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<document id="E5923CE20BFF50AF553BBAE0B64B3E74" ID-CLB-Dataset="56063" ID-DOI="10.5281/zenodo.5305725" ID-GBIF-Dataset="31cb15bf-28f5-4b27-86df-6a1b28cbce41" ID-ISSN="0374-1036" ID-Zenodo-Dep="5305725" ID-ZooBank="E24F1028-C6AC-4323-9ED5-C9B7FF3434ACD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630111997363" checkinUser="marcus" docAuthor="Sekerka, Lukáš" docDate="2016" docId="677FCF20FFCBFFE0FE6EFF3AFF09FC18" docLanguage="en" docName="ActaEntMusNatPra.2016.56.1.275-344.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Charidotis nucleata Boheman 1855" docType="treatment" docVersion="8" lastPageNumber="292" masterDocId="9B46B758FFDAFFF1FFFCFFE6FFBEFFB0" masterDocTitle="Taxonomic and nomenclatural changes in Cassidinae (Coleoptera: Chrysomelidae)" masterLastPageNumber="344" masterPageNumber="275" pageNumber="292" updateTime="1699159820365" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6DE6C091F3EE1C62CA3D7BEC8F771DE0">Taxonomic and nomenclatural changes in Cassidinae (Coleoptera: Chrysomelidae)</mods:title>
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<paragraph id="EF697E36FFCBFFE0FE6EFF3AFC9EFF43" blockId="17.[402,800,220,243]" box="[402,800,220,243]" pageId="17" pageNumber="292">
<heading id="B421C95AFFCBFFE0FE6EFF3AFC9EFF43" bold="true" box="[402,800,220,243]" centered="true" fontSize="8" level="2" pageId="17" pageNumber="292" reason="3">
<taxonomicName id="28D605B5FFCBFFE0FE6EFF3AFC9EFF43" ID-CoL="TNKX" authority="Boheman, 1855" authorityName="Boheman" authorityYear="1855" box="[402,800,220,243]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="nucleata">
<emphasis id="DDA2A224FFCBFFE0FE6EFF3AFC9EFF43" bold="true" box="[402,800,220,243]" pageId="17" pageNumber="292">
<emphasis id="DDA2A224FFCBFFE0FE6EFF3AFDD5FF43" bold="true" box="[402,619,220,243]" italics="true" pageId="17" pageNumber="292">Charidotis nucleata</emphasis>
<bibRefCitation id="8B4703C7FFCBFFE0FD8DFF3AFC9EFF43" author="Boheman" box="[625,800,220,243]" firstAuthor="Boheman" pageId="17" pageNumber="292" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFCBFFE0FF8DFEE7FD67FE9F" pageId="17" pageNumber="292" type="reference_group">
<paragraph id="EF697E36FFCBFFE0FF8DFEE7FD37FEA4" blockId="17.[113,729,257,303]" box="[113,649,257,276]" pageId="17" pageNumber="292">
<treatmentCitationGroup id="CFC65918FFCBFFE0FF8DFEE7FD37FEA4" box="[113,649,257,276]" pageId="17" pageNumber="292">
<treatmentCitation id="6E775827FFCBFFE0FF8DFEE7FE76FEA4" author="BOHEMAN C. H." box="[113,456,257,276]" page="18" pageId="17" pageNumber="292" year="1855">
<taxonomicName id="28D605B5FFCBFFE0FF8DFEE7FE76FEA4" ID-CoL="TNKX" authority="Boheman, 1855: 18" authorityName="Boheman" authorityPageNumber="18" authorityYear="1855" box="[113,456,257,276]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="nucleata">
<emphasis id="DDA2A224FFCBFFE0FF8DFEE7FEA5FEA4" box="[113,283,257,276]" italics="true" pageId="17" pageNumber="292">Charidotis nucleata</emphasis>
<bibRefCitation id="8B4703C7FFCBFFE0FEDDFEE7FE76FEA4" author="Boheman" box="[289,456,257,276]" firstAuthor="Boheman" page="18" pageId="17" pageNumber="292" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855: 18</bibRefCitation>
</taxonomicName>
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(original description).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EF697E36FFCBFFE0FF8DFEFAFD67FE9F" blockId="17.[113,729,257,303]" box="[113,729,284,303]" pageId="17" pageNumber="292">
<treatmentCitationGroup id="CFC65918FFCBFFE0FF8DFEFAFD67FE9F" box="[113,729,284,303]" pageId="17" pageNumber="292">
<treatmentCitation id="6E775827FFCBFFE0FF8DFEFAFE7BFE9F" author="BOHEMAN C. H." box="[113,453,284,303]" page="14" pageId="17" pageNumber="292" year="1855">
<taxonomicName id="28D605B5FFCBFFE0FF8DFEFAFE7BFE9F" ID-CoL="TNL5" authority="Boheman, 1855: 14" authorityName="Boheman" authorityPageNumber="14" authorityYear="1855" box="[113,453,284,303]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="orbifera">
<emphasis id="DDA2A224FFCBFFE0FF8DFEFAFEA6FE9F" box="[113,280,284,303]" italics="true" pageId="17" pageNumber="292">Charidotis orbifera</emphasis>
<bibRefCitation id="8B4703C7FFCBFFE0FEE1FEFAFE7BFE9F" author="Boheman" box="[285,453,284,303]" firstAuthor="Boheman" page="14" pageId="17" pageNumber="292" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">Boheman, 1855: 14</bibRefCitation>
</taxonomicName>
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(original description),
<emphasis id="DDA2A224FFCBFFE0FD76FEFAFD67FE9F" bold="true" box="[650,729,284,303]" pageId="17" pageNumber="292">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFCBFFE0FF8DFEA0FE37FE49" pageId="17" pageNumber="292" type="materials_examined">
<paragraph id="EF697E36FFCBFFE0FF8DFEA0FC3BFEE9" blockId="17.[113,1089,326,505]" box="[113,901,326,345]" pageId="17" pageNumber="292">
<emphasis id="DDA2A224FFCBFFE0FF8DFEA0FF46FEE9" bold="true" box="[113,248,326,345]" pageId="17" pageNumber="292">Type localities.</emphasis>
<taxonomicName id="28D605B5FFCBFFE0FF01FEA0FE19FEE9" authorityName="Boheman" authorityYear="1855" box="[253,423,326,345]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="nucleata">
<emphasis id="DDA2A224FFCBFFE0FF01FEA0FE19FEE9" box="[253,423,326,345]" italics="true" pageId="17" pageNumber="292">Charidotis nucleata</emphasis>
</taxonomicName>
: Brasilia;
<taxonomicName id="28D605B5FFCBFFE0FDF3FEA0FDD3FEE9" authorityName="Boheman" authorityYear="1855" box="[527,621,326,345]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="orbifera">
<emphasis id="DDA2A224FFCBFFE0FDF3FEA0FDD3FEE9" box="[527,621,326,345]" italics="true" pageId="17" pageNumber="292">C. orbifera</emphasis>
</taxonomicName>
: Brasilia ad flumen Amazonis.
</paragraph>
<paragraph id="EF697E36FFCBFFE0FF8DFE87FE2CFE74" blockId="17.[113,1089,326,505]" pageId="17" pageNumber="292">
<emphasis id="DDA2A224FFCBFFE0FF8DFE87FEEFFEC4" bold="true" box="[113,337,353,372]" pageId="17" pageNumber="292">Type material examined.</emphasis>
<materialsCitation id="5FBE746BFFCBFFE0FEA9FE87FD87FE3E" ID-GBIF-Occurrence="3750508342" collectionCode="NHRS" location="Para" pageId="17" pageNumber="292" specimenCode="000021815" specimenCount="1" stateProvince="Para" typeStatus="syntype">
<taxonomicName id="28D605B5FFCBFFE0FEA9FE87FE43FEC4" authorityName="Boheman" authorityYear="1855" box="[341,509,353,372]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="nucleata">
<emphasis id="DDA2A224FFCBFFE0FEA9FE87FE43FEC4" box="[341,509,353,372]" italics="true" pageId="17" pageNumber="292">
<originalDetermination id="CCC8C6CAFFCBFFE0FEA9FE87FE43FEC4" box="[341,509,353,372]" pageId="17" pageNumber="292">Charidotis nucleata</originalDetermination>
</emphasis>
</taxonomicName>
:
<typeStatus id="306DC094FFCBFFE0FDF4FE87FDF1FEC2" box="[520,591,353,372]" pageId="17" pageNumber="292" type="syntype">SYNTYPE</typeStatus>
:
<specimenCount id="F9D0B5BFFFCBFFE0FDA5FE87FD28FEC4" box="[601,662,353,372]" pageId="17" pageNumber="292" type="generic">1 spec.</specimenCount>
, pinned,
<collectingRegion id="2D12B0D4FFCBFFE0FD1FFE87FCB3FEC4" box="[739,781,353,372]" country="Suriname" name="Para" pageId="17" pageNumber="292">Para</collectingRegion>
[w, hw, s] || Bhn. [w, p, s] || Type. [w, p, s] || NHRS-JLKB |
<specimenCode id="BF70D64DFFCBFFE0FED7FE9DFE35FE3E" box="[299,395,379,398]" pageId="17" pageNumber="292">000021815</specimenCode>
[w, p, cb] (
<collectionCode id="89C7E6F3FFCBFFE0FE0EFE9DFD8EFE3E" box="[498,560,379,398]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="17" pageNumber="292">NHRS</collectionCode>
).
</materialsCitation>
<materialsCitation id="5FBE746BFFCBFFE0FDC1FE9DFE2CFE74" ID-GBIF-Occurrence="3750508394" pageId="17" pageNumber="292">
<specimenCount id="F9D0B5BFFFCBFFE0FDC1FE9DFDC5FE3E" box="[573,635,379,398]" pageId="17" pageNumber="292" type="generic">1 spec.</specimenCount>
, pinned,
<collectingRegion id="2D12B0D4FFCBFFE0FD37FE9DFD4BFE3E" box="[715,757,379,398]" country="Suriname" name="Para" pageId="17" pageNumber="292">Para</collectingRegion>
[w, hw, s] || Baly Coll. | 1905—54. [w, p, cb] ||
<taxonomicName id="28D605B5FFCBFFE0FF49FE70FEB0FE19" authorityName="Boheman" authorityYear="1854" box="[181,270,406,425]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="genus">Charidotis</taxonomicName>
| orbifera | Boh | Amazons [w, hw by Baly, cb] || Type Boh | Col Jekel [hw by Baly on underside of the preceding label] (
<collectionCode id="89C7E6F3FFCBFFE0FEBFFE57FE36FE74" box="[323,392,433,452]" pageId="17" pageNumber="292">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="EF697E36FFCBFFE0FF6CFE2DFE37FE49" blockId="17.[113,1089,326,505]" pageId="17" pageNumber="292">
<materialsCitation id="5FBE746BFFCBFFE0FF6CFE2DFE37FE49" ID-GBIF-Occurrence="3750508467" collectionCode="NHRS" country="Brazil" location="Rio Amazonas" pageId="17" pageNumber="292" specimenCode="000021818" specimenCount="1" typeStatus="holotype">
<taxonomicName id="28D605B5FFCBFFE0FF6CFE2DFE88FE6E" authorityName="Boheman" authorityYear="1855" box="[144,310,459,478]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="orbifera">
<emphasis id="DDA2A224FFCBFFE0FF6CFE2DFE88FE6E" box="[144,310,459,478]" italics="true" pageId="17" pageNumber="292">
<originalDetermination id="CCC8C6CAFFCBFFE0FF6CFE2DFE88FE6E" box="[144,310,459,478]" pageId="17" pageNumber="292">Charidotis orbifera</originalDetermination>
</emphasis>
</taxonomicName>
:
<typeStatus id="306DC094FFCBFFE0FEBCFE2DFE2AFE6C" box="[320,404,459,478]" pageId="17" pageNumber="292" type="holotype">HOLOTYPE</typeStatus>
: pinned, Amazo | na [w, hw, s] || Jekel [w, hw, s] || Type. [w, p, s] || NHRS-JLKB |
<specimenCode id="BF70D64DFFCBFFE0FF87FE00FF65FE49" box="[123,219,486,505]" pageId="17" pageNumber="292">000021818</specimenCode>
[w, p, cb] (
<collectionCode id="89C7E6F3FFCBFFE0FEBEFE00FE3EFE49" box="[322,384,486,505]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="17" pageNumber="292">NHRS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFCBFFE0FF8DFDF7FE04FCD8" pageId="17" pageNumber="292" type="discussion">
<paragraph id="EF697E36FFCBFFE0FF8DFDF7FE04FCD8" blockId="17.[113,1089,529,936]" pageId="17" pageNumber="292">
<emphasis id="DDA2A224FFCBFFE0FF8DFDF7FF5EFD98" bold="true" box="[113,224,529,552]" pageId="17" pageNumber="292">Remarks.</emphasis>
<bibRefCitation id="8B4703C7FFCBFFE0FF18FDF7FE2DFD98" author="BOHEMAN" box="[228,403,529,552]" firstAuthor="BOHEMAN" pageId="17" pageNumber="292" refId="ref47955" refString="BOHEMAN C. H. 1855: Monographia Cassididarum. Tomus Tertius. Officina Norstedtiana, Holmiae, 543 pp. + 1 pl." type="book" year="1855">BOHEMAN (1855)</bibRefCitation>
described
<taxonomicName id="28D605B5FFCBFFE0FDFEFDF7FDC4FD98" authorityName="Boheman" authorityYear="1855" box="[514,634,529,552]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="nucleata">
<emphasis id="DDA2A224FFCBFFE0FDFEFDF7FDC4FD98" box="[514,634,529,552]" italics="true" pageId="17" pageNumber="292">C. nucleata</emphasis>
</taxonomicName>
and
<taxonomicName id="28D605B5FFCBFFE0FD57FDF7FCA1FD98" authorityName="Boheman" authorityYear="1855" box="[683,799,529,552]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="orbifera">
<emphasis id="DDA2A224FFCBFFE0FD57FDF7FCA1FD98" box="[683,799,529,552]" italics="true" pageId="17" pageNumber="292">C. orbifera</emphasis>
</taxonomicName>
in the same publication and separated them based on colouration and size. Both taxa have a black ring on elytra, becoming brownish or reddish towards their centre. Boheman described the colouration of
<taxonomicName id="28D605B5FFCBFFE0FC36FDB7FBFEFDD8" authorityName="Boheman" authorityYear="1855" box="[970,1088,593,616]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="orbifera">
<emphasis id="DDA2A224FFCBFFE0FC36FDB7FBFEFDD8" box="[970,1088,593,616]" italics="true" pageId="17" pageNumber="292">C. orbifera</emphasis>
</taxonomicName>
as being darker, almost black, only with a ferruginous tip of scutellum, while in
<taxonomicName id="28D605B5FFCBFFE0FC3AFD97FBFFFD38" authorityName="Boheman" authorityYear="1855" box="[966,1089,625,648]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="nucleata">
<emphasis id="DDA2A224FFCBFFE0FC3AFD97FBFFFD38" box="[966,1089,625,648]" italics="true" pageId="17" pageNumber="292">C. nucleata</emphasis>
</taxonomicName>
it has a distinct brownish-red inner ring. I have studied and compared
<typeStatus id="306DC094FFCBFFE0FCABFD77FC31FD18" box="[855,911,657,680]" pageId="17" pageNumber="292">types</typeStatus>
of both taxa and could not find reasonable characters to separate the two taxa besides colouration. The shape of clypeus, ratio of antennomeres and punctation of elytra are identical and I hereby synonymize the two taxa. As both were described in the same publication I followed the Principle of the first reviser (
<bibRefCitation id="8B4703C7FFCBFFE0FEBFFCF7FE7AFC98" author="ICZN" box="[323,452,785,808]" firstAuthor="ICZN" pageId="17" pageNumber="292" refId="ref49784" refString="ICZN 1999: International code of zoological nomenclature. Fourth edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Article 24.2) and chose
<taxonomicName id="28D605B5FFCBFFE0FD29FCF7FCECFC98" authorityName="Boheman" authorityYear="1855" box="[725,850,785,808]" class="Insecta" family="Chrysomelidae" genus="Charidotis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="292" phylum="Arthropoda" rank="species" species="nucleata">
<emphasis id="DDA2A224FFCBFFE0FD29FCF7FCECFC98" box="[725,850,785,808]" italics="true" pageId="17" pageNumber="292">C. nucleata</emphasis>
</taxonomicName>
as the valid name for this species as it appears to be always correctly applied and historic material identified in museums agrees with the
<typeStatus id="306DC094FFCBFFE0FE79FCB7FE08FCD8" box="[389,438,849,872]" pageId="17" pageNumber="292">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A7CC2DBDFFCBFFE0FF8DFC97FF09FC18" pageId="17" pageNumber="292" type="distribution">
<paragraph id="EF697E36FFCBFFE0FF8DFC97FF09FC18" blockId="17.[113,1089,529,936]" pageId="17" pageNumber="292">
<emphasis id="DDA2A224FFCBFFE0FF8DFC97FEBAFC38" bold="true" box="[113,260,881,904]" pageId="17" pageNumber="292">Distribution.</emphasis>
<collectingCountry id="97C13EA6FFCBFFE0FEF7FC97FEEEFC38" box="[267,336,881,904]" name="Brazil" pageId="17" pageNumber="292">Brazil</collectingCountry>
: Pará (
<bibRefCitation id="8B4703C7FFCBFFE0FE66FC97FD9AFC38" author="SPAETH" box="[410,548,881,904]" firstAuthor="SPAETH" pageId="17" pageNumber="292" pagination="66 - 99" refId="ref51359" refString="SPAETH F. 1936: Bestimmungstabelle der Gattung Charidotis (Col. Chrys. Cassid.). Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze 14: 66 - 99." type="journal article" year="1936">SPAETH 1936</bibRefCitation>
,
<bibRefCitation id="8B4703C7FFCBFFE0FDD3FC97FD69FC38" author="BOROWIEC" box="[559,727,881,904]" firstAuthor="BOROWIEC" pageId="17" pageNumber="292" pagination="43 - 138" refId="ref48380" refString="BOROWIEC L. 2002: New records of Neotropical Cassidinae, with description of three new species (Coleoptera: Chrysomelidae). Genus 13: 43 - 138." type="journal article" year="2002">BOROWIEC 2002</bibRefCitation>
) and
<collectingRegion id="2D12B0D4FFCBFFE0FCEFFC97FC11FC38" box="[787,943,881,904]" country="Brazil" name="Rio de Janeiro" pageId="17" pageNumber="292">Rio de Janeiro</collectingRegion>
(
<bibRefCitation id="8B4703C7FFCBFFE0FC42FC97FF15FC18" author="FLINTE" etAl="et al." firstAuthor="FLINTE" pageId="17" pageNumber="292" pagination="571 - 614" refId="ref49297" refString="FLINTE V., BOROWIEC L., FERITAS S., VIANA J. H., FERNANDES F. R., NOGUEIRA-DE-SA F., MACEDO M. V. &amp; MONTEIRO R. F. 2009: Tortoise beetles of the State of Rio de Janeiro, Brazil (Coleoptera: Chrysomelidae: Cassidinae). Genus 20: 571 - 614." type="journal article" year="2009">FLINTE et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>