E" box="[1200,1344,1631,1655]" italics="true" pageId="15" pageNumber="362">A. geminana</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="35A74D53E97DFFBA13A71670F6283BB5" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="christiandiana">
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<emphasis id="C0D3EAC2E97DFFBA13A71670F6283BB5" italics="true" pageId="15" pageNumber="362">A. christiandiana</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="35A74D53E97DFFBA175216ABF6F73BB5" box="[369,484,1667,1692]" class="Insecta" family="Tortricidae" genus="Ancylis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="362" phylum="Arthropoda" rank="species" species="saliana">
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<emphasis id="C0D3EAC2E97DFFBA175216ABF6F73BB5" box="[369,484,1667,1692]" italics="true" pageId="15" pageNumber="362">A. saliana</emphasis>
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</taxonomicName>
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are described as new. As a result of these taxonomic changes, none of the species in this group is considered to be Holarctic in distribution.
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</paragraph>
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</subSubSection>
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<subSubSection id="BABD655BE97DFFA516E416E4F6693E74" lastPageId="16" lastPageNumber="363" pageId="15" pageNumber="362" type="description">
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<paragraph id="F21836D0E97DFFBA16E416E4F6AD3A66" blockId="15.[151,1437,1595,2015]" pageId="15" pageNumber="362">Species identification in this group is difficult and usually relies on minor differences in wing pattern. Male genitalia are variable and useless in discriminating species; we illustrate each species to display the range of variability in this group. Female genitalia are uniform with species-specific characters for only a couple taxa. Shared characters include:</paragraph>
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<paragraph id="F21836D0E97DFFBA16E41773F2793AF6" blockId="15.[151,1437,1595,2015]" pageId="15" pageNumber="362">
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<emphasis id="C0D3EAC2E97DFFBA16E41773F6203A5A" box="[199,307,1884,1907]" italics="true" pageId="15" pageNumber="362">Forewing</emphasis>
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. Fasciae are undefined; instead the wing is divided longitudinally along the radius and/or cubitus into a darker costal half and a lighter dorsal half, creating a two-toned appearance in some species. The border between the costal and dorsal halves creates a sinuate “longitudinal line,” often bordered in white, which runs from base to apex. The shape and degree of expression of the longitudinal line is often useful in diagnosing species.
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</paragraph>
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<paragraph id="F21836D0E962FFA516E410B8F5733C4C" blockId="16.[151,1437,151,861]" pageId="16" pageNumber="363">
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<emphasis id="C0D3EAC2E962FFA516E410B8F67F3D99" box="[199,364,151,176]" italics="true" pageId="16" pageNumber="363">Male genitalia</emphasis>
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. Uncus bifid to 0.75–0.9 its total length, well developed in all species. Socii large membranous setose pads. Valva with costal margin concave to nearly straight; saccular angle moderately to strongly developed with variable triangular terminal projection; neck width variable, from moderate to wide; cucullus with rounded dorsal lobe and narrow fingerlike ventral lobe, outer margin concave to straight, variably setose with rows of short setae along dorsal lobe and outer margin. Caulis 0.5–0.7 as long as phallus; phallus 0.4–0.5 as long as valva; vesica with> 100 deciduous lanceolate cornuti.
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</paragraph>
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<paragraph id="F21836D0E962FFA516E41140F6953F15" blockId="16.[151,1437,151,861]" pageId="16" pageNumber="363">
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<emphasis id="C0D3EAC2E962FFA516E41140F6963CA1" box="[199,389,367,392]" italics="true" pageId="16" pageNumber="363">Female genitalia</emphasis>
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. Papillae anales moderately setose. Apophyses posteriores the same length or 0.75 as long as apophyses anteriores. Lamella postvaginalis membranous, undefined; lamella antevaginalis sclerotized, projecting ventrally above ostium, often with slight median indentation, forming a weakly sclerotized funnel-shaped antrum. Colliculum present as two small lateral sclerites continuous with anterior sclerotization of antrum. Ductus seminalis arising posterior to junction of ductus and corpus bursae. Corpus bursae large, oval; two blade- or hornshaped signa present.
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</paragraph>
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<paragraph id="F21836D0E962FFA516E41266F6693E74" blockId="16.[151,1437,151,861]" pageId="16" pageNumber="363">
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<emphasis id="C0D3EAC2E962FFA516E41266F63C3F49" box="[199,303,585,608]" italics="true" pageId="16" pageNumber="363">Variation</emphasis>
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. The shape of the male valva varies greatly. The width of the neck, and corresponding degree of development of the dorsal lobe of the cucullus, can vary extensively within species (e.g.,
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<figureCitation id="6A9C2A55E962FFA512B41243F2663FAD" box="[1175,1397,620,645]" captionStart="FIGURES 67 – 74" captionStartId="11.[151,264,1851,1874]" captionTargetBox="[175,1410,223,1816]" captionTargetId="figure@11.[170,1418,223,1817]" captionTargetPageId="11" captionText="FIGURES 67 – 74. Male genitalia. 67, A. geminana ([no data], TMG 618). 68, A. christiandiana (Austria, TOR 464 P. Huemer). 69, A. diminutana (Germany, TMG 616). 70 – 71, A. diminuatana (70, Maryland, TMG 688; 71, Nebraska, TMG 635). 72, A. saliana (Florida, TMG 676). 73 – 74, A. subarcuana (73, Austria, TMG 619; 74, Germany, TMG 703)." httpUri="https://zenodo.org/record/258418/files/figure.png" pageId="16" pageNumber="363">Figs. 70–71, 73–74</figureCitation>
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) or even within the same individual (e.g.,
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<figureCitation id="6A9C2A55E962FFA5147712A0F5BA3F81" box="[596,681,655,680]" captionStart="FIGURES 67 – 74" captionStartId="11.[151,264,1851,1874]" captionTargetBox="[175,1410,223,1816]" captionTargetId="figure@11.[170,1418,223,1817]" captionTargetPageId="11" captionText="FIGURES 67 – 74. Male genitalia. 67, A. geminana ([no data], TMG 618). 68, A. christiandiana (Austria, TOR 464 P. Huemer). 69, A. diminutana (Germany, TMG 616). 70 – 71, A. diminuatana (70, Maryland, TMG 688; 71, Nebraska, TMG 635). 72, A. saliana (Florida, TMG 676). 73 – 74, A. subarcuana (73, Austria, TMG 619; 74, Germany, TMG 703)." httpUri="https://zenodo.org/record/258418/files/figure.png" pageId="16" pageNumber="363">Fig. 71</figureCitation>
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). The shape of the ventral lobe of the cucullus also varies within species and individuals. Note that the appearance of valval shape is influenced by the method of slide preparation; any tilting or twisting of the genitalia during mounting will alter the appearance of the cucullus and the sacculus. The male genitalia figures presented here (
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<figureCitation id="6A9C2A55E962FFA5145F12D3F4173E3C" box="[636,772,764,789]" captionStart="FIGURES 67 – 74" captionStartId="11.[151,264,1851,1874]" captionTargetBox="[175,1410,223,1816]" captionTargetId="figure@11.[170,1418,223,1817]" captionTargetPageId="11" captionText="FIGURES 67 – 74. Male genitalia. 67, A. geminana ([no data], TMG 618). 68, A. christiandiana (Austria, TOR 464 P. Huemer). 69, A. diminutana (Germany, TMG 616). 70 – 71, A. diminuatana (70, Maryland, TMG 688; 71, Nebraska, TMG 635). 72, A. saliana (Florida, TMG 676). 73 – 74, A. subarcuana (73, Austria, TMG 619; 74, Germany, TMG 703)." httpUri="https://zenodo.org/record/258418/files/figure.png" pageId="16" pageNumber="363">Figs. 67–74</figureCitation>
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) illustrate the range of variability found within this group; however, none of the variability is specific to a particular species and any of the figures could represent any of the species treated here.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |