343 lines
46 KiB
XML
343 lines
46 KiB
XML
<document id="4D74DCDB1DEA27DFB43B8AE7B79830F7" ID-DOI="10.11646/zootaxa.5375.4.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10202772" ID-ZooBank="648DD26D-F271-4BD2-ADFB-0C5C31191FF1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1700813825895" checkinUser="plazi" docAuthor="Magnien, Philippe" docDate="2023" docId="294A879EFFDD8E5C37A8FA7925F4DBCB" docLanguage="en" docName="zootaxa.5375.4.5.pdf" docOrigin="Zootaxa 5375 (4)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.4.5/52344" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oukhypencha Magnien 2023, gen. nov." docType="treatment" docUuid="1C8FE412-DB94-461B-B9F3-D52CFAC1B860" docUuidSource="ZooBank" docVersion="6" lastPageNumber="536" masterDocId="D573FFE6FFDC8E5F373FFFFB2113DB32" masterDocTitle="New genera, species and subspecies of tessaratomine true bugs (Heteroptera: Tessaratomidae, Tessaratominae)" masterLastPageNumber="549" masterPageNumber="533" pageNumber="534" updateTime="1701864501225" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="BF6347931C6F62003C113038E562D4BC">New genera, species and subspecies of tessaratomine true bugs (Heteroptera: Tessaratomidae, Tessaratominae)</mods:title>
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<mods:namePart id="4EAA3B239B5985C96C81E2259A90781A">Magnien, Philippe</mods:namePart>
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<mods:title id="0C5E93F89C2B5CB33C576FB1FD0B98BE">Zootaxa</mods:title>
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<mods:part id="E0AD556BDE13784996197E58FC81E025">
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<mods:date id="A42D33AB4C771B74BE1483C53DECFBF0">2023</mods:date>
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<mods:number id="34409B1AFFB406A780D14090F3EB1554">2023-11-24</mods:number>
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<mods:number id="3B22C20B3E87DAC41D389ECF8A893E4D">5375</mods:number>
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<mods:url id="EDD143297C901B6DD1A91DB41E50AA66">https://www.mapress.com/zt/article/download/zootaxa.5375.4.5/52344</mods:url>
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<mods:classification id="227419AB942A34292D083E5C17ABCCBF">journal article</mods:classification>
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<mods:identifier id="2B90ABBEA219C0F00D3E04980F755458" type="DOI">10.11646/zootaxa.5375.4.5</mods:identifier>
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<mods:identifier id="FBDDB1CC6B723C3E7035320ADD00865F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="294A879EFFDD8E5C37A8FA7925F4DBCB" LSID="urn:lsid:zoobank.org:act:1C8FE412-DB94-461B-B9F3-D52CFAC1B860" httpUri="http://treatment.plazi.org/id/294A879EFFDD8E5C37A8FA7925F4DBCB" lastPageId="3" lastPageNumber="536" pageId="1" pageNumber="534">
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<subSubSection id="E9F96503FFDD8E5E37A8FA7920A5DEAF" box="[151,438,1410,1437]" pageId="1" pageNumber="534" type="nomenclature">
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<paragraph id="A15C3688FFDD8E5E37A8FA7920A5DEAF" blockId="1.[151,438,1410,1437]" box="[151,438,1410,1437]" pageId="1" pageNumber="534">
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<heading id="FA1481E4FFDD8E5E37A8FA7920A5DEAF" bold="true" box="[151,438,1410,1437]" fontSize="11" level="1" pageId="1" pageNumber="534" reason="1">
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<emphasis id="9397EA9AFFDD8E5E37A8FA7920A5DEAF" bold="true" box="[151,438,1410,1437]" pageId="1" pageNumber="534">
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<taxonomicName id="66E34D0BFFDD8E5E37A8FA792053DEAF" authority="Magnien, 2023" authorityName="Magnien" authorityYear="2023" box="[151,320,1410,1437]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="534" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="9397EA9AFFDD8E5E37A8FA792053DEAF" bold="true" box="[151,320,1410,1437]" italics="true" pageId="1" pageNumber="534">Oukhypencha</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="88A457E1FFDD8E5E3677FA7820A5DEAF" box="[328,438,1411,1437]" pageId="1" pageNumber="534" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9F96503FFDD8E5E37A8FA302243DED7" box="[151,848,1482,1509]" pageId="1" pageNumber="534" type="type_taxon">
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<paragraph id="A15C3688FFDD8E5E37A8FA302243DED7" blockId="1.[151,1436,1482,2013]" box="[151,848,1482,1509]" pageId="1" pageNumber="534">
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<emphasis id="9397EA9AFFDD8E5E37A8FA302021DED6" bold="true" box="[151,306,1482,1509]" pageId="1" pageNumber="534">
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<typeStatus id="7E58882AFFDD8E5E37A8FA3021C1DED7" box="[151,210,1483,1509]" pageId="1" pageNumber="534">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="66E34D0BFFDD8E5E3606FA3020CDDED7" authority="Magnien, 2023" authorityName="Magnien" authorityYear="2023" box="[313,478,1483,1509]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="534" phylum="Arthropoda" rank="species" species="clemenceae" status="sp. nov.">
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<emphasis id="9397EA9AFFDD8E5E3606FA3020CDDED7" box="[313,478,1483,1509]" italics="true" pageId="1" pageNumber="534">O. clemenceae</emphasis>
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</taxonomicName>
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<emphasis id="9397EA9AFFDD8E5E36DAFA30232CDED7" bold="true" box="[485,575,1483,1509]" pageId="1" pageNumber="534">
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<taxonomicNameLabel id="88A457E1FFDD8E5E36DAFA30232CDED7" box="[485,575,1483,1509]" pageId="1" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="E9F96503FFDD8E5E37F8FA1523A1DD47" pageId="1" pageNumber="534" type="diagnosis">
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<paragraph id="A15C3688FFDD8E5E37F8FA1523A1DD47" blockId="1.[151,1436,1482,2013]" pageId="1" pageNumber="534">
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<emphasis id="9397EA9AFFDD8E5E37F8FA152051DD3A" bold="true" box="[199,322,1518,1544]" pageId="1" pageNumber="534">Diagnosis.</emphasis>
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Medium sized
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<taxonomicName id="66E34D0BFFDD8E5E36CCFA142387DD3B" authorityName="Stal" authorityYear="1865" box="[499,660,1519,1545]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="534" phylum="Arthropoda" rank="subTribe" subTribe="Tessaratomina">Tessaratomina</taxonomicName>
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(
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<quantity id="661B9B6DFFDD8E5E359AFA142235DD3B" box="[677,806,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.6" metricValueMin="1.9" pageId="1" pageNumber="534" unit="mm" value="22.5" valueMax="26.0" valueMin="19.0">19–26 mm</quantity>
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), characterized by the following set of characters: head very small, with eyes and ocellae relatively large (ocular index 1.1–1.3, ocellar index 1.1–1.9), antennae long, little more than half (0.50–0.69) of habitus length; connexivum bicolourous; prosternal process lanceolate, rounded, sides not flattened, reaching or surpassing fore coxae.
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</paragraph>
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</subSubSection>
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<subSubSection id="E9F96503FFDD8E5D37F8F98525CFDAC7" lastPageId="2" lastPageNumber="535" pageId="1" pageNumber="534" type="description">
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<paragraph id="A15C3688FFDD8E5E37F8F9852063DDD3" blockId="1.[151,1436,1482,2013]" pageId="1" pageNumber="534">
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<emphasis id="9397EA9AFFDD8E5E37F8F985204ADDAA" bold="true" box="[199,345,1662,1688]" pageId="1" pageNumber="534">Description.</emphasis>
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Body of the common pentatomoid shape (
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<figureCitation id="39D82A0DFFDD8E5E3470F98422ADDDAB" box="[847,958,1663,1689]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–6" captionStart-2="FIGURES 7–9" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="6.[151,264,1914,1938]" captionStartId-2="9.[151,264,1943,1967]" captionTargetBox-0="[222,1365,179,1912]" captionTargetBox-1="[157,1431,315,1885]" captionTargetBox-2="[340,1247,185,1915]" captionTargetId-0="figure-23@4.[216,1371,179,1917]" captionTargetId-1="figure-88@6.[151,1435,309,1891]" captionTargetId-2="figure-15@9.[336,1251,181,1919]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="9" captionText-0="FIGURES 1–3: habitus.—1–2 Oukhypencha clemenceae n.sp.; 1:—♂ (paratype, Sabah)—2: ♀ (West Kalimantan)—3: Oukhypencha florenceae n. sp. (holotype ♂). Scale 10 mm" captionText-1="FIGURES 4–6: habitus.—4 Oukhypencha ophthalmica ophthalmica Stål ♀ (holotype); 5–6: Oukhypencha ophthalmica sumaterensis n. sp.—5: ♂ (holotype)—6: ♀ (paratype). Scale 10 mm" captionText-2="FIGURES 7–9: habitus.—7 Oukhypencha stali n. sp. ♀ (holotype); 8–9: Homoiacoris annemarieae n. sp.—8: ♂ (holotype)— 9: ♀ (paratype). Scale 10 mm" figureDoi-0="http://doi.org/10.5281/zenodo.10202774" figureDoi-1="http://doi.org/10.5281/zenodo.10202776" figureDoi-2="http://doi.org/10.5281/zenodo.10202780" httpUri-0="https://zenodo.org/record/10202774/files/figure.png" httpUri-1="https://zenodo.org/record/10202776/files/figure.png" httpUri-2="https://zenodo.org/record/10202780/files/figure.png" pageId="1" pageNumber="534">Figs. 1–7</figureCitation>
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), general hue bronze brownish above, of a lighter brownish ventrally; apex of fourth antennomere, apex of scutellum and forepart of each connexivum segment yellowish.
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</paragraph>
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<paragraph id="A15C3688FFDD8E5E37F8F9102552DC1B" blockId="1.[151,1436,1482,2013]" pageId="1" pageNumber="534">Head. Triangular, mandibular plates surpassing tylus, entirely punctate, save for a small area around ocellae; antennae 4-segmented, long; eyes and ocellae relatively large, ocular index 1.1–1.6.</paragraph>
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<paragraph id="A15C3688FFDD8E5E37F8F8C823BCDCEF" blockId="1.[151,1436,1482,2013]" pageId="1" pageNumber="534">
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Thorax. Pronotum sexangular, anterior, posterior and anterolateral margins nearly straight, posterolateral margins slightly bisinuous; callosities conspicuous, slightly marked. Scutellum triangular, longer than wide, its apex lanceolate. Metasternum with an elevated plate between coxae, bearing a rounded lanceolate process reaching or surpassing fore coxae, curved upwards at apex; evaporatoria small (
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<figureCitation id="39D82A0DFFDD8E5E348DF8642510DC8B" box="[946,1027,1951,1977]" captionStart="FIGURES 19–23" captionStartId="11.[151,264,1312,1336]" captionTargetBox="[155,1431,183,1289]" captionTargetId="figure-15@11.[151,1435,181,1289]" captionTargetPageId="11" captionText="FIGURES 19–23: Oukhypencha, female genitalia, evaporatoria.—19–22: female genitalia.—23: evaporatoria.—19, 23: Oukhypencha clemenceae n. sp.—20: Oukhypencha ophthalmica ophthalmica—21: Oukhypencha ophthalmica sumaterensis— 22: Oukhypencha stali. Abbreviations: ad—anterior duct, ev—evaporatoria field, ip—intermediate part, o—ostiole, pd—posterior duct, pw—posterior wall of vagina, r—receptacle, rs—ring sclerite, sd—striated dilation All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202786" httpUri="https://zenodo.org/record/10202786/files/figure.png" pageId="1" pageNumber="534">Fig. 23</figureCitation>
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), in accordance with the description given for the tribe in Kment &Vilimova (2010).
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</paragraph>
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<paragraph id="A15C3688FFDE8E5D37F8FF6C2279DBE7" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">Abdomen. Connexivum and urosternite margins straight; spiracles closer to margin than to middle of the sternites, trichobothria closer to middle, aft of transverse furrow.</paragraph>
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<paragraph id="A15C3688FFDE8E5D37F8FF2421FEDA57" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">Male genitalia (Figs. 11–13, 15–18). Pygophore of various shapes. Parameres having shape of a recurved blade, rounded at apophysis, sensorial lobe conspicuous, marked by a brush of setae about as long as parameral width; phallus bearing three or four processus, two of them strongly sclerotized, lateroventral process strong, scythe blade-shaped.</paragraph>
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<paragraph id="A15C3688FFDE8E5D37F8FE9425CFDAC7" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">
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Female genitalia (
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<figureCitation id="39D82A0DFFDE8E5D36A2FE94233EDABB" box="[413,557,367,393]" captionStart="FIGURES 19–23" captionStartId="11.[151,264,1312,1336]" captionTargetBox="[155,1431,183,1289]" captionTargetId="figure-15@11.[151,1435,181,1289]" captionTargetPageId="11" captionText="FIGURES 19–23: Oukhypencha, female genitalia, evaporatoria.—19–22: female genitalia.—23: evaporatoria.—19, 23: Oukhypencha clemenceae n. sp.—20: Oukhypencha ophthalmica ophthalmica—21: Oukhypencha ophthalmica sumaterensis— 22: Oukhypencha stali. Abbreviations: ad—anterior duct, ev—evaporatoria field, ip—intermediate part, o—ostiole, pd—posterior duct, pw—posterior wall of vagina, r—receptacle, rs—ring sclerite, sd—striated dilation All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202786" httpUri="https://zenodo.org/record/10202786/files/figure.png" pageId="2" pageNumber="535">Figs. 19–22</figureCitation>
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). Genital plates of general shape found in
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<taxonomicName id="66E34D0BFFDE8E5D3309FE9425C5DABB" authorityName="Stal" authorityYear="1865" box="[1078,1238,367,393]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="tribe" tribe="Tessaratomini">Tessaratomini</taxonomicName>
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. Spermatheca of common
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<typeStatus id="7E58882AFFDE8E5D363BFE682027DA9F" box="[260,308,403,429]" pageId="2" pageNumber="535">type</typeStatus>
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in
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<taxonomicName id="66E34D0BFFDE8E5D365FFE682313DA9F" authorityName="Stal" authorityYear="1865" box="[352,512,403,429]" class="Insecta" family="Tessaratomidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="tribe" tribe="Tessaratomini">Tessaratomini</taxonomicName>
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; apical receptacle (distal bulb) spherical with a basal “neck”, intermediate part (pump apparatus) with two well-developed flanges; spermathecal duct bipartite, anterior duct slender, posterior duct much wider, its anterior part slightly ovoid and longitudinally striated, rear part somewhat longer.
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</paragraph>
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</subSubSection>
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<subSubSection id="E9F96503FFDE8E5D37F8FE042581D90F" pageId="2" pageNumber="535" type="etymology">
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<paragraph id="A15C3688FFDE8E5D37F8FE042581D90F" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">
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<emphasis id="9397EA9AFFDE8E5D37F8FE04205BD92B" bold="true" box="[199,328,511,537]" pageId="2" pageNumber="535">Etymology</emphasis>
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. The name
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<taxonomicName id="66E34D0BFFDE8E5D36EBFE042355D92B" authorityName="Amyot & Serville" authorityYear="1843" box="[468,582,511,537]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
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<emphasis id="9397EA9AFFDE8E5D36EBFE042355D92B" box="[468,582,511,537]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
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</taxonomicName>
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was crafted by its authors from the Greek ύπό, under and ἕγχος, sword. Ι added the prefix
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<emphasis id="9397EA9AFFDE8E5D366BFDD82090D90F" box="[340,387,547,573]" italics="true" pageId="2" pageNumber="535">Ouk</emphasis>
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, from the Greek οὑκ, not, to imply it was not belonging in
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<taxonomicName id="66E34D0BFFDE8E5D3326FDD82598D90F" authorityName="Amyot & Serville" authorityYear="1843" box="[1049,1163,547,573]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
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<emphasis id="9397EA9AFFDE8E5D3326FDD82598D90F" box="[1049,1163,547,573]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="E9F96503FFDE8E5D37F8FDBD2595D953" box="[199,1158,582,609]" pageId="2" pageNumber="535" type="distribution">
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||
<paragraph id="A15C3688FFDE8E5D37F8FDBD2595D953" blockId="2.[151,1437,151,2043]" box="[199,1158,582,609]" pageId="2" pageNumber="535">
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<emphasis id="9397EA9AFFDE8E5D37F8FDBD204AD952" bold="true" box="[199,345,582,608]" pageId="2" pageNumber="535">Distribution</emphasis>
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. Peninsular
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<collectingCountry id="D9F47618FFDE8E5D36DBFDBC2359D953" box="[484,586,583,609]" name="Thailand" pageId="2" pageNumber="535">Thailand</collectingCountry>
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,
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<collectingCountry id="D9F47618FFDE8E5D3569FDBC23ACD953" box="[598,703,583,609]" name="Malaysia" pageId="2" pageNumber="535">Malaysia</collectingCountry>
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,
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<collectingCountry id="D9F47618FFDE8E5D35F5FDBC2224D953" box="[714,823,583,609]" name="Indonesia" pageId="2" pageNumber="535">Indonesia</collectingCountry>
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(
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<collectingRegion id="6327F86AFFDE8E5D3479FDBC226FD953" box="[838,892,583,609]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="535">Java</collectingRegion>
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,
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<collectingRegion id="6327F86AFFDE8E5D34B8FDBC22F4D953" box="[903,999,583,609]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="535">Sumatra</collectingRegion>
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,
|
||
<collectingRegion id="6327F86AFFDE8E5D34CCFDBC2568D953" box="[1011,1147,583,609]" country="Indonesia" name="Kalimantan Utara" pageId="2" pageNumber="535">Kalimantan</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9F96503FFDE8E5C37F8FD9125F4DBCB" lastPageId="3" lastPageNumber="536" pageId="2" pageNumber="535" type="discussion">
|
||
<paragraph id="A15C3688FFDE8E5D37F8FD912573D80B" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">
|
||
<emphasis id="9397EA9AFFDE8E5D37F8FD912056D9B6" bold="true" box="[199,325,618,644]" pageId="2" pageNumber="535">Discussion</emphasis>
|
||
. During this study, it became very apparent that previously identified material of
|
||
<taxonomicName id="66E34D0BFFDE8E5D33D3FD90248FD9B7" authorityName="Stal" authorityYear="1863" box="[1260,1436,619,645]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="species" species="ophthalmica">
|
||
<emphasis id="9397EA9AFFDE8E5D33D3FD90248FD9B7" box="[1260,1436,619,645]" italics="true" pageId="2" pageNumber="535">H. ophthalmica</emphasis>
|
||
</taxonomicName>
|
||
was in fact a composite of at least four different species. The difficulties were a result of problems in the proper identification of the male of Stål’s species, and in the true generic placement of these taxa. Regarding the first problem, unfortunately, no specimens from Peninsular
|
||
<collectingCountry id="D9F47618FFDE8E5D35CBFD2C224BD9C3" box="[756,856,727,753]" name="Thailand" pageId="2" pageNumber="535">Thailand</collectingCountry>
|
||
or
|
||
<collectingCountry id="D9F47618FFDE8E5D3445FD2C22F2D9C3" box="[890,993,727,753]" name="Malaysia" pageId="2" pageNumber="535">Malaysia</collectingCountry>
|
||
were available for study, so the decision was made to select the taxon with the most similar female genitalia as representing
|
||
<taxonomicName id="66E34D0BFFDE8E5D3301FD0025FFD827" authorityName="Stal" authorityYear="1863" box="[1086,1260,763,789]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="species" species="ophthalmica">
|
||
<emphasis id="9397EA9AFFDE8E5D3301FD0025FFD827" box="[1086,1260,763,789]" italics="true" pageId="2" pageNumber="535">H. ophthalmica</emphasis>
|
||
</taxonomicName>
|
||
. It is one of the species from Sumatra, which is also among the closest to the
|
||
<typeStatus id="7E58882AFFDE8E5D3478FCE42264D80B" box="[839,887,799,825]" pageId="2" pageNumber="535">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="D9F47618FFDE8E5D34CAFCE4254FD80B" box="[1013,1116,799,825]" name="Thailand" pageId="2" pageNumber="535">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A15C3688FFDE8E5D37F8FCB82527D8FB" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">
|
||
Regarding the generic placement, it was quickly determined that for the involved taxa for which male specimens were available for study, none of them could fit in
|
||
<taxonomicName id="66E34D0BFFDE8E5D3589FC9C223BD8B3" authorityName="Amyot & Serville" authorityYear="1843" box="[694,808,871,897]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3589FC9C223BD8B3" box="[694,808,871,897]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
. In fact, the variation in the male genitalia observed could have justified the creation of several different genera. Despite this variability, I have chosen to take a conservative approach and place all of these taxa in a single new genus,
|
||
<taxonomicName id="66E34D0BFFDE8E5D3412FC5422D4D8FB" authority="Magnien, 2023" authorityName="Magnien" authorityYear="2023" box="[813,967,943,969]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="9397EA9AFFDE8E5D3412FC5422D4D8FB" box="[813,967,943,969]" italics="true" pageId="2" pageNumber="535">Oukhypencha</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="88A457E1FFDE8E5D34F1FC542527D8FB" box="[974,1076,943,969]" pageId="2" pageNumber="535" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="A15C3688FFDE8E5D37F8FC282273DFDB" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">
|
||
Its habitus is similar to that of other tessaratomine genera, the closest being
|
||
<taxonomicName id="66E34D0BFFDE8E5D3327FC282599D8DF" authorityName="Amyot & Serville" authorityYear="1843" box="[1048,1162,979,1005]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3327FC282599D8DF" box="[1048,1162,979,1005]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
, the genus in which Stål originally placed
|
||
<taxonomicName id="66E34D0BFFDE8E5D3666FC0C2315DF23" authorityName="Stal" authorityYear="1863" box="[345,518,1015,1041]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="species" species="ophthalmica">
|
||
<emphasis id="9397EA9AFFDE8E5D3666FC0C2315DF23" box="[345,518,1015,1041]" italics="true" pageId="2" pageNumber="535">H. ophthalmica</emphasis>
|
||
</taxonomicName>
|
||
. In general, the male genitalia are indeed very good characters to use in delineating tessaratomine genera. That is, the male genitalia are usually very similar among species within each genus, but they usually vary distinctly between genera (except for a couple, e.g.,
|
||
<taxonomicName id="66E34D0BFFDE8E5D34BFFBC42541DF6B" authority="Stal" authorityName="Stal" box="[896,1106,1086,1113]" class="Insecta" family="Tessaratomidae" genus="Embolosterna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D34BFFBC4250EDF6B" box="[896,1053,1087,1113]" italics="true" pageId="2" pageNumber="535">Embolosterna</emphasis>
|
||
Stål
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="66E34D0BFFDE8E5D33B4FBC42497DF6B" authority="Berthold" authorityName="Berthold" authorityYear="1827" box="[1163,1412,1087,1113]" class="Insecta" family="Tessaratomidae" genus="Tessaratoma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D33B4FBC4240ADF6A" box="[1163,1305,1087,1112]" italics="true" pageId="2" pageNumber="535">Tessaratoma</emphasis>
|
||
Berthold
|
||
</taxonomicName>
|
||
<superScript id="56969BC0FFDE8E5D32BBFBC5249FDF7E" attach="left" box="[1412,1420,1086,1100]" fontSize="6" pageId="2" pageNumber="535">1</superScript>
|
||
), to the point that it is usually fairly easy to identify a genus from the examination of the male genitalia alone (see
|
||
<bibRefCitation id="C5724B79FFDE8E5D37A8FB7C208EDF93" author="Kumar R. & Ghauri M. S. K." box="[151,413,1159,1185]" pageId="2" pageNumber="535" pagination="1 - 32" refId="ref10342" refString="Kumar R. & Ghauri M. S. K. (1970) Morphology and relationships of the Pentatomoidea (Heteroptera) 2 - World genera of Tessaratomini (Tessaratomidae), Deutsche Entomologische Zeitschrift, 17 (I - III), 1 - 32. https: // doi. org / 10.1002 / mmnd. 19700170102" type="journal article" year="1970">Kumar & Ghauri, 1970</bibRefCitation>
|
||
). The phallus of
|
||
<taxonomicName id="66E34D0BFFDE8E5D356BFB7C23D5DF93" authorityName="Amyot & Serville" authorityYear="1843" box="[596,710,1159,1185]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D356BFB7C23D5DF93" box="[596,710,1159,1185]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
, according to
|
||
<bibRefCitation id="C5724B79FFDE8E5D345CFB7C2596DF93" author="Kumar R. & Ghauri M. S. K." box="[867,1157,1159,1185]" pageId="2" pageNumber="535" pagination="1 - 32" refId="ref10342" refString="Kumar R. & Ghauri M. S. K. (1970) Morphology and relationships of the Pentatomoidea (Heteroptera) 2 - World genera of Tessaratomini (Tessaratomidae), Deutsche Entomologische Zeitschrift, 17 (I - III), 1 - 32. https: // doi. org / 10.1002 / mmnd. 19700170102" type="journal article" year="1970">Kumar and Ghauri (1970)</bibRefCitation>
|
||
, can be characterized by:
|
||
<emphasis id="9397EA9AFFDE8E5D37A8FB50200BDFF7" box="[151,280,1195,1221]" italics="true" pageId="2" pageNumber="535">
|
||
“
|
||
<taxonomicName id="66E34D0BFFDE8E5D379EFB50200BDFF7" authorityName="Amyot & Serville" authorityYear="1843" box="[161,280,1195,1221]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">Hypencha</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="66E34D0BFFDE8E5D3618FB502084DFF6" authorityName="Amyot & Serville" authorityYear="1843" box="[295,407,1195,1220]" class="Insecta" family="Tessaratomidae" genus="Mucanum" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3618FB502084DFF6" box="[295,407,1195,1220]" italics="true" pageId="2" pageNumber="535">Mucanum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="66E34D0BFFDE8E5D369AFB502351DFF7" box="[421,578,1195,1221]" class="Insecta" family="Tessaratomidae" genus="Embolosterna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D369AFB502351DFF7" box="[421,578,1195,1221]" italics="true" pageId="2" pageNumber="535">Embolosterna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="66E34D0BFFDE8E5D3545FB50221BDFF6" authorityName="Berthold" authorityYear="1827" box="[634,776,1195,1220]" class="Insecta" family="Tessaratomidae" genus="Tessaratoma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3545FB50221BDFF6" box="[634,776,1195,1220]" italics="true" pageId="2" pageNumber="535">Tessaratoma</emphasis>
|
||
</taxonomicName>
|
||
, all have three pairs of conjunctival processes.”
|
||
<taxonomicName id="66E34D0BFFDE8E5D3214FB50248EDFF7" authorityName="Amyot & Serville" authorityYear="1843" box="[1323,1437,1195,1221]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3214FB50248EDFF7" box="[1323,1437,1195,1221]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
is distinct in having two pairs of ventral conjunctival processes.
|
||
</paragraph>
|
||
<paragraph id="A15C3688FFDE8E5D37F8FB0822FFDE03" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">They add that the anteroventral one is membranous, and that the paramere is elongate, apically curved. Those descriptions do not match the organisation of the genitalia in the new genus.</paragraph>
|
||
<paragraph id="A15C3688FFDE8E5D37F8FAC020D6DDD3" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">
|
||
The conjunctiva of
|
||
<taxonomicName id="66E34D0BFFDE8E5D3697FAC02351DE67" authorityName="Magnien" authorityYear="2023" box="[424,578,1339,1365]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3697FAC02351DE67" box="[424,578,1339,1365]" italics="true" pageId="2" pageNumber="535">Oukhypencha</emphasis>
|
||
</taxonomicName>
|
||
bears three or four pairs of processes, but even in those species with three pairs, their disposition is not homologous with those of
|
||
<taxonomicName id="66E34D0BFFDE8E5D343AFAA42264DE4B" authorityName="Amyot & Serville" authorityYear="1843" box="[773,887,1375,1401]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D343AFAA42264DE4B" box="[773,887,1375,1401]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="66E34D0BFFDE8E5D34BBFAA422E5DE4B" authorityName="Amyot & Serville" authorityYear="1843" box="[900,1014,1375,1401]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D34BBFAA422E5DE4B" box="[900,1014,1375,1401]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="39D82A0DFFDE8E5D333AFAA42545DE4B" box="[1029,1110,1374,1401]" captionStart="FIGURES 15–18" captionStartId="10.[151,264,1797,1821]" captionTargetBox="[155,1430,348,1770]" captionTargetId="figure-115@10.[151,1435,345,1774]" captionTargetPageId="10" captionText="FIGURES 15–18: male genitalia, phallus and paramere—top: phallus (lateral view)—middle: ventral processus (ventral view, right side)—bottom: paramere (dorsal and lateral view, pilosity omitted in lateral view)—15 Hypencha sp. (same specimen as Fig. 10)—16: Oukhypencha clemenceae n. sp.—17: Oukhypencha florenceae n. sp.—18: Oukhypencha ophthalmica sumaterensis n. subsp. Abbreviations: a—apophysis, c—conjunctiva, d—membranous dorsal process, lv—latero-ventral processus, mv—membranous ventral process, ph—phallobase, sd—sclerified dorsal processus, sl—sensorial lobe, v—vesica, vp—ventral processus. All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202784" httpUri="https://zenodo.org/record/10202784/files/figure.png" pageId="2" pageNumber="535">Fig. 15</figureCitation>
|
||
) lacks the ventral sclerotized and membranous processes, while
|
||
<taxonomicName id="66E34D0BFFDE8E5D3523FA7823A5DEAF" authorityName="Magnien" authorityYear="2023" box="[540,694,1411,1437]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3523FA7823A5DEAF" box="[540,694,1411,1437]" italics="true" pageId="2" pageNumber="535">Oukhypencha</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="39D82A0DFFDE8E5D35FAFA782258DEAF" box="[709,843,1411,1437]" captionStart="FIGURES 15–18" captionStartId="10.[151,264,1797,1821]" captionTargetBox="[155,1430,348,1770]" captionTargetId="figure-115@10.[151,1435,345,1774]" captionTargetPageId="10" captionText="FIGURES 15–18: male genitalia, phallus and paramere—top: phallus (lateral view)—middle: ventral processus (ventral view, right side)—bottom: paramere (dorsal and lateral view, pilosity omitted in lateral view)—15 Hypencha sp. (same specimen as Fig. 10)—16: Oukhypencha clemenceae n. sp.—17: Oukhypencha florenceae n. sp.—18: Oukhypencha ophthalmica sumaterensis n. subsp. Abbreviations: a—apophysis, c—conjunctiva, d—membranous dorsal process, lv—latero-ventral processus, mv—membranous ventral process, ph—phallobase, sd—sclerified dorsal processus, sl—sensorial lobe, v—vesica, vp—ventral processus. All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202784" httpUri="https://zenodo.org/record/10202784/files/figure.png" pageId="2" pageNumber="535">Figs. 16–18</figureCitation>
|
||
) species lack the proximoventral process. The dorsal process is sclerotized in
|
||
<taxonomicName id="66E34D0BFFDE8E5D3691FA5C2333DEF3" authorityName="Amyot & Serville" authorityYear="1843" box="[430,544,1447,1473]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3691FA5C2333DEF3" box="[430,544,1447,1473]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
, while it is membranous in the new genus, except for
|
||
<taxonomicName id="66E34D0BFFDE8E5D33B5FA5C2431DEF3" authorityName="Magnien" authorityYear="2023" box="[1162,1314,1447,1473]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="species" species="florenceae">
|
||
<emphasis id="9397EA9AFFDE8E5D33B5FA5C2431DEF3" box="[1162,1314,1447,1473]" italics="true" pageId="2" pageNumber="535">O. florenceae</emphasis>
|
||
</taxonomicName>
|
||
which has two dorsal processes, the distal process is membranous, while the proximal process is sclerotized. Furthermore, the parameres (
|
||
<figureCitation id="39D82A0DFFDE8E5D367AFA1420DEDD3B" box="[325,461,1519,1545]" captionStart="FIGURES 15–18" captionStartId="10.[151,264,1797,1821]" captionTargetBox="[155,1430,348,1770]" captionTargetId="figure-115@10.[151,1435,345,1774]" captionTargetPageId="10" captionText="FIGURES 15–18: male genitalia, phallus and paramere—top: phallus (lateral view)—middle: ventral processus (ventral view, right side)—bottom: paramere (dorsal and lateral view, pilosity omitted in lateral view)—15 Hypencha sp. (same specimen as Fig. 10)—16: Oukhypencha clemenceae n. sp.—17: Oukhypencha florenceae n. sp.—18: Oukhypencha ophthalmica sumaterensis n. subsp. Abbreviations: a—apophysis, c—conjunctiva, d—membranous dorsal process, lv—latero-ventral processus, mv—membranous ventral process, ph—phallobase, sd—sclerified dorsal processus, sl—sensorial lobe, v—vesica, vp—ventral processus. All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202784" httpUri="https://zenodo.org/record/10202784/files/figure.png" pageId="2" pageNumber="535">Figs. 16–18</figureCitation>
|
||
) are not digitiform as those found in species of
|
||
<taxonomicName id="66E34D0BFFDE8E5D34CDFA142577DD3B" authorityName="Amyot & Serville" authorityYear="1843" box="[1010,1124,1519,1545]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D34CDFA142577DD3B" box="[1010,1124,1519,1545]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="39D82A0DFFDE8E5D334BFA1425DBDD3B" box="[1140,1224,1518,1545]" captionStart="FIGURES 15–18" captionStartId="10.[151,264,1797,1821]" captionTargetBox="[155,1430,348,1770]" captionTargetId="figure-115@10.[151,1435,345,1774]" captionTargetPageId="10" captionText="FIGURES 15–18: male genitalia, phallus and paramere—top: phallus (lateral view)—middle: ventral processus (ventral view, right side)—bottom: paramere (dorsal and lateral view, pilosity omitted in lateral view)—15 Hypencha sp. (same specimen as Fig. 10)—16: Oukhypencha clemenceae n. sp.—17: Oukhypencha florenceae n. sp.—18: Oukhypencha ophthalmica sumaterensis n. subsp. Abbreviations: a—apophysis, c—conjunctiva, d—membranous dorsal process, lv—latero-ventral processus, mv—membranous ventral process, ph—phallobase, sd—sclerified dorsal processus, sl—sensorial lobe, v—vesica, vp—ventral processus. All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10202784" httpUri="https://zenodo.org/record/10202784/files/figure.png" pageId="2" pageNumber="535">Fig. 15</figureCitation>
|
||
), and they possess a conspicuous sensorial lobe with a dense brush of setae, which is absent in all examined specimens of
|
||
<taxonomicName id="66E34D0BFFDE8E5D321BF9E82485DD1F" authorityName="Amyot & Serville" authorityYear="1843" box="[1316,1430,1555,1581]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D321BF9E82485DD1F" box="[1316,1430,1555,1581]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
. Some differences, probably of less generic significance, listed in the diagnosis, can be found in the habitus. The antennae are relatively long for a tessaratomine, from about 50 to 70 % of the habitus length, whereas in
|
||
<taxonomicName id="66E34D0BFFDE8E5D3214F9A0248EDD47" authorityName="Amyot & Serville" authorityYear="1843" box="[1323,1437,1627,1653]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3214F9A0248EDD47" box="[1323,1437,1627,1653]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
specimens, the antennae are about one-third of the habitus length, except for
|
||
<taxonomicName id="66E34D0BFFDE8E5D34C5F984259CDDAB" authorityName="Bergroth" authorityYear="1922" box="[1018,1167,1663,1689]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="species" species="aeroplana">
|
||
<emphasis id="9397EA9AFFDE8E5D34C5F984259CDDAB" box="[1018,1167,1663,1689]" italics="true" pageId="2" pageNumber="535">H. aeroplana</emphasis>
|
||
</taxonomicName>
|
||
in which it reaches less than half the habitus length. The ocellae are very large. The free part of the sternal process is rounded, not flattened, as is the case in
|
||
<taxonomicName id="66E34D0BFFDE8E5D3673F93C20ADDDD3" authorityName="Amyot & Serville" authorityYear="1843" box="[332,446,1735,1761]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3673F93C20ADDDD3" box="[332,446,1735,1761]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A15C3688FFDE8E5D37F8F9102219DC43" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">
|
||
The connexivum is bicolorous in
|
||
<taxonomicName id="66E34D0BFFDE8E5D3579F91023F3DC37" authorityName="Magnien" authorityYear="2023" box="[582,736,1771,1797]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3579F91023F3DC37" box="[582,736,1771,1797]" italics="true" pageId="2" pageNumber="535">Oukhypencha</emphasis>
|
||
</taxonomicName>
|
||
, contrasting with unicolorous in
|
||
<taxonomicName id="66E34D0BFFDE8E5D3365F910243FDC37" box="[1114,1324,1771,1797]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="9397EA9AFFDE8E5D3365F91025DFDC37" box="[1114,1228,1771,1797]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
The apex of the scutellum is large and concolorous in
|
||
<taxonomicName id="66E34D0BFFDE8E5D35B9F8F423EBDC1B" authorityName="Amyot & Serville" authorityYear="1843" box="[646,760,1807,1833]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D35B9F8F423EBDC1B" box="[646,760,1807,1833]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
, while it is thin and more or less yellowish in
|
||
<taxonomicName id="66E34D0BFFDE8E5D33C4F8F42486DC1B" authorityName="Magnien" authorityYear="2023" box="[1275,1429,1807,1833]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D33C4F8F42486DC1B" box="[1275,1429,1807,1833]" italics="true" pageId="2" pageNumber="535">Oukhypencha</emphasis>
|
||
</taxonomicName>
|
||
. The size of the eyes and ocellae is larger for
|
||
<taxonomicName id="66E34D0BFFDE8E5D35A3F8C82225DC7F" authorityName="Magnien" authorityYear="2023" box="[668,822,1843,1869]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D35A3F8C82225DC7F" box="[668,822,1843,1869]" italics="true" pageId="2" pageNumber="535">Oukhypencha</emphasis>
|
||
</taxonomicName>
|
||
(ocular index 1.1–1.6, ocellar index 1.1–3.5) than for
|
||
<taxonomicName id="66E34D0BFFDE8E5D37A8F8AC201ADC43" authorityName="Amyot & Serville" authorityYear="1843" box="[151,265,1879,1905]" class="Insecta" family="Tessaratomidae" genus="Hypencha" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D37A8F8AC201ADC43" box="[151,265,1879,1905]" italics="true" pageId="2" pageNumber="535">Hypencha</emphasis>
|
||
</taxonomicName>
|
||
(ocular index 1.3–1.6, ocellar index 2.0–5.5).
|
||
</paragraph>
|
||
<paragraph id="A15C3688FFDE8E5D37A8F860249EDCC8" blockId="2.[151,1437,151,2043]" pageId="2" pageNumber="535">
|
||
1 The synonymy between these two taxa should be examined on this basis; no character, except for the shape and colouration of the post-frenal part of the scutellum, has been found that can reliably separate the two genera. The conclusion of
|
||
<bibRefCitation id="C5724B79FFDE8E5D3214F8442034DCC9" author="Kumar R. & Ghauri M. S. K." pageId="2" pageNumber="535" pagination="1 - 32" refId="ref10342" refString="Kumar R. & Ghauri M. S. K. (1970) Morphology and relationships of the Pentatomoidea (Heteroptera) 2 - World genera of Tessaratomini (Tessaratomidae), Deutsche Entomologische Zeitschrift, 17 (I - III), 1 - 32. https: // doi. org / 10.1002 / mmnd. 19700170102" type="journal article" year="1970">Kumar and Ghauri (1970)</bibRefCitation>
|
||
on this point is not sustainable, as their key would place
|
||
<taxonomicName id="66E34D0BFFDE8E5D3455F81F255CDCC9" authority="Martin, 1902" authorityName="Martin" authorityYear="1902" box="[874,1103,2019,2043]" class="Insecta" family="Tessaratomidae" genus="Embolosterna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="species" species="vacca">
|
||
<emphasis id="9397EA9AFFDE8E5D3455F81F226ADCC9" box="[874,889,2020,2043]" italics="true" pageId="2" pageNumber="535">E</emphasis>
|
||
.
|
||
<emphasis id="9397EA9AFFDE8E5D34B8F81F22D1DCC9" box="[903,962,2020,2043]" italics="true" pageId="2" pageNumber="535">vacca</emphasis>
|
||
Martin, 1902
|
||
</taxonomicName>
|
||
, within the genus
|
||
<taxonomicName id="66E34D0BFFDE8E5D3238F81F249BDCC9" authorityName="Berthold" authorityYear="1827" box="[1287,1416,2020,2043]" class="Insecta" family="Tessaratomidae" genus="Tessaratoma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9397EA9AFFDE8E5D3238F81F249BDCC9" box="[1287,1416,2020,2043]" italics="true" pageId="2" pageNumber="535">Tessaratoma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A15C3688FFDF8E5C37F8FF6C25F4DBCB" blockId="3.[151,1436,151,249]" pageId="3" pageNumber="536">
|
||
The choice to designate
|
||
<taxonomicName id="66E34D0BFFDF8E5C36E8FF6C236FDB83" authority="Magnien, 2023" authorityName="Magnien" authorityYear="2023" box="[471,636,151,177]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="536" phylum="Arthropoda" rank="species" species="clemenceae" status="sp. nov.">
|
||
<emphasis id="9397EA9AFFDF8E5C36E8FF6C236FDB83" box="[471,636,151,177]" italics="true" pageId="3" pageNumber="536">O. clemenceae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9397EA9AFFDF8E5C35BDFF6C23D3DB83" bold="true" box="[642,704,151,177]" pageId="3" pageNumber="536">
|
||
<taxonomicNameLabel id="88A457E1FFDF8E5C35BDFF6C23D3DB83" box="[642,704,151,177]" pageId="3" pageNumber="536" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
as the
|
||
<typeStatus id="7E58882AFFDF8E5C3430FF6C222CDB83" box="[783,831,151,177]" pageId="3" pageNumber="536">type</typeStatus>
|
||
species for this new genus was made on the basis that both male and female specimens where readily available in good numbers, which is not the case with the previously described
|
||
<taxonomicName id="66E34D0BFFDF8E5C3636FF2420ABDBCB" authorityName="Stal" authorityYear="1863" box="[265,440,223,249]" class="Insecta" family="Tessaratomidae" genus="Oukhypencha" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="536" phylum="Arthropoda" rank="species" species="ophthalmica">
|
||
<emphasis id="9397EA9AFFDF8E5C3636FF2420ABDBCB" box="[265,440,223,249]" italics="true" pageId="3" pageNumber="536">O. ophthalmica</emphasis>
|
||
</taxonomicName>
|
||
for which male specimens are still unknown for the nominal subspecies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |