375 lines
45 KiB
XML
375 lines
45 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1175.107636" ID-Pensoft-Pub="1313-2970-1175-163" ID-Pensoft-UUID="CE03B1B394CF56B48E8F7E5026CD639C" ID-ZooBank="E88FF072E8C945AEA4944B2A5F7DD7E8" ModsDocID="1313-2970-1175-163" checkinTime="1692265381537" checkinUser="pensoft" docAuthor="Yin, Wenqi, Zhu, Pingzhou & Shi, Hongliang" docDate="2023" docId="2B37CF9BBA7554048D7E727CE42E0AEB" docLanguage="en" docName="ZooKeys 1175: 163-185" docOrigin="ZooKeys 1175" docPubDate="2023-08-16" docSource="http://dx.doi.org/10.3897/zookeys.1175.107636" docTitle="Pterostichus (Orientostichus) pemphis Yin & Zhu & Shi 2023, sp. nov." docType="treatment" docUuid="69D91D53-ECC3-4E35-B0FE-7CE16F72A5B6" docUuidSource="ZooBank" docVersion="1" id="CE03B1B394CF56B48E8F7E5026CD639C" lastPageNumber="163" masterDocId="CE03B1B394CF56B48E8F7E5026CD639C" masterDocTitle="Taxonomic study on the subgenus Orientostichus, the Pterostichus pulcher species group (Coleoptera, Carabidae, Pterostichus)" masterLastPageNumber="185" masterPageNumber="163" pageNumber="163" updateTime="1692265381537" updateUser="pensoft">
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<mods:titleInfo id="25B5CEEDEA5357579DF94ACC7626F174">
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<mods:title id="694345C566FC4434DB4C523373FF7A26">Taxonomic study on the subgenus Orientostichus, the Pterostichus pulcher species group (Coleoptera, Carabidae, Pterostichus)</mods:title>
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<mods:namePart id="8F466B12738125042A7EA8499FA9BA52">Yin, Wenqi</mods:namePart>
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<mods:affiliation id="2B4DE451EB3F7D06E7163D49915E8017">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<mods:roleTerm id="965CDBD9CCA24DE77227810B29821F1E">Author</mods:roleTerm>
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<mods:namePart id="DA72F4B88D531644B3C1006D3B38175B">Zhu, Pingzhou</mods:namePart>
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<mods:nameIdentifier id="24906450060E3A3BABDA4D3BD37B6AFC" type="ORCID">https://orcid.org/0000-0003-2139-6764</mods:nameIdentifier>
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<mods:affiliation id="03ACEF4EC0FF52AE23B7AEE3DD6E4A54">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100093, China</mods:affiliation>
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<mods:name id="9ECF5B2F81B692CDAFD32B794E18D344" type="personal">
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<mods:namePart id="EE7FF11515DA6DA7280E65C993FC7285">Shi, Hongliang</mods:namePart>
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<mods:nameIdentifier id="395BC55F9FC07079E9C1248E0E228E40" type="ORCID">https://orcid.org/0000-0002-9989-5830</mods:nameIdentifier>
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<mods:affiliation id="75123A17C09270BF496F7753CF61AE21">College of Forestry, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
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<mods:nameIdentifier id="7260E3078C21897D14B454E208004FA0" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
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<mods:title id="33BC1E5CAB808F535842D3327C611720">ZooKeys</mods:title>
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<mods:date id="FDFFB59ABCF193F9DB1F85865AEBB498">2023</mods:date>
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<mods:number id="F9DA5149771B2D683C8E3DE608ECF6FE">2023-08-16</mods:number>
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<mods:number id="310C7FB802F2E0761F270BF865D88DC4">1175</mods:number>
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<mods:url id="F29944C9B53766B55AC90425C3216D7D">http://dx.doi.org/10.3897/zookeys.1175.107636</mods:url>
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<treatment id="2B37CF9BBA7554048D7E727CE42E0AEB" LSID="urn:lsid:zoobank.org:act:69D91D53-ECC3-4E35-B0FE-7CE16F72A5B6" httpUri="http://treatment.plazi.org/id/2B37CF9BBA7554048D7E727CE42E0AEB" lastPageNumber="163" pageId="0" pageNumber="163">
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<subSubSection id="89D5E339DC716C03DFBA2F39D1BF2413" pageId="0" pageNumber="163" type="nomenclature">
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<paragraph id="1B85100E7283D473A8C19D1F139E3C97" pageId="0" pageNumber="163">
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<taxonomicName id="18170A1AF948246EF0D29E754077ADC1" LSID="https://zoobank.org/69D91D53-ECC3-4E35-B0FE-7CE16F72A5B6" authority="Yin & Zhu & Shi, 2023" authorityName="Yin & Zhu & Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) pemphis" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pemphis" status="sp. nov." subGenus="Orientostichus">Pterostichus (Orientostichus) pemphis</taxonomicName>
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<taxonomicNameLabel id="51D1758F1A692799DFC0DB34C7DBB717" pageId="0" pageNumber="163">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="AA8A274EBD4265C62AA08B226661994F" pageId="0" pageNumber="163" type="description">
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<paragraph id="A48F616203F13C929BF1E31F1EE9CA35" pageId="0" pageNumber="163">
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<figureCitation id="AFA3D2E8DEBDF47908B4920577277648" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus and labels of Pterostichus pulcher species group 1 P. pemphis sp. nov., holotype 2 P. orbicollis sp. nov., holotype 3 P. leo sp. nov., holotype 4 P. liyuani sp. nov., holotype 5 P. condylus sp. nov., holotype 6 P. jialini sp. nov., holotype. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1175.107636.figures1-6" httpUri="https://binary.pensoft.net/fig/893313" pageId="0" pageNumber="163">Figs 1</figureCitation>
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<figureCitation id="E0E13ECF92ED2804530C5F98C54801E5" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">, 7</figureCitation>
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<figureCitation id="EF61C09107BA97E812FFE2BEC5118186" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">, 13</figureCitation>
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<figureCitation id="94743A2DE843BC1F63E592B6534FF8DF" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky & Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">, 19</figureCitation>
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<figureCitation id="87C6E1901E2C98287DF79CBFD9A7468E" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky & Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">, 34</figureCitation>
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<figureCitation id="585642473B69C5BB3A5F465F10166F6C" captionStart="Figures 40, 41" captionStartId="F8" captionText="Figures 40, 41. Female genitalia of Pterostichus (Orientostichus) pemphis sp. nov., paratype from Shuihaizi, Puge county 40 ventral view and inner lateral view of ovipositor. 41. Female reproductive system. Scale bars: 0.5 mm (40); 1 mm (41). Abbreviations: g 1: gonocoxite 1, g 2: gonocoxite 2, es: ensiform setae, ns, nematiform setae, bc: bursa copulatrix, co: common oviduct, sc: seminal canal, rc: receptaculum, sg: spermathecal gland, spc: spermathecal canal." figureDoi="10.3897/zookeys.1175.107636.figures40-41" httpUri="https://binary.pensoft.net/fig/893320" pageId="0" pageNumber="163">, 40</figureCitation>
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<figureCitation id="2ED70BDDEC2ABD6D97C59AD4BCEF67F1" captionStart="Figures 40, 41" captionStartId="F8" captionText="Figures 40, 41. Female genitalia of Pterostichus (Orientostichus) pemphis sp. nov., paratype from Shuihaizi, Puge county 40 ventral view and inner lateral view of ovipositor. 41. Female reproductive system. Scale bars: 0.5 mm (40); 1 mm (41). Abbreviations: g 1: gonocoxite 1, g 2: gonocoxite 2, es: ensiform setae, ns, nematiform setae, bc: bursa copulatrix, co: common oviduct, sc: seminal canal, rc: receptaculum, sg: spermathecal gland, spc: spermathecal canal." figureDoi="10.3897/zookeys.1175.107636.figures40-41" httpUri="https://binary.pensoft.net/fig/893320" pageId="0" pageNumber="163">
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, 41 Chinese vernacular name:
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<normalizedToken id="27B55BA46CA5748B81CA342DA5E3B3C1" originalValue="泡通缘步甲">泡通缘步甲</normalizedToken>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EB6769724DF00FA6A5DE0870E5F48347" pageId="0" pageNumber="163" type="type locality">
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<paragraph id="A3C9D180EE005C96C5CBA8EEDE604FD0" pageId="0" pageNumber="163">Type locality.</paragraph>
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<paragraph id="428E76E37EAA3E385C528DE67EFC1138" pageId="0" pageNumber="163">
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China, Sichuan province: Liangshan Yi Autonomous Prefecture, Puge county, Shuihaizi wind power station (
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<geoCoordinate id="ECB9A1C575A66A7013C4E7108E6E4564" degrees="27.33" direction="north" orientation="latitude" precision="555" value="27.33">27.33N</geoCoordinate>
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,
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<geoCoordinate id="FEB80D77D147F222339494CD7FEB78EE" degrees="102.45" direction="east" orientation="longitude" precision="555" value="102.45">102.45E</geoCoordinate>
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, alt 3515 m).
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</paragraph>
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</subSubSection>
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<subSubSection id="CF63A78377080FB2FE48D5B392C731E0" pageId="0" pageNumber="163" type="type material">
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<paragraph id="5FD5850AEF58EF549BD3602318BBA35A" pageId="0" pageNumber="163">Type material.</paragraph>
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<paragraph id="084E85223917B3A07FAEE4348A1AB9BF" pageId="0" pageNumber="163">
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<emphasis id="4551AFB4EFA2F7B724FB347AA75B78BB" bold="true" italics="true" pageId="0" pageNumber="163">Holotype</emphasis>
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: ♂: "China: Sichuan prov., Puge County, Shuihaizi wind power station, meadow + rhodo. + fir. 3515 m, N27.3355, E102.4461", "2018.VII.21, pitfall trap. Shi HL, Yan WF, Zhu PZ & Jiang ZY lgt., IZAS & BJFU exp. 2018", "HOLOTYPE ♂
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<taxonomicName id="B3D974ADFFADD4A97ED425C3E3C446E3" authorityName="Yin & Zhu & Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) pemphis" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pemphis" subGenus="Orientostichus">Pterostichus (Orientostichus) pemphis</taxonomicName>
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sp. nov. det. Yin & Shi, 2022" [red label];
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<emphasis id="69F81636AE4F0884A3B88E477544B297" bold="true" italics="true" pageId="0" pageNumber="163">Paratypes</emphasis>
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(a total of 96 ♂ and 79 ♀): 19 ♂ and 24 ♀: the same data as holotype but labeled as paratype; 1 ♂ 2 ♀: "China: Sichuan prov., Butuo county, Juesa vill., Pukui mt., alpine meadow, 3248 m, N27.4233, E102.7604", "2015.VII.26, pitfall trap, Shi HL, Liu B & Ma YL lgt., BJFU exp. 2015", "PARATYPE of
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<taxonomicName id="6FFC7273A90DB3854E98C11250D0E952" authorityName="Yin & Zhu & Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) pemphis" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pemphis" subGenus="Orientostichus">Pterostichus (Orientostichus) pemphis</taxonomicName>
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sp. nov. det. Yin & Shi, 2022" [red label]; 76 ♂ and 53 ♀ (CHYL): "China, Sichuan province, Liangshan autonomous prefecture, Zhaojue county, Jiefanggou town, 3000 m, 2022.VI, leg by Li Yuan", "PARATYPE of
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<taxonomicName id="41BB06CEB8130CD9ECF776289816D30E" authorityName="Yin & Zhu & Shi" authorityYear="2023" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Orientostichus) pemphis" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pemphis" subGenus="Orientostichus">Pterostichus (Orientostichus) pemphis</taxonomicName>
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sp. nov. det. Yin & Shi, 2022" [red label].
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</paragraph>
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</subSubSection>
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<subSubSection id="B60E1170ABB41E0B8CC2A2E46D9E7FBA" pageId="0" pageNumber="163" type="diagnosis">
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<paragraph id="39BCD60F8BDDB885E009CAD08F6BE2BF" pageId="0" pageNumber="163">Diagnosis.</paragraph>
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<paragraph id="31F349A8A3692569B7490A9198A6C125" pageId="0" pageNumber="163">
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Elytral intervals 3 and 5 each with ≥ 3 large foveate discal pores, interval 7 without discal pore. Antennomere 3 with accessory setae. Pronotum nearly circular, evenly curved before posterior angles; posterior angles rounded, apex with an additional blunt denticle which distinctly prominent (Fig.
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<figureCitation id="8CFF80E73582AE821FC4764129A583FC" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">7</figureCitation>
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); lateral margins with 2-4 mid-lateral setae. Male sternite VII with a well-defined smooth tubercle (Fig.
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<figureCitation id="AEFB04D409F39CBEF1173BA28965C36C" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky & Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">34</figureCitation>
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). Apical lamella of aedeagus wide and short (LL/LW = 1.35-1.55), apex rounded (Fig.
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<figureCitation id="8937ADC353AA6A5C1F309010B0A1348F" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">13</figureCitation>
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); endophallus with right branch of bsp tuberculate on dorsal surface (Fig.
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<figureCitation id="8A46A89F7079FD1442A6E70FA935E54D" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky & Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">19</figureCitation>
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).
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</paragraph>
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<caption id="58791A3E892D12DFA9CDADDA3FF8CA61" doi="10.3897/zookeys.1175.107636.figures1-6" httpUri="https://binary.pensoft.net/fig/893313" pageId="0" pageNumber="163" start="Figures 1–6" startId="F1">
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<paragraph id="120DDEAEF48942330F84C9C321E98D26" pageId="0" pageNumber="163">
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<emphasis id="20D36CD5A4231AEBA649B895C8F1FB1F" bold="true" pageId="0" pageNumber="163">Figures 1-6.</emphasis>
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Habitus and labels of
|
||
<taxonomicName id="EDF2223A175432CCE4FA02E7394FAF52" authorityName="Sciaky & Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher">
|
||
<emphasis id="26D78DC2017098F80D5C710FB8CC021E" italics="true" pageId="0" pageNumber="163">Pterostichus pulcher</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
<emphasis id="8DA94C80CB942ED8592523F446C64E18" bold="true" pageId="0" pageNumber="163">1</emphasis>
|
||
<taxonomicName id="1F2B4EE8009ABA3EA37D8FB87B32625C" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
|
||
<emphasis id="B5633FADF0A7F2C77C5A2045E557888D" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="B715FBD56B57CF0B33D16C5333C7E5C6" bold="true" pageId="0" pageNumber="163">2</emphasis>
|
||
<taxonomicName id="469838E16CA121ADC5A3FA62DEA13A34" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
|
||
<emphasis id="BC347549590D63CBA38F23B6CC38F1D7" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="E64D4A366DED6748CC45786267006DA7" bold="true" pageId="0" pageNumber="163">3</emphasis>
|
||
<taxonomicName id="A5C6E112DA1C66CDF308D69DF2DAD1FC" lsidName="P. leo" pageId="0" pageNumber="163" rank="species" species="leo">
|
||
<emphasis id="A2A7DB66979C16A4239FDC07DBAAF57A" italics="true" pageId="0" pageNumber="163">P. leo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="CF27BC32763E2FCBD891916A0E1F4D0C" bold="true" pageId="0" pageNumber="163">4</emphasis>
|
||
<taxonomicName id="4043CECF05099EFF415635AEF16081D7" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
|
||
<emphasis id="45F1C253891E37F00CBE2774E1619010" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="B82FCC92E2A91BEE3F8350873ED174BD" bold="true" pageId="0" pageNumber="163">5</emphasis>
|
||
<taxonomicName id="8082255A278E342E8C7308C14AAB5DF5" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
|
||
<emphasis id="20C590CAEB23F8AF5FAB4FF4697EAC6B" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="9BF76F0F501FC45119CA7AD3CBFC1B3C" bold="true" pageId="0" pageNumber="163">6</emphasis>
|
||
<taxonomicName id="D92B104D133130F541B03591805C303F" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
|
||
<emphasis id="5B2AD4F80586EAD123D559C5E894042F" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="BC2DC9A554023F98C46AAE102D4F6E0A" doi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163" start="Figures 7–12" startId="F2">
|
||
<paragraph id="73010D83E079D4BE273496C96336EA81" pageId="0" pageNumber="163">
|
||
<emphasis id="2EC4620C9620CACE05FC723709B47DFE" bold="true" pageId="0" pageNumber="163">Figures 7-12.</emphasis>
|
||
Pronota of
|
||
<taxonomicName id="9DA01BDC60F18E454A30A71DB0EAB030" authorityName="Sciaky & Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher">
|
||
<emphasis id="5B28FB6C48C2635338E172CF8689CDC4" italics="true" pageId="0" pageNumber="163">Pterostichus pulcher</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
<emphasis id="68C371C78586755126621332F8774B36" bold="true" pageId="0" pageNumber="163">7</emphasis>
|
||
<taxonomicName id="F464F99979D1A67A234E68FDC8E098C7" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
|
||
<emphasis id="469D099D1D8F05E94C108A86803802F5" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="E478A0C3B3074DCE9148B5009574D58A" bold="true" pageId="0" pageNumber="163">8</emphasis>
|
||
<taxonomicName id="D254FA35600B5150583C38359260AAC6" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
|
||
<emphasis id="A815C50A58EF2D3C65494914CD9B4648" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="A76E70E208E89AD12E4710F8355AF0ED" bold="true" pageId="0" pageNumber="163">9</emphasis>
|
||
<taxonomicName id="2E6A9DA287F1B8B80321D67EA6F10CE2" lsidName="P. leo" pageId="0" pageNumber="163" rank="species" species="leo">
|
||
<emphasis id="873DE0599B4689B0EF16BF0160981675" italics="true" pageId="0" pageNumber="163">P. leo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="1A24C0A705996F53C9815B1B17A10874" bold="true" pageId="0" pageNumber="163">10</emphasis>
|
||
<taxonomicName id="02F4104C85FB9D4693A6B8B1646FA88A" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
|
||
<emphasis id="A4F649F0F773A699417B0F79EBF68576" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="C73E932AD4783DAA6ED23D1E3A9705FD" bold="true" pageId="0" pageNumber="163">11</emphasis>
|
||
<taxonomicName id="C9509F3E8E3B119A78AAA4D51E93FD78" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
|
||
<emphasis id="43512F7B764E7ED9905A1EF7547687CA" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="1698C4BA4F214A7A004EE0E2B0ED8B98" bold="true" pageId="0" pageNumber="163">12</emphasis>
|
||
<taxonomicName id="E129E08B35116018BC327D7963F77189" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
|
||
<emphasis id="CAA2EFD52769978890067C3D8ACE5996" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="3ECE2F3247A47C350ACF83112E49F9CE" pageId="0" pageNumber="163" type="comparisons">
|
||
<paragraph id="78852382FBCA646AA8FEB587E45CF00B" pageId="0" pageNumber="163">Comparison.</paragraph>
|
||
<paragraph id="0A77C58DD230CDF2C8EF0D9E94CFB67D" pageId="0" pageNumber="163">
|
||
From the external features,
|
||
<taxonomicName id="B8A76661D125317E5E827D0888F1D732" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
|
||
<emphasis id="FBA9A948369252CF4D6A1ACED2F42AEA" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to the following two new species,
|
||
<taxonomicName id="74F8B8B0A3C8EEF41387E97C7B008EB0" lsidName="P. leo" pageId="0" pageNumber="163" rank="species" species="leo">
|
||
<emphasis id="748526A9D41272CF5FBE666E8A816788" italics="true" pageId="0" pageNumber="163">P. leo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="07B4DA8C474E4893DAFAC66318CF70EA" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
|
||
<emphasis id="C54086837CC89753DFB9057372AC390C" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. as all these three have accessory setae on antennomere 3, nearly circular pronotum, and dorsally tuberculate bsp on endophallus.
|
||
</paragraph>
|
||
<paragraph id="DBAC5A5DFCC512554E4CE8A4D1C499A6" pageId="0" pageNumber="163">
|
||
The present new species is different from
|
||
<taxonomicName id="7ADA152ED292EB213CA10FC2C24D2BED" lsidName="P. leo" pageId="0" pageNumber="163" rank="species" species="leo">
|
||
<emphasis id="E61AA1683A9106715173FC5973B6E303" italics="true" pageId="0" pageNumber="163">P. leo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by: (1) pronotum disc with distinct transverse wrinkles, but at most with very faint wrinkles in the latter species; (2) pronotal lateral margins evenly arched before posterior angles, posterior angles with a distinctly prominent denticle (Fig.
|
||
<figureCitation id="4DE02C9C67C3BD70EED3F39DD7E39073" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">7</figureCitation>
|
||
), but in the latter species, pronotal lateral margins slightly sinuate before posterior angles, posterior angles with a smaller denticle (Fig.
|
||
<figureCitation id="4507FD1681E175B29FC4C8634B7A8771" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">9</figureCitation>
|
||
); (3) lateral margins with ≥ 2 mid-lateral setae, but with only one mid-lateral seta in the latter species; (4) male sternite VII with a smooth tubercle, but not modified in the latter species; (5) apical lamella of male genitalia short and straight, but strongly elongate and turned upward in the latter species.
|
||
</paragraph>
|
||
<paragraph id="78509CFB6D203C112B7E8A73134659F6" pageId="0" pageNumber="163">
|
||
Compared to
|
||
<taxonomicName id="0961274A3DA2A8F3A30BD215FF34EDDB" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
|
||
<emphasis id="5895D778BDDAECCF29179C0F8292AFE0" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="90C3A33BEDB7458C9FE8E24EB9128CD3" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
|
||
<emphasis id="922A5EF9230742AAF2E90B96B7545195" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is different in: (1) posterior angles with a distinctly prominent denticle (Fig.
|
||
<figureCitation id="907461405E95F9ACF89B2862DB4DF2AF" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">7</figureCitation>
|
||
), but not dentate in
|
||
<taxonomicName id="AB44532E534F97158138C5B72D0F44D7" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
|
||
<emphasis id="08220CC84F8F2CAFCFDDA380874BC0BF" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="9AB9B7B6E8F4B16779CC0290AFEA149F" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">8</figureCitation>
|
||
); (2) lateral margins with ≥ 2 mid-lateral setae, but with only one such seta in the latter species; (3) apical lamella of aedeagus a little longer than in
|
||
<taxonomicName id="E6AA3F66B55E1337F6B3376731477B24" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
|
||
<emphasis id="6D28B9CDD38748C2555DA0D47E3B5F8F" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
|
||
</taxonomicName>
|
||
(LL/LW = 1.35-1.55 vs 1.1-1.25).
|
||
</paragraph>
|
||
<paragraph id="A1BE686B4D5640F3978C70AAC5E5CFE9" pageId="0" pageNumber="163">
|
||
The present new species cohabitates with
|
||
<taxonomicName id="4EB95517C9B0A2B5E1762992DBB99FB5" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
|
||
<emphasis id="0DB9DEF8F25D2F6979790E02BDA7CF54" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in Shuihaizi of Puge county. These two species can be easily distinguished by the differences on the shape of pronotum, the chaetotaxy on antennomere 3, and the sexual modification on male sternite VII (
|
||
<taxonomicName id="934EC6DBAE461400EB61CC568A3DA9B3" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
|
||
<emphasis id="A4378A7BEB075609FE41EC1C80838EE2" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. with a larger tubercle).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E1EA0118FCF61B14A9FA1A54938773B9" pageId="0" pageNumber="163" type="description">
|
||
<paragraph id="9ED501557D44B5EAEC2072AFD5E394B2" pageId="0" pageNumber="163">Description.</paragraph>
|
||
<paragraph id="C49F2618B0D56323AA60165B739C7425" pageId="0" pageNumber="163">
|
||
BL = 14.5-18.2 mm, BW = 5.0-6.0 mm, dorsal surface and appendages black, elytra often a little reddish brown. Antennomere 3 with accessory setae on apical 2/3 in addition to the primary setae forming apical ring. Pronotum nearly circular, PW/PL = 1.20-1.24, widest near anterior 1/3; anterior margin wider than posterior margin; strongly narrowed to base (PBW/PW = 0.61-0.65); lateral margins evenly arched from anterior angles to posterior angles, not sinuate before posterior angles; posterior angles rounded, apex with an additional blunt denticle which distinctly prominent; lateral margins with 2-4 mid-lateral setae near maximum width; basal foveae impunctate, inner and outer grooves straight, partly fused at base, outer groove a little shorter than inner one, area between them depressed (Fig.
|
||
<figureCitation id="29D916D3433864186D6B05C2C0AFA706" captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Pronota of Pterostichus pulcher species group 7 P. pemphis sp. nov., holotype 8 P. orbicollis sp. nov., holotype 9 P. leo sp. nov., holotype 10 P. liyuani sp. nov., holotype 11 P. condylus sp. nov., holotype 12 P. jialini sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures7-12" httpUri="https://binary.pensoft.net/fig/893314" pageId="0" pageNumber="163">7</figureCitation>
|
||
); disc with fine transverse wrinkles aside median line. Elytra oblong; parascutellar pore usually absent, but present in some specimens from Zhaojue county; intervals 3 and 5 each with ≥ 3 large foveate discal pores; interval 7 without discal pore. Male sternite VII with well-defined smooth tubercle on middle (Fig.
|
||
<figureCitation id="A6D990C26AE5E99C73EF72E074D143F3" captionStart="Figures 34–39" captionStartId="F7" captionText="Figures 34 - 39. Male sternum VII of Pterostichus pulcher species group spp. 34 P. pemphis sp. nov., holotype 35 P. orbicollis sp. nov., holotype 36 P. liyuani sp. nov., holotype 37 P. condylus sp. nov., holotype 38 P. jialini sp. nov., holotype 39 P. pulcher Sciaky & Allegro, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures34-39" httpUri="https://binary.pensoft.net/fig/893319" pageId="0" pageNumber="163">34</figureCitation>
|
||
). Median lobe of aedeagus stout, strongly curved near basal 1/3; apical lamella gradually narrowed and deflected ventrally; apical lamella slightly twisted longitudinally, relatively short (LL/LW = 1.35-1.55), apex rounded (Fig.
|
||
<figureCitation id="1DDB34DE8BC236C8D7078DF1D31CBF63" captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Male genitalia of Pterostichus pulcher species group, left lateral view and dorsal view of median lobe of aedeagus 13 P. pemphis sp. nov., holotype 14 P. orbicollis sp. nov., holotype 15 P. leo sp. nov., holotype 16 P. liyuani sp. nov., holotype 17 P. condylus sp. nov., holotype 18 P. jialini sp. nov., holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163">13</figureCitation>
|
||
). Endophallus long, straightly directed ventrally, gonopore opened to ventral-basal direction of aedeagus; vb distinct, oblate, rounded at both apexes; bsp deeply grooved, nearly U-shaped, dorsal surface of right branch folded and strongly hooked forming a tubercle (Fig.
|
||
<figureCitation id="14C078517F060F5A6FE82F44C71D3CBB" captionStart="Figures 19–24" captionStartId="F4" captionText="Figures 19 - 24. Endophallus of Pterostichus pulcher species group, left lateral view and right lateral view 19 P. pemphis sp. nov., paratype from Zhaojue county 20 P. orbicollis sp. nov., paratype from Longzhoushan mountain, Huili county 21 P. leo sp. nov., paratype from Shizishan mountain, Jinyang county 22 P. liyuani sp. nov., paratype from Luojishan mountain, Puge county 23 P. condylus sp. nov., paratype from Mianning county 24 P. pulcher Sciaky & Allegro, paratype from Yizi Yakou, E'bian county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe." figureDoi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163">19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="85EE5C94606BB5FBF1136C0D4C4F352C" doi="10.3897/zookeys.1175.107636.figures13-18" httpUri="https://binary.pensoft.net/fig/893315" pageId="0" pageNumber="163" start="Figures 13–18" startId="F3">
|
||
<paragraph id="6D858E550ABF685974A8814CAB09C425" pageId="0" pageNumber="163">
|
||
<emphasis id="EA06CD782F69DDEE17E9A9908CC6FE71" bold="true" pageId="0" pageNumber="163">Figures 13-18.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="CA1A1711BAA8CF71C8551C6CFE7796BB" authorityName="Sciaky & Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher">
|
||
<emphasis id="D0581E03F536C984E80B18FCB01F9B1B" italics="true" pageId="0" pageNumber="163">Pterostichus pulcher</emphasis>
|
||
</taxonomicName>
|
||
species group, left lateral view and dorsal view of median lobe of aedeagus
|
||
<emphasis id="D11C5BAD67326F368ECE70542A3E1689" bold="true" pageId="0" pageNumber="163">13</emphasis>
|
||
<taxonomicName id="67C0743176467C36ABA8C3FAE1BAC96A" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
|
||
<emphasis id="EE6C7D681D3D5573343C06E3EAC98A52" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="40113C5A0DAE07E9D7DF17D6BA9C23BC" bold="true" pageId="0" pageNumber="163">14</emphasis>
|
||
<taxonomicName id="1A9BA6019B179C570DACFCF8E7F32D58" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
|
||
<emphasis id="04D3877321A15EEF42D7385B96B07106" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="D10D618AB516FC9C5245ECA02D9D0070" bold="true" pageId="0" pageNumber="163">15</emphasis>
|
||
<taxonomicName id="641FD3352B2E1F8C1210EF7E0B3B8260" lsidName="P. leo" pageId="0" pageNumber="163" rank="species" species="leo">
|
||
<emphasis id="18DC125F31ADB312EFC1F1A12F77A151" italics="true" pageId="0" pageNumber="163">P. leo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="96027AF092853309B16900546F1C45CA" bold="true" pageId="0" pageNumber="163">16</emphasis>
|
||
<taxonomicName id="3CF44FDC564E8F673809B45FC059F9F4" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
|
||
<emphasis id="A3EB15DF7FE3D569D4608C73A7B78A81" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="B24B290E63C07ABD616FF85DED35140E" bold="true" pageId="0" pageNumber="163">17</emphasis>
|
||
<taxonomicName id="5DC424FC83D30CA073869CB7F7421104" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
|
||
<emphasis id="E37D2D3ADF7889ACAAA2589D2CDD411D" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype
|
||
<emphasis id="9F326D20816CB94D3ACAA202B6825BFD" bold="true" pageId="0" pageNumber="163">18</emphasis>
|
||
<taxonomicName id="FDBADBAFB4C84123D7718BB848DE5C62" lsidName="P. jialini" pageId="0" pageNumber="163" rank="species" species="jialini">
|
||
<emphasis id="0ACFBF6844744A6DA786CC331416778A" italics="true" pageId="0" pageNumber="163">P. jialini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="FE818AE30AC6C2215994A2D2A5A5C734" doi="10.3897/zookeys.1175.107636.figures19-24" httpUri="https://binary.pensoft.net/fig/893316" pageId="0" pageNumber="163" start="Figures 19–24" startId="F4">
|
||
<paragraph id="E1843E51B59E08124B2801E815975DD4" pageId="0" pageNumber="163">
|
||
<emphasis id="992E8693B6BEFB73ABC9D3529F907AD0" bold="true" pageId="0" pageNumber="163">Figures 19-24.</emphasis>
|
||
Endophallus of
|
||
<taxonomicName id="285E337B347A3B72F4C5CAF9B812C71A" authorityName="Sciaky & Allegro" authorityYear="2013" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pulcher" order="Coleoptera" pageId="0" pageNumber="163" phylum="Arthropoda" rank="species" species="pulcher">
|
||
<emphasis id="81274B15E911CDAD678A121EF7E45CC0" italics="true" pageId="0" pageNumber="163">Pterostichus pulcher</emphasis>
|
||
</taxonomicName>
|
||
species group, left lateral view and right lateral view
|
||
<emphasis id="5E7D4EEEBCF81C1E498DB28A595F5819" bold="true" pageId="0" pageNumber="163">19</emphasis>
|
||
<taxonomicName id="ECF023D37ABEAD8D5D9BD858133FF789" lsidName="P. pemphis" pageId="0" pageNumber="163" rank="species" species="pemphis">
|
||
<emphasis id="C80DFE0A3A0C4DE2AE8AD637067CCC98" italics="true" pageId="0" pageNumber="163">P. pemphis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype from Zhaojue county
|
||
<emphasis id="B21F3CE21AECDA55DDC2EAA41BD1CCE7" bold="true" pageId="0" pageNumber="163">20</emphasis>
|
||
<taxonomicName id="2A55DB0957B92105B2ECA1241CBE1B0D" lsidName="P. orbicollis" pageId="0" pageNumber="163" rank="species" species="orbicollis">
|
||
<emphasis id="D0D5D6F38BDCCF687A5F2DABE621EC36" italics="true" pageId="0" pageNumber="163">P. orbicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype from Longzhoushan mountain, Huili county
|
||
<emphasis id="D65D8C7D084A28F8440CDE6E52C66F49" bold="true" pageId="0" pageNumber="163">21</emphasis>
|
||
<taxonomicName id="5CADC7CC6C0754344B4F167E08681D09" lsidName="P. leo" pageId="0" pageNumber="163" rank="species" species="leo">
|
||
<emphasis id="DC570D7CFD1AC334433160234246C6F6" italics="true" pageId="0" pageNumber="163">P. leo</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype from Shizishan mountain, Jinyang county
|
||
<emphasis id="265AA3998BFE318F80AE554E9CE417F9" bold="true" pageId="0" pageNumber="163">22</emphasis>
|
||
<taxonomicName id="BB40C268D33607B05CE54484ABE42E74" lsidName="P. liyuani" pageId="0" pageNumber="163" rank="species" species="liyuani">
|
||
<emphasis id="BA26F196EA64973F8E7080F26B24F722" italics="true" pageId="0" pageNumber="163">P. liyuani</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype from Luojishan mountain, Puge county
|
||
<emphasis id="7F34A6C3E8CD6F0600D649F1396E38E4" bold="true" pageId="0" pageNumber="163">23</emphasis>
|
||
<taxonomicName id="88152F5660E09569E8E5C13704793DE1" lsidName="P. condylus" pageId="0" pageNumber="163" rank="species" species="condylus">
|
||
<emphasis id="511E1DCEE53BA2FCEC57D334327C66F8" italics="true" pageId="0" pageNumber="163">P. condylus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., paratype from Mianning county
|
||
<emphasis id="657BBDAA2BA1EE3CAF9B22A5E48EA199" bold="true" pageId="0" pageNumber="163">24</emphasis>
|
||
<taxonomicName id="0308FA6CDBA3F54E22339C7CF8EF5D75" lsidName="P. pulcher" pageId="0" pageNumber="163" rank="species" species="pulcher">
|
||
<emphasis id="252C9F9436FD9269E0AA931C7FFE6280" italics="true" pageId="0" pageNumber="163">P. pulcher</emphasis>
|
||
</taxonomicName>
|
||
Sciaky & Allegro, paratype from Yizi Yakou,
|
||
<normalizedToken id="F87C03D761F6FE52A6DDC546F3E621FB" originalValue="E’bian">E'bian</normalizedToken>
|
||
county to Meigu county. Scale bar: 1 mm. Abbreviations: gp: gonopore, gpl: gonopore lobe, bsp: basal sclerotized projection, vb: ventral basal lobe.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="305251C5C88F9E961D104FC8706DB679" pageId="0" pageNumber="163" type="distribution">
|
||
<paragraph id="2E013D7C9180792F6270923BA3DDF250" pageId="0" pageNumber="163">Distribution.</paragraph>
|
||
<paragraph id="663FD7F6D3C74053F10E980FDE422A31" pageId="0" pageNumber="163">
|
||
This species is relatively widely distributed from Puge county to Zhaojue county, in Liangshan Yi autonomous prefecture (Fig.
|
||
<figureCitation id="B824C509AC8C70C5D00D78AAAE95EB4A" captionStart="Figure 42" captionStartId="F9" captionText="Figure 42. Distribution of the taxa of Pterostichus pulcher species group in Sichuan province, China. Brown: P. pemphis sp. nov. Yellow: P. orbicollis sp. nov. Red: P. leo sp. nov. Purple: P. liyuani sp. nov. Blue: P. condylus sp. nov. Green: P. jialini sp. nov. Orange: P. pulcher Sciaky & Allegro." figureDoi="10.3897/zookeys.1175.107636.figure42" httpUri="https://binary.pensoft.net/fig/893321" pageId="0" pageNumber="163">42</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9C1D9BFA9B2169697E9150B8AF41894" pageId="0" pageNumber="163" type="etymology">
|
||
<paragraph id="06DAF43C892869426C3B92E2D2A46FF1" pageId="0" pageNumber="163">Etymology.</paragraph>
|
||
<paragraph id="FD896DA24BF1CBA4BDABE272BF6E8FA4" pageId="0" pageNumber="163">
|
||
The scientific name of the new species is derived from the Greek root
|
||
<emphasis id="2522AB27D8711ED0FD1AC4867F572830" italics="true" pageId="0" pageNumber="163">pemph</emphasis>
|
||
-, meaning blister, referring to the bump-shaped elytral intervals 3 and 5 of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |