485 lines
52 KiB
XML
485 lines
52 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1176.104980" ID-Pensoft-Pub="1313-2970-1176-13" ID-Pensoft-UUID="C178263C62205EE685739607736AAB36" ID-ZooBank="5C628D2388934955B260F4052F4D8193" ModsDocID="1313-2970-1176-13" checkinTime="1692723655473" checkinUser="pensoft" docAuthor="Miller, Kelly B., Benetti, Cesar J. & Michat, Mariano C." docDate="2023" docId="34C12EF8D2285E0982A3EA2FCBDCF4D0" docLanguage="en" docName="ZooKeys 1176: 13-28" docOrigin="ZooKeys 1176" docPubDate="2023-08-22" docSource="http://dx.doi.org/10.3897/zookeys.1176.104980" docTitle="Miradessus beni Miller, Benetti & Michat 2023, sp. nov." docType="treatment" docUuid="6A3EEA04-9DEA-41DA-A2F4-40E795270841" docUuidSource="ZooBank" docVersion="1" id="C178263C62205EE685739607736AAB36" lastPageNumber="13" masterDocId="C178263C62205EE685739607736AAB36" masterDocTitle="Miradessus gen. nov. from South America described for two species previously in Amarodytes Regimbart, 1900 and two new species (Arthropoda, Insecta, Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Bidessini)" masterLastPageNumber="28" masterPageNumber="13" pageNumber="13" updateTime="1692723655473" updateUser="pensoft">
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<mods:title id="290B3B36AEB916EFCEBA901DC30ADBCE">Miradessus gen. nov. from South America described for two species previously in Amarodytes Regimbart, 1900 and two new species (Arthropoda, Insecta, Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Bidessini)</mods:title>
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<mods:name id="5FC992AA69FC086FB525B1F7428BCFC4" type="personal">
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<mods:roleTerm id="D2175A9C27579A804FE6F8CB7B0514B5">Author</mods:roleTerm>
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<mods:namePart id="0B7DEE99AEA6F8AE595DC6243DFDEF9C">Miller, Kelly B.</mods:namePart>
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<mods:affiliation id="27A5C4F5A0C41173B664478F55776F67">Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
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<mods:nameIdentifier id="2D6A09BB090C9EA61FCFF6C7AA217045" type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:name id="2AAA4AD9200F11B45C0EB94F8B4C004A" type="personal">
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<mods:roleTerm id="51524FC5E096C01E2E6B546C4FF3139C">Author</mods:roleTerm>
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<mods:namePart id="44C6DF613B5E9854D5F0FF3CAECD404F">Benetti, Cesar J.</mods:namePart>
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<mods:nameIdentifier id="AA048752AA444F19A6A5B30BBD80C974" type="ORCID">https://orcid.org/0000-0001-6795-2194</mods:nameIdentifier>
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<mods:affiliation id="9A6844BFC6B4EC33D4F7F1B724507A85">Departamento de Biodiversidad y Gestion Ambiental, Facultad de Ciencias Biologicas y Ambientales, Universidad de Leon, Campus de Vegazana, 24071, Leon, Spain & Coordenacao de Biodiversidade, Programa de Pos-Graduacao em Entomologia (PPGEnt), Instituto Nacional de Pesquisas da Amazonia (INPA), Av. Andre Araujo 2936, CEP 69067 - 375, Manaus, AM, Brazil</mods:affiliation>
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<mods:namePart id="D7DD04118D3F2A262233397EA998A5B4">Michat, Mariano C.</mods:namePart>
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<mods:nameIdentifier id="044B4136779E7CB0127ECADAFC31E002" type="ORCID">https://orcid.org/0000-0002-1962-7976</mods:nameIdentifier>
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<mods:affiliation id="CD5836373A4CF15C735A61C48F86FE44">Faculty of Exact and Natural Sciences, Department of Biodiversity and Experimental Biology, Laboratory of Entomology, Institute of Biodiversity and Experimental and Applied Biology (IBBEA), CONICET-University of Buenos Aires, Buenos Aires, Argentina</mods:affiliation>
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<mods:typeOfResource id="80F738763BBC29A683917AD7BB2C883C">text</mods:typeOfResource>
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<mods:title id="FD3E3F8AE14DDA0426C10E59807F009F">ZooKeys</mods:title>
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<mods:part id="384140E3EB09F0B355593004015D5590">
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<mods:date id="55FFFB2C0E0762E0875B3B0791455F4A">2023</mods:date>
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<mods:number id="E3F1CF417400A0ED9BAE991C6FA9D383">2023-08-22</mods:number>
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<mods:number id="88E59A10AEAA8A83C5AD6C4542AD82B9">1176</mods:number>
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<mods:start id="BA62C832DF9D8241A3890267AB567BA6">13</mods:start>
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<mods:url id="3F38A5A47970129F5A784E7805253232">http://dx.doi.org/10.3897/zookeys.1176.104980</mods:url>
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<mods:classification id="9BFEDAC04010286FCCBCC0CF902035EF">journal article</mods:classification>
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<mods:identifier id="34792A05ABD4B50E029ACCE9EF1267A0" type="DOI">http://dx.doi.org/10.3897/zookeys.1176.104980</mods:identifier>
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<treatment id="34C12EF8D2285E0982A3EA2FCBDCF4D0" LSID="urn:lsid:zoobank.org:act:6A3EEA04-9DEA-41DA-A2F4-40E795270841" httpUri="http://treatment.plazi.org/id/34C12EF8D2285E0982A3EA2FCBDCF4D0" lastPageNumber="13" pageId="0" pageNumber="13">
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<subSubSection id="3D1B800D6B1B79302FE3F00D9BA0FC6C" pageId="0" pageNumber="13" type="nomenclature">
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<paragraph id="18F891CB54431D3BDAAD7289B014EBA3" pageId="0" pageNumber="13">
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<taxonomicName id="7B4C53697286CF1A195FB3F9C1496CD6" LSID="https://zoobank.org/6A3EEA04-9DEA-41DA-A2F4-40E795270841" authority="Miller, Benetti & Michat" authorityName="Miller, Benetti & Michat" authorityYear="2023" family="Dytiscidae" genus="Miradessus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Miradessus beni" order="Coleoptera" pageId="0" pageNumber="13" rank="species" species="beni" status="sp. nov.">Miradessus beni Miller, Benetti & Michat</taxonomicName>
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<taxonomicNameLabel id="7CAA5348693BB5991A12970C79868BBE" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="136F2F875BA5C0864905125C91AED4ED" pageId="0" pageNumber="13" type="description">
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<paragraph id="745E89E11891E33BD80FF4B65EEB53B1" pageId="0" pageNumber="13">
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<figureCitation id="277F7A48E8A6688A4186BCF59B07F046" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">Figs 1</figureCitation>
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<figureCitation id="8A61FB9F68398C329BA7A106D07C4997" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">, 5</figureCitation>
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<figureCitation id="28BDCEF032C89AC5B019CEF2175B120F" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">, 9</figureCitation>
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<figureCitation id="C29316E4AE6284F0265796356ACD2605" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">, 10</figureCitation>
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<figureCitation id="5BBC7FF79DBD1424F0DA22AB4BF07933" captionStart="Figures 18–29" captionStartId="F3" captionText="Figures 18 - 29. Miradessus species, primary type specimens and labels 18 - 20 M. beni, holotype specimen 18 dorsal habitus 19 ventral habitus 20 specimen labels 21 - 23 M. plaumanni, lectotype specimen 21 dorsal habitus 22 ventral habitus 23 specimen labels 24 - 26 M. pulchellus, holotype specimen 24 dorsal habitus 25 ventral habitus 26 specimen labels 27 - 29 M. rikae, holotype specimen 27 dorsal habitus 28 ventral habitus 29 specimen labels. 21 - 23 courtesy of M. Schwarz (OLML) 24 - 26 courtesy of M. A. Balke (ZSMG)." figureDoi="10.3897/zookeys.1176.104980.figures18-29" httpUri="https://binary.pensoft.net/fig/895236" pageId="0" pageNumber="13">, 18-20</figureCitation>
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<figureCitation id="D2AF346BE3C3F821C3653499847FA58C" captionStart="Figure 30" captionStartId="F4" captionText="Figure 30. Miradessus species, distributions." figureDoi="10.3897/zookeys.1176.104980.figure30" httpUri="https://binary.pensoft.net/fig/895237" pageId="0" pageNumber="13">, 30</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="09CFA5C0B6C49D46B1D53F4B963DEFED" pageId="0" pageNumber="13" type="type locality">
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<paragraph id="95C63911CE9CC99C50113732859F752E" pageId="0" pageNumber="13">Type locality.</paragraph>
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<paragraph id="F1AA74E3716211C85E09BDC14A533961" pageId="0" pageNumber="13">
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Bolivia, Departmento de La Paz, Provincia de Abel Iturralde, San Miguel del Bala,
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<geoCoordinate id="D44BD3CB7A01A20DA4EEC31503B3B402" degrees="14" direction="south" minutes="30.602" orientation="latitude" precision="1" value="-14.510034">14°30.602'S</geoCoordinate>
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,
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<geoCoordinate id="4748776B70455650B5E29D40CEFBB4D0" degrees="67" direction="west" minutes="29.555" orientation="longitude" precision="1" value="-67.492584">67°29.555'W</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="3D7664879FC6BC7E6C4E061BC3013D8A" pageId="0" pageNumber="13" type="diagnosis">
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<paragraph id="F20FEB4E8A5913073B08994A56C13145" pageId="0" pageNumber="13">Diagnosis.</paragraph>
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<paragraph id="587EE5968D706290641156F5F4715F6B" pageId="0" pageNumber="13">
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This species and
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<taxonomicName id="D58887C2D51FA29AD901C3FA2A51F269" lsidName="M. pulchellus" pageId="0" pageNumber="13" rank="species" species="pulchellus">
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<emphasis id="8DAB0F869A54A17E01166DECFEFE5044" italics="true" pageId="0" pageNumber="13">M. pulchellus</emphasis>
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</taxonomicName>
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are extremely similar externally, with similar overall shape, color pattern, and other features (Figs
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<figureCitation id="6B2C88970CB5A9CBE471E861A096ED31" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">1</figureCitation>
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,
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<figureCitation id="A89F0EF7142C987EB47ECB0FB51FEDEC" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">3</figureCitation>
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). The external differences between them are subtle, including the shape of the prosternal process, which is more prominently laterally carinate and deeply sulcate in
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<taxonomicName id="7F9A0006444C5E00ACFC6F675B7E6858" lsidName="M. pulchellus" pageId="0" pageNumber="13" rank="species" species="pulchellus">
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<emphasis id="3C77EA54704A4E7F79B2112A15605BBB" italics="true" pageId="0" pageNumber="13">M. pulchellus</emphasis>
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</taxonomicName>
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than in
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<taxonomicName id="D043A4EC37672B29EE993243A363836D" lsidName="M. beni" pageId="0" pageNumber="13" rank="species" species="beni">
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<emphasis id="BC5D79CAE0200473BC6AF1113E247432" italics="true" pageId="0" pageNumber="13">M. beni</emphasis>
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</taxonomicName>
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. Also, the ventral surface is darker in most specimens of
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<taxonomicName id="779556A5BA240C49F7C88C6E89D359D7" lsidName="M. pulchellus" pageId="0" pageNumber="13" rank="species" species="pulchellus">
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<emphasis id="416F66DCECDEEC4A7B2B3DC4C82B745E" italics="true" pageId="0" pageNumber="13">M. pulchellus</emphasis>
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</taxonomicName>
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than in
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<taxonomicName id="C5C553EDBEE2964280122EA4D22FA469" lsidName="M. beni" pageId="0" pageNumber="13" rank="species" species="beni">
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<emphasis id="01F97794708D356CC67263C9174B0B35" italics="true" pageId="0" pageNumber="13">M. beni</emphasis>
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</taxonomicName>
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. The main differences between these species are in the male genitalia, which are distinctive and characteristic. Both have the median lobe trilobed (with two side portions and a middle portion), but in
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<taxonomicName id="45764D7FEA5A9129894F44744C804323" lsidName="M. pulchellus" pageId="0" pageNumber="13" rank="species" species="pulchellus">
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<emphasis id="359C918C9785222116EACF9F97B04BDC" italics="true" pageId="0" pageNumber="13">M. pulchellus</emphasis>
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</taxonomicName>
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the middle portion is nearly as long as the side portions and the apex has a distinct, fine pencil of setae on each side that are divergent (Fig.
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<figureCitation id="610F478887031FBB534E7AFFFB8CDE73" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">13</figureCitation>
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). In
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<taxonomicName id="25A149D3A181EBAA5E1104B47AC55D97" lsidName="M. beni" pageId="0" pageNumber="13" rank="species" species="beni">
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<emphasis id="FDB84CF0BA752BAE7FC7AEBC576DFD38" italics="true" pageId="0" pageNumber="13">M. beni</emphasis>
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</taxonomicName>
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the middle portion is considerably shorter than the side portions and has a series of setae along its apex (Fig.
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<figureCitation id="DD0B10867E1E2B8A1354032043290E19" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">9</figureCitation>
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). In lateral aspect, the lateral lobe of
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<taxonomicName id="FC523F4DA3CA835B4EEC6DAFF71D5C77" lsidName="M. pulchellus" pageId="0" pageNumber="13" rank="species" species="pulchellus">
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<emphasis id="F4EEF04054F8C853533F449C0749EA73" italics="true" pageId="0" pageNumber="13">M. pulchellus</emphasis>
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</taxonomicName>
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has the apical segment nearly as long as the basal segment and it is curved ventrad (Fig.
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<figureCitation id="8FF4EAF2C7ADA3BD68648958B405780C" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">14</figureCitation>
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). In
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<taxonomicName id="989EDC4B183670253DC8A391B7DEBA3C" lsidName="M. beni" pageId="0" pageNumber="13" rank="species" species="beni">
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<emphasis id="B06D2BCB62F6C0361C2C8D2964C7E02C" italics="true" pageId="0" pageNumber="13">M. beni</emphasis>
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</taxonomicName>
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the lateral lobe has the apical segment only about 1/3 the length of the basal segment and it is curved dorsad (Fig.
|
||
<figureCitation id="2C1A74B02515AB96518B017503A4EAB6" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2BAF394F8DCAF9100C507D8FBF684CE2" pageId="0" pageNumber="13" type="description">
|
||
<paragraph id="893B4E4602DDC26A5C6B6E254E53695E" pageId="0" pageNumber="13">Description.</paragraph>
|
||
<paragraph id="C2B1C3F321B0E539C156E134807BF493" pageId="0" pageNumber="13">
|
||
<emphasis id="7D84EC3B109C14BDA341F005CFF6EBBD" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="A3F1CFAE474AF6D15F412ADE751F2E4F" italics="true" pageId="0" pageNumber="13">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
TL = 2.7-2.8 mm, GW = 1.3-1.4 mm, PW = 1.0-1.1 mm, HW = 0.7-0.8 mm, EW = 0.3-0.4 mm, TL/GW = 2.0-2.1, HW/EW = 1.8-1.9, FL/RL = 2.2-2.3.
|
||
</paragraph>
|
||
<paragraph id="03EED5F797AFD3D3B1F668BB7D665D0C" pageId="0" pageNumber="13">
|
||
<emphasis id="9BDD2A60988E7ECD2AB0F78900A550DC" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="7F2FE68E1A919EB1B3917E975D2FF2B4" italics="true" pageId="0" pageNumber="13">Habitus</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body shape elongate-oval, lateral outline somewhat discontinuous between pronotum and elytron, posterior apex narrowly rounded (Fig.
|
||
<figureCitation id="838676C9A13C2C6AA543A24376F85D0C" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B513FC74F167CD0B4F1B9D0B52A3F6B9" pageId="0" pageNumber="13">
|
||
<emphasis id="E72EAC84A6F2AA7656DB8F8E094DA62C" bold="true" italics="true" pageId="0" pageNumber="13">Coloration</emphasis>
|
||
(Fig.
|
||
<figureCitation id="A7F7DE63DB055D0BA038505374CDAE22" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">1</figureCitation>
|
||
). Head yellow-orange. Pronotum yellow-orange with a narrow, rounded lobe of black broadly along each side of posterior margin. Elytron maculate, yellow-orange and black, margins of maculae strongly demarcated; with large, transverse yellow-orange regions anteriorly, medially, and apically, not extending medially to suture, with narrow band of black along entire length of suture, margins of maculae irregular and variously lobed (Fig.
|
||
<figureCitation id="B6F7DD1651304EA6C0EDEA6B1D66951C" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">1</figureCitation>
|
||
). Ventral surfaces mostly orange; legs, epipleuron, and lateral areas of prothorax and head lighter orange-yellow, some sutural margins darker, infuscate, mesothoracic ventrites and prosternal process strongly infuscate to nearly black.
|
||
</paragraph>
|
||
<paragraph id="A9B88419EF1844BDB800A966FC4F8A46" pageId="0" pageNumber="13">
|
||
<emphasis id="CF1F4EC2D79EE064C8179B160FAD5774" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="DDF3C438457C4C403325CCA8760D65E4" italics="true" pageId="0" pageNumber="13">Sculpture and structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Head shiny and smooth, nearly impunctate, with small micropunctures sparsely distributed; eyes moderately large (HW/EW = 1.8-1.9); antennae slender, unmodified. Pronotum with lateral margins moderately curved anteriorly, shallowly curved posteriorly; with narrow bead along entire margin; surface shiny, moderately and evenly punctate; lateral pronotal plica weakly impressed, sublinear, extending about 1/3 distance across pronotum. Elytron with lateral margin evenly and broadly rounded; surface shiny, moderately and evenly punctate. Prosternum medially moderately broad, medially not protruberant, mediolaterally somewhat granular; prosternal process moderately broad medially with slight tubercle, apical blade large, covered with fine setae, laterally with prominent rounded ridges along entire length, medially longitudinally sulcate, lateral margins slightly convergent to narrowly rounded apex (Fig.
|
||
<figureCitation id="746796DD6BF48F2204036D4F6F39526F" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">5</figureCitation>
|
||
). Metaventer and metaventral wings smooth and shiny, covered with shallow punctures (Fig.
|
||
<figureCitation id="3CF37B843656857BB9B750B3765CF77E" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">5</figureCitation>
|
||
). Metacoxa with medial portion moderately broad, metacoxal lines distinct, evenly divergent anteriorly to posterior margin of metaventrite, lateral portion large, medially and anteriorly covered with shallow punctures; metatrochanter about 1/3 length of metafemur (Fig.
|
||
<figureCitation id="FDBB819BE27E013CEBBFD1CAE0BE7CF5" captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Miradessus species 1 - 4 dorsal habitus 1 M. beni 2 M. plaumanni 3 M. pulchellus 4 M. rikae 5 - 8 prosternal process left surfaces of metaventrite metacoxa metatrochanter and metafemur 5 Miradessus beni 6 M. plaumanni 7 M. pulchellus 8 M. rikae. Scale bars: 1.0 mm (1 - 4); 0.5 mm (5 - 8)." figureDoi="10.3897/zookeys.1176.104980.figures1-8" httpUri="https://binary.pensoft.net/fig/895234" pageId="0" pageNumber="13">5</figureCitation>
|
||
). Abdominal ventrites impunctate except II and III laterally with shallow, indistinct punctures; VI strongly concave in lateral aspect, apex pointed.
|
||
</paragraph>
|
||
<paragraph id="7E0DFE98227F1B19C90FB41FE072FDB8" pageId="0" pageNumber="13">
|
||
<emphasis id="5BAE504CEA2C73542594A6C386BFF97C" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="A002D2E74D34FB82A46C7959662318FF" italics="true" pageId="0" pageNumber="13">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Median lobe in ventral aspect broad, trilobed, with dorso-medial, elongate, slender, apically pointed ramus and another ventral, elongate, apically bilobedramus, with each ramus elongate and slender, medial dorsal ramus much shorter than ventral ramus (Fig.
|
||
<figureCitation id="234C68E1DAFEE5E56F2CE900C032C1E9" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">9</figureCitation>
|
||
); lateral lobe in lateral aspect robust, basal segment very broad, robust, apical segment short, robust, curved dorsad, apex obliquely truncate (Fig.
|
||
<figureCitation id="9287DC03B5808254D4E22810BFAF96E1" captionStart="Figures 9–17" captionStartId="F2" captionText="Figures 9 - 17. Miradessus species 9, 10 M. beni, male aedeagus 9 median lobe and left lateral lobe, dorsal aspect 10 right lateral lobe, right lateral aspect 11, 12 M. plaumanni, male aedeagus 11 median lobe and lateral lobes, dorsal aspect 12 right lateral lobe, right lateral aspect 13, 14 M. pulchellus, male aedeagus 13 median lobe and left lateral lobe, dorsal aspect 14 right lateral lobe, right lateral aspect 15, 16 M. rikae, male aedeagus 15 median lobe and lateral lobes, dorsal aspect 16 median lobe and right lateral lobe, right lateral aspect 17 M. pulchellus, female genitalia, ventral aspect. Scale bar: 0.25 mm." figureDoi="10.3897/zookeys.1176.104980.figures9-17" httpUri="https://binary.pensoft.net/fig/895235" pageId="0" pageNumber="13">10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3A9A304338B623D31D9EE8E51BAC47A2" pageId="0" pageNumber="13">
|
||
<emphasis id="91F2BA436875A546D9C0C133A4880418" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="D7B16C8482CBC6ECF65A309EA80F7CD1" italics="true" pageId="0" pageNumber="13">Sexual dimorphism</emphasis>
|
||
.
|
||
</emphasis>
|
||
Males have the pro- and mesotarsomeres I-III slightly but distinctly broader than in females. Abdominal ventrite VI distinctly concave in lateral aspect in females, but medially more expanded and apically somewhat depressed in males.
|
||
</paragraph>
|
||
<paragraph id="060B57BD051B6987EA4D7288CE259A4A" pageId="0" pageNumber="13">
|
||
<emphasis id="C380CBDA2AA49F5F6202885E0BB7FBE3" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="F4FEE08B6B31A68AE9EC96CF8DD5DE82" italics="true" pageId="0" pageNumber="13">Variation</emphasis>
|
||
.
|
||
</emphasis>
|
||
There is some minor variation in shape and extent of coloration of the dorsal and ventral surfaces but otherwise specimens are similar.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11316D78C91223213A1CC5B00B364234" pageId="0" pageNumber="13" type="etymology">
|
||
<paragraph id="5B74A370CDA65B4BEFA84F83BE940551" pageId="0" pageNumber="13">Etymology.</paragraph>
|
||
<paragraph id="BB368936EC10CA4EB2AE52DD688932EF" pageId="0" pageNumber="13">
|
||
This species is named
|
||
<emphasis id="FACE97C4A1E11B07022045261BEBF1AF" italics="true" pageId="0" pageNumber="13">beni</emphasis>
|
||
after the name
|
||
<normalizedToken id="536A587888F71432A6408E47E2D350BD" originalValue="Río">Rio</normalizedToken>
|
||
Beni, the river at the type locality of the species. The name is a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D6F7930E15CD7D7A20BF9B6643D55729" pageId="0" pageNumber="13" type="distribution">
|
||
<paragraph id="6E8985C024C35665E8D85B2646CBE38C" pageId="0" pageNumber="13">Distribution.</paragraph>
|
||
<paragraph id="5020776BC5ED7AF59EB8CA9166AF9029" pageId="0" pageNumber="13">
|
||
<taxonomicName id="5C9CCD4FA61B5804CFAB890D22D730E5" authorityName="Miller, Benetti & Michat" authorityYear="2023" family="Dytiscidae" genus="Miradessus" kingdom="Animalia" lsidName="Miradessus beni" order="Coleoptera" pageId="0" pageNumber="13" rank="species" species="beni">
|
||
<emphasis id="99BD0E186E75861733EB645801752B10" italics="true" pageId="0" pageNumber="13">Miradessus beni</emphasis>
|
||
</taxonomicName>
|
||
is known from the type locality in lowland Departmento de La Paz, Bolivia and from two sites in lowland Peru (Fig.
|
||
<figureCitation id="CB181BF865645F6115942F65461B8961" captionStart="Figure 30" captionStartId="F4" captionText="Figure 30. Miradessus species, distributions." figureDoi="10.3897/zookeys.1176.104980.figure30" httpUri="https://binary.pensoft.net/fig/895237" pageId="0" pageNumber="13">30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06A08C55798EEBC63608DD092058813F" pageId="0" pageNumber="13" type="habitat">
|
||
<paragraph id="BD1E1C5340291A844A3A682A8C5484E4" pageId="0" pageNumber="13">Habitat.</paragraph>
|
||
<paragraph id="43CBB152C4EBB62EBEDF18B288567D84" pageId="0" pageNumber="13">The type locality is a heavily forested area of the Andean foothills next to a large river. However, nothing is known of the specific collection habitat of this species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="125F27315BBE3C7E3B843EEAE3B1F6F2" pageId="0" pageNumber="13" type="materials_examined">
|
||
<paragraph id="D7F42CA6507C1DB2905AF4CEAD16503E" pageId="0" pageNumber="13">Material examined.</paragraph>
|
||
<paragraph id="B1B040365C579669CDD8E8AEB2DB6475" pageId="0" pageNumber="13">
|
||
<emphasis id="5C0A64685C8FED1DE931574677F09D55" bold="true" pageId="0" pageNumber="13">
|
||
<emphasis id="CD8D9B06E64017AA0AA728D4E46C10F2" italics="true" pageId="0" pageNumber="13">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="511BBFFA15984B08244DBA9320D3CC33" collectingDate="2007-09-24" collectingDateMax="2007-09-30" collectingDateMin="2007-09-24" country="BOLIVIA" latitude="-14.510034" location="Figs" longLatPrecision="1" longitude="-67.492584" specimenCount="1" stateProvince="La Paz" typeStatus="Holotype">
|
||
<emphasis id="C875988D3B57308B652959DB24E3A2A1" italics="true" pageId="0" pageNumber="13">
|
||
<typeStatus id="C5C0F9111FF3885B91A19EC48C94611F">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<location id="DA7470D0AB8204618EA337BC68BE794B" LSID="urn:lsid:plazi:treatment:34C12EF8D2285E0982A3EA2FCBDCF4D0:DA7470D0AB8204618EA337BC68BE794B" country="BOLIVIA" latitude="-14.510034" longLatPrecision="1" longitude="-67.492584" name="Figs" stateProvince="La Paz">Figs</location>
|
||
<figureCitation id="969C6A29D507AEB940E175969FFA1EB5" captionStart="Figures 18–29" captionStartId="F3" captionText="Figures 18 - 29. Miradessus species, primary type specimens and labels 18 - 20 M. beni, holotype specimen 18 dorsal habitus 19 ventral habitus 20 specimen labels 21 - 23 M. plaumanni, lectotype specimen 21 dorsal habitus 22 ventral habitus 23 specimen labels 24 - 26 M. pulchellus, holotype specimen 24 dorsal habitus 25 ventral habitus 26 specimen labels 27 - 29 M. rikae, holotype specimen 27 dorsal habitus 28 ventral habitus 29 specimen labels. 21 - 23 courtesy of M. Schwarz (OLML) 24 - 26 courtesy of M. A. Balke (ZSMG)." figureDoi="10.3897/zookeys.1176.104980.figures18-29" httpUri="https://binary.pensoft.net/fig/895236" pageId="0" pageNumber="13">18-20</figureCitation>
|
||
) in MIZA, male labeled, "
|
||
<collectingCountry id="44064E8B7F1E6B5F32333B8442F738C3" name="Bolivia">BOLIVIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="CD15BA09DD078E998D59E85E5D41D483" country="Bolivia" name="La Paz">La Paz</collectingRegion>
|
||
Dept. Ituralde [sic] Prov., San Miguel
|
||
<geoCoordinate id="8BE7A7E00B5C1DF9DC724945B70125B6" degrees="14" direction="south" minutes="30.602" orientation="latitude" precision="1" value="-14.510034">14°30.602'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D05F0EBC60309D65DCE11DA06AFF5798" degrees="67" direction="west" minutes="29.555" orientation="longitude" precision="1" value="-67.492584">67°29.555'W</geoCoordinate>
|
||
,
|
||
<collectingDate id="25D821DF90E5DB43EF012796C6900F92" value="2007-09-24" valueMax="2007-09-30" valueMin="2007-09-24">24-30 Sept. 2007</collectingDate>
|
||
KB Miller KBMC24090701/
|
||
<typeStatus id="39E903EDFDA5B09A336F302E8C650649">HOLOTYPE</typeStatus>
|
||
<taxonomicName id="84749E763AF7B3CE7AA0E3B5E2172D6C" authorityName="Miller, Benetti & Michat" authorityYear="2023" family="Dytiscidae" genus="Miradessus" kingdom="Animalia" lsidName="Miradessus beni" order="Coleoptera" pageId="0" pageNumber="13" rank="species" species="beni">
|
||
<emphasis id="CE52754BC963336763C8852182125BB8" italics="true" pageId="0" pageNumber="13">Miradessus beni</emphasis>
|
||
</taxonomicName>
|
||
Miller, Benetti & Michat, 2023 [red label with black line border]."
|
||
</materialsCitation>
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="15357C4571F22FA045315EDDF204966E" doi="10.3897/zookeys.1176.104980.figures18-29" httpUri="https://binary.pensoft.net/fig/895236" pageId="0" pageNumber="13" start="Figures 18–29" startId="F3">
|
||
<paragraph id="BC5C6881DA711B2CA7FBD797C62D640D" pageId="0" pageNumber="13">
|
||
<emphasis id="47587C360260A11DF85B388608872F94" bold="true" pageId="0" pageNumber="13">Figures 18-29.</emphasis>
|
||
<taxonomicName id="C09212ECD193410C95F7DE7E2B3D4D78" authorityName="Miller, Benetti & Michat" authorityYear="2023" family="Dytiscidae" genus="Miradessus" kingdom="Animalia" lsidName="Miradessus" order="Coleoptera" pageId="0" pageNumber="13" rank="genus">
|
||
<emphasis id="6D451E45FAE463081503317E87A248AE" italics="true" pageId="0" pageNumber="13">Miradessus</emphasis>
|
||
</taxonomicName>
|
||
species, primary type specimens and labels
|
||
<emphasis id="200B19E08623440445D65E0BAF814212" bold="true" pageId="0" pageNumber="13">
|
||
18-
|
||
<specimenCount id="78333040F1A2D5434C3027CB302816A0" type="generic">
|
||
20
|
||
<taxonomicName id="F95ECAB18CE9B8648081C65BBEEA4D85" lsidName="M. beni" pageId="0" pageNumber="13" rank="species" species="beni">
|
||
<emphasis id="5FA7AC1CC1D1596FF4F52CD53260640E" italics="true" pageId="0" pageNumber="13">M. beni</emphasis>
|
||
</taxonomicName>
|
||
</specimenCount>
|
||
</emphasis>
|
||
,
|
||
<typeStatus id="174B7777B406F8D6FEAE9E9E7EF665AC">holotype</typeStatus>
|
||
specimen
|
||
<emphasis id="DC68A4862BB883B2ECE81C0516903106" bold="true" pageId="0" pageNumber="13">18</emphasis>
|
||
dorsal habitus
|
||
<emphasis id="37E97EEC44958D7E5DC5C2A745349412" bold="true" pageId="0" pageNumber="13">19</emphasis>
|
||
ventral habitus
|
||
<specimenCount id="BA7CE844AB9405D051DCD1194F42DB8D" type="generic">
|
||
<emphasis id="E18F52869EA21FB9F5420EC17D2B5435" bold="true" pageId="0" pageNumber="13">20</emphasis>
|
||
specimen
|
||
</specimenCount>
|
||
labels
|
||
<emphasis id="34AFBEF6FF329DAB6017B8641B659740" bold="true" pageId="0" pageNumber="13">
|
||
21-
|
||
<specimenCount id="58BB90B73B7F2793F6C85203DFBEB56F" type="generic">
|
||
23
|
||
<taxonomicName id="865F2DCE2FD33A422FBE8B8FDCAA2095" lsidName="M. plaumanni" pageId="0" pageNumber="13" rank="species" species="plaumanni">
|
||
<emphasis id="000EE1242614F9B9F86DC72B285BE720" italics="true" pageId="0" pageNumber="13">M. plaumanni</emphasis>
|
||
</taxonomicName>
|
||
</specimenCount>
|
||
</emphasis>
|
||
,
|
||
<typeStatus id="E7E9006C4DA12F6C5B6A0411CB074ED8">lectotype</typeStatus>
|
||
specimen
|
||
<emphasis id="85BC760A48729C94D49250AD2B41EA04" bold="true" pageId="0" pageNumber="13">21</emphasis>
|
||
dorsal habitus
|
||
<emphasis id="F7306FB41B07B7DC459640A8A395046A" bold="true" pageId="0" pageNumber="13">22</emphasis>
|
||
ventral habitus
|
||
<specimenCount id="62CE47DC409F95A81FAE6250B2D1C121" type="generic">
|
||
<emphasis id="21107998EF786C721EB326EFBE7E9A1A" bold="true" pageId="0" pageNumber="13">23</emphasis>
|
||
specimen
|
||
</specimenCount>
|
||
labels
|
||
<emphasis id="1FCA2DD5862328B943E940D11BED0A5C" bold="true" pageId="0" pageNumber="13">
|
||
24-
|
||
<specimenCount id="91A2675B49F5F7AC70EF6DB8A0EE8AB7" type="generic">
|
||
26
|
||
<taxonomicName id="3D5497416ED3AD6AD60930ABDE31921D" lsidName="M. pulchellus" pageId="0" pageNumber="13" rank="species" species="pulchellus">
|
||
<emphasis id="E1369090933A01DA7806E7AB4C0EC17A" italics="true" pageId="0" pageNumber="13">M. pulchellus</emphasis>
|
||
</taxonomicName>
|
||
</specimenCount>
|
||
</emphasis>
|
||
,
|
||
<typeStatus id="1B629A6B3EC2B7FE8156DBAFCD9F4B56">holotype</typeStatus>
|
||
specimen
|
||
<emphasis id="81DCCB33FF875C1E443D3F7E9BCBDE7A" bold="true" pageId="0" pageNumber="13">24</emphasis>
|
||
dorsal habitus
|
||
<emphasis id="8D8DE67C28233B9E72C406301DDAF9AE" bold="true" pageId="0" pageNumber="13">25</emphasis>
|
||
ventral habitus
|
||
<specimenCount id="8EB8ED39B7CB9527DED3AA716F22011C" type="generic">
|
||
<emphasis id="B009B3466433D1ED2C6998D35D9318EB" bold="true" pageId="0" pageNumber="13">26</emphasis>
|
||
specimen
|
||
</specimenCount>
|
||
labels
|
||
<emphasis id="B4E152EFA02EF909194A2B4D2087FC98" bold="true" pageId="0" pageNumber="13">
|
||
27-
|
||
<specimenCount id="0399B434B145AE9991AF4E1AEB870A70" type="generic">
|
||
29
|
||
<taxonomicName id="48A558E44144E572D4CAD969E940D1E5" lsidName="M. rikae" pageId="0" pageNumber="13" rank="species" species="rikae">
|
||
<emphasis id="BEB3766D5FF7E0516F8C4C32EBB1AD35" italics="true" pageId="0" pageNumber="13">M. rikae</emphasis>
|
||
</taxonomicName>
|
||
</specimenCount>
|
||
</emphasis>
|
||
,
|
||
<typeStatus id="9DFD687054800308FDD8673632A5D47A">holotype</typeStatus>
|
||
specimen
|
||
<emphasis id="EB65568E127D282F2852C3FC8A1A741B" bold="true" pageId="0" pageNumber="13">27</emphasis>
|
||
dorsal habitus
|
||
<emphasis id="EEB682900BF615CA81D7A6D665990E93" bold="true" pageId="0" pageNumber="13">28</emphasis>
|
||
ventral habitus
|
||
<specimenCount id="AFB3E4EEE541962A82834613DB55E7AF" type="generic">
|
||
<emphasis id="1DA574E7FF3596F25282C131F304C743" bold="true" pageId="0" pageNumber="13">29</emphasis>
|
||
specimen
|
||
</specimenCount>
|
||
labels.
|
||
<emphasis id="6C175F9385F22F5AB07BD5893248FEFC" bold="true" pageId="0" pageNumber="13">21-23</emphasis>
|
||
courtesy of M. Schwarz (OLML)
|
||
<emphasis id="16415695033211A8F115E5BA60737FEB" bold="true" pageId="0" pageNumber="13">24-26</emphasis>
|
||
courtesy of M.A. Balke (ZSMG).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DB660444FC7BC224CABEAB0980D4E3DF" pageId="0" pageNumber="13">
|
||
<emphasis id="21B408ACFD8C66FEB89A1ACF7443D9A4" bold="true" italics="true" pageId="0" pageNumber="13">
|
||
<typeStatus id="E9D9E9FDB6CD00F26D90E316C88982BF">Paratypes</typeStatus>
|
||
</emphasis>
|
||
, 41 total;
|
||
<materialsCitation id="A5975B6664428262EB2BD680020D5ED1" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="PERU" location="Cuzco Prov." specimenCount="14" stateProvince="Cuzco Prov." typeStatus="holotype">
|
||
<specimenCount id="51601A5BA620D5116710BD7471381D5B" type="generic" typeStatus="paratypes">6 paratypes</specimenCount>
|
||
(MIZA, MSBA) labeled same as
|
||
<typeStatus id="5F3979C9A5292A07D55FFEEF5D170BD2">holotype</typeStatus>
|
||
;"
|
||
<specimenCount id="4520AD7790446633BE62081466E3C64A" type="generic" typeStatus="paratypes">8 paratypes</specimenCount>
|
||
(USNM) labeled, "
|
||
<collectingCountry id="EC859BB6BE49923EC1477BCF118DF476" name="Peru">PERU</collectingCountry>
|
||
:Dpt.
|
||
<collectingRegion id="EB3717967B75AFC8B2F691097C9910A5" country="Peru" name="Cusco">Cuzco Prov.</collectingRegion>
|
||
Quispicanchis Quincemil 6-II-X-1976 Robert Gordon"
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F1BB3A73B4E6FD3E701A8EB4A28EDDB5" collectingDate="2003-04-06" collectingDateMax="2003-04-17" collectingDateMin="2003-04-06" collectorName="E. - G. Burmeister" country="Peru" county="Prov." latitude="-9.616667" location="Biol. Stat. Panguana. Oestl. Ort." longLatPrecision="1299" longitude="-74.933334" municipality="Rio Yuyapichis" specimenCount="12" stateProvince="Huanuco" typeStatus="paratype">
|
||
<specimenCount id="5C3B7D3A5366F18A11543F6B810B0CD4" type="generic" typeStatus="paratypes">12 paratypes</specimenCount>
|
||
(ZSMG) labeled, "
|
||
<taxonomicName id="EF40B00E80299B4CB5A50467F22E1CB8" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Bidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bidessus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
|
||
cruciatus? [handwritten] /
|
||
<taxonomicName id="2966460EF4C3B847FC94715B9F11EE2D" authorityName="Miller & Benetti & Michat" authorityYear="2023" baseAuthorityName="Miller & Benetti & Michat" baseAuthorityYear="2023" class="Insecta" family="Dytiscidae" genus="Amarodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amarodytes pulchellus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="pulchellus">Amarodytes pulchellus</taxonomicName>
|
||
[handwritten] /
|
||
<collectingCountry id="0DCD406F90AEF51A865D28AE366F682B" name="Peru">Peru</collectingCountry>
|
||
,
|
||
<collectingCounty id="7A945F0B6305F8BBF204B13F72B49F4E">Prov.</collectingCounty>
|
||
<collectingRegion id="D642103EAEB074C4D699AC03DD618FCF" country="Peru" name="Huanuco">Huanuco</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="2ACA97A70903DD97C96E96C7420D6C73">Rio Yuyapichis</collectingMunicipality>
|
||
,
|
||
<location id="0EF7E380E4F3AE7462E229DFE0A3A7E0" LSID="urn:lsid:plazi:treatment:34C12EF8D2285E0982A3EA2FCBDCF4D0:0EF7E380E4F3AE7462E229DFE0A3A7E0" country="Peru" county="Prov." latitude="-9.616667" longLatPrecision="1299" longitude="-74.933334" municipality="Rio Yuyapichis" name="Biol. Stat. Panguana. Oestl. Ort." stateProvince="Huanuco">
|
||
Biol. Stat. Panguana.
|
||
<normalizedToken id="A907639E06D50D64FBD4B5D6D47E34F5" originalValue="Östl">Oestl</normalizedToken>
|
||
. Ort.
|
||
</location>
|
||
<geoCoordinate id="D3C820ECDDCCE014DD94D54AE3730BBA" degrees="9" direction="south" minutes="37" orientation="latitude" precision="925" value="-9.616667">9°37'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FAD7E6AF16612BF4F87475ED63BE5D38" degrees="74" direction="west" minutes="56" orientation="longitude" precision="925" value="-74.933334">74°56'W</geoCoordinate>
|
||
<collectingDate id="C9E734EEB7393D8A92690C2F23DEAF4A" value="2003-04-06" valueMax="2003-04-17" valueMin="2003-04-06">6.-17.April 2003</collectingDate>
|
||
, Malaise [handwritten], leg. H., J. u.
|
||
<collectorName id="222C055AB9EF52B4AA4075FD75FCA647">E. -G. Burmeister</collectorName>
|
||
"
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="A30FEF8B0AED47D14265A6C05688134E" collectingDate="2015-12" collectorName="J. Monzon" country="PERU" county="Dept." latitude="-9.616667" location="Rio Yuyapichis" longLatPrecision="1299" longitude="-74.933334" municipality="Panguana" specimenCount="2" stateProvince="Huanuco" typeStatus="paratype">
|
||
<specimenCount id="9262513D66F5FF58C494FB3C394505E9" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
|
||
(ZSMG) labeled, "
|
||
<collectingCountry id="2DA52A26CD7B6850A73DEFB9AD4D69C3" name="Peru">PERU</collectingCountry>
|
||
,
|
||
<collectingCounty id="AA8BB310297F7F9AE9B7EC0D67076CC7">Dept.</collectingCounty>
|
||
<collectingRegion id="8727F0B60697417B0E27C2279E40657F" country="Peru" name="Huanuco">Huanuco</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="C2150A45C622AEAAC296B37A0D6B3A00">Panguana</collectingMunicipality>
|
||
ACP,
|
||
<location id="E735C4FD07BDC8468D97EC73D198264C" LSID="urn:lsid:plazi:treatment:34C12EF8D2285E0982A3EA2FCBDCF4D0:E735C4FD07BDC8468D97EC73D198264C" country="PERU" county="Dept." latitude="-9.616667" longLatPrecision="1299" longitude="-74.933334" municipality="Panguana" name="Rio Yuyapichis" stateProvince="Huanuco">Rio Yuyapichis</location>
|
||
<geoCoordinate id="B74026E36D33A33E4DACE6A9775B8F23" degrees="9" direction="south" minutes="37" orientation="latitude" precision="925" value="-9.616667">9°37'S</geoCoordinate>
|
||
-
|
||
<geoCoordinate id="4F981D6AE1ECF6EC7F8BFE96DA6BBA17" degrees="74" direction="west" minutes="56" orientation="longitude" precision="925" value="-74.933334">74°56'W</geoCoordinate>
|
||
, at blacklight,
|
||
<collectingDate id="4FA6571B86BC8AE17E294E8E4FB4AC59" value="2015-12">XII.2015</collectingDate>
|
||
<collectorName id="FB1494E0BE3937460CA384A1979985D3">J. Monzon</collectorName>
|
||
leg."
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="62FE6CA12B35B391BC6895BFD2DC744B" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectingMethod="blacklight trap" country="Peru" elevation="230" specimenCount="4" stateProvince="Huanuco" typeStatus="paratype">
|
||
<specimenCount id="F7FAA69C6A364D3C2C1B1547856B50E4" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
|
||
(ZSMG) same as previous except
|
||
<normalizedToken id="E5D9A1536D9DD0507A44EEAA12EB5EBA" originalValue="“…">"...</normalizedToken>
|
||
/
|
||
<quantity id="83A998F0FC1EB8B98BACA9B278C28E13" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">
|
||
<elevation id="28C797F71398957480DE2BA436F33045" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">230 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="173D2A7CC56859EE242B095BE6602713">blacklight trap</collectingMethod>
|
||
/
|
||
<taxonomicName id="D4EF2D70F5BC00C7CB8406E550A18453" class="Insecta" family="Dytiscidae" genus="Hydrodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrodessus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">Hydrodessus</taxonomicName>
|
||
sp. 1 [handwritten], Hendrich dt. 2020"
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="5D254FF3B0DDCA9FEA3254773892AE1A" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
|
||
(ZSMG) same as previous except
|
||
<normalizedToken id="78EAA414B1EB5EC3BAC1F2BCF67D3820" originalValue="“…/I.2016”">
|
||
".../
|
||
<date id="39DB1371559CF18FEC8C6D86C1AC3F00" value="2016-01">I.2016</date>
|
||
"
|
||
</normalizedToken>
|
||
;
|
||
<materialsCitation id="5630B0CA8B8F2178E09CC1EF82D4FC27" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Peru" determinerName="H. J. Bremer" specimenCount="4" stateProvince="Huanuco" typeStatus="paratype">
|
||
<specimenCount id="07BAD2930537451D0669315545A53579" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
|
||
(ZSMG) same as previous except
|
||
<normalizedToken id="B756A42AA33D2489521C6CC390CEB6D4" originalValue="“…">"...</normalizedToken>
|
||
/V-VI.2017/
|
||
<taxonomicName id="288671A145BFE8E5F553B0CD2BEF5730" class="Insecta" family="Dytiscidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
|
||
?
|
||
<taxonomicName id="0A00E0AAE5B010EEF81AFC12C77E5F0E" class="Insecta" family="Dytiscidae" genus="Hydrodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrodessus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">Hydrodessus</taxonomicName>
|
||
sp. [handwritten],
|
||
<determinerName id="9EF1810515F8D51E3AD1BD15DB7ABA30">H.J. Bremer</determinerName>
|
||
det. 2018"
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="CF3617ABA5A1A924D619F8DB42176F9C" collectingDate="2007-10-09" collectorName="Burmeister., Each" country="Peru" location="Lux" specimenCount="2" stateProvince="Huanuco" typeStatus="paratype">
|
||
<specimenCount id="04C4D443F93F78D69323A2A0402DF5B5" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
|
||
(ZSMG) same as previous except
|
||
<normalizedToken id="75CBD34246B5DF847E67799EB9F3F239" originalValue="“…">"...</normalizedToken>
|
||
/
|
||
<location id="7AFAD4DCF64FC631496BC9202070B3FB" LSID="urn:lsid:plazi:treatment:34C12EF8D2285E0982A3EA2FCBDCF4D0:7AFAD4DCF64FC631496BC9202070B3FB" country="Peru" name="Lux" stateProvince="Huanuco">Lux</location>
|
||
- 20.9.-
|
||
<collectingDate id="F6DE4E8C5C0D45A1775D8836ECEFE9C5" value="2007-10-09">9.10.2007</collectingDate>
|
||
leg.
|
||
<collectorName id="BA7EFF4BDE9271D4C08303E0D3515E98">Burmeister.</collectorName>
|
||
"
|
||
<collectorName id="2CFE741265E3E5124AFE92F62F62DE6C">Each</collectorName>
|
||
<typeStatus id="2B8AC581B17D94634C249D5274054A99">paratype</typeStatus>
|
||
with
|
||
<normalizedToken id="92527611E7BA402DB98BF17F55248628" originalValue="“…">"...</normalizedToken>
|
||
/
|
||
<typeStatus id="639FF0C196811FA05375B438B95EF18E">PARATYPE</typeStatus>
|
||
<taxonomicName id="C869059BDC4E262F60978F4B7EF1B7F7" authorityName="Miller, Benetti & Michat" authorityYear="2023" family="Dytiscidae" genus="Miradessus" kingdom="Animalia" lsidName="Miradessus beni" order="Coleoptera" pageId="0" pageNumber="13" rank="species" species="beni">
|
||
<emphasis id="A06E105D85E1852FF342910DB9278A67" italics="true" pageId="0" pageNumber="13">Miradessus beni</emphasis>
|
||
</taxonomicName>
|
||
Miller, Benetti & Michat, 2023 [blue label with black line border]."
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |