treatments-xml/data/03/E4/BA/03E4BA27FF99FFBDFDC6B7CBFBC0FA57.xml
2024-06-21 12:22:17 +02:00

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<mods:title id="9884A3C338B9EE8B96423A6324F21C5E">Maratus banyowla, a new peacock spider in the personatus group from Western Australia (Araneae: Salticidae: Euophryini)</mods:title>
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<mods:namePart id="2DC903C1A948EE17D8680146A41ED051">Otto, Jürgen C.</mods:namePart>
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<mods:namePart id="4F955F1CD42146FF3737C12A44F08730">Hill, David E.</mods:namePart>
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<paragraph id="8BF20B31FF99FFA8FDC6B7CBFBE2FF0B" blockId="1.[587,1045,172,202]" box="[587,1045,172,202]" pageId="1" pageNumber="2">
<heading id="D0BABC5DFF99FFA8FDC6B7CBFBE2FF0B" bold="true" box="[587,1045,172,202]" centered="true" fontSize="12" level="2" pageId="1" pageNumber="2" reason="0">
<emphasis id="B939D723FF99FFA8FDC6B7CBFBE2FF0B" bold="true" box="[587,1045,172,202]" pageId="1" pageNumber="2">
<taxonomicName id="4C4D70B2FF99FFA8FDC6B7CBFCA1FF0B" ID-CoL="95G7Z" authority="Otto &amp; Hill, 2019" authorityName="Otto &amp; Hill" authorityYear="2019" box="[587,854,172,202]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="banyowla" status="sp. nov.">
<emphasis id="B939D723FF99FFA8FDC6B7CBFCA1FF0B" bold="true" box="[587,854,172,202]" italics="true" pageId="1" pageNumber="2">Maratus banyowla</emphasis>
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,
<taxonomicNameLabel id="A20A6A58FF99FFA8FCEBB7C9FBE2FF0B" box="[870,1045,173,202]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C35758BAFF99FFA8FFEDB790FB8FFE47" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BF20B31FF99FFA8FFEDB790FB8FFE47" blockId="1.[96,1534,247,390]" pageId="1" pageNumber="2">
<emphasis id="B939D723FF99FFA8FFEDB790FEC7FED4" box="[96,304,247,277]" italics="true" pageId="1" pageNumber="2">Type specimens</emphasis>
. The
<materialsCitation id="3B25016CFF99FFA8FE0AB79EFB83FE47" ID-GBIF-Occurrence="2592359492" collectingDate="2019-09-11" country="Australia" latitude="-32.06286" location="Ellis Brook Valley, Banyowla Regional Park, Rushton Road, Martin, Perth" longLatPrecision="1" longitude="116.03307" municipality="David Knowles and Kerry Stuart" pageId="1" pageNumber="2" specimenCode="#4" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Western Australia" typeStatus="holotype">
<typeStatus id="54F6B593FF99FFA8FE0AB79EFDF7FED7" box="[391,512,249,278]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
<specimenCount id="9D4BC0B8FF99FFA8FD85B79EFDBCFED7" box="[520,587,249,278]" pageId="1" pageNumber="2" type="male">male</specimenCount>
(
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<specimenCode id="DBEBA34AFF99FFA8FD0DB79EFD5EFED7" box="[640,681,249,278]" pageId="1" pageNumber="2">#4</specimenCode>
), six
<typeStatus id="54F6B593FF99FFA8FD7DB79EFC9CFED4" box="[752,875,249,277]" pageId="1" pageNumber="2" type="paratype">paratype</typeStatus>
<specimenCount id="9D4BC0B8FF99FFA8FCFEB79EFC33FED7" box="[883,964,249,278]" pageId="1" pageNumber="2" type="male">males</specimenCount>
(
<specimenCount id="9D4BC0B8FF99FFA8FC54B79EFC06FED4" box="[985,1009,249,277]" pageId="1" pageNumber="2" type="male"></specimenCount>
#1-3, 5-7), and four
<typeStatus id="54F6B593FF99FFA8FA9FB79EFA7AFED4" box="[1298,1421,249,277]" pageId="1" pageNumber="2" type="paratype">paratype</typeStatus>
<specimenCount id="9D4BC0B8FF99FFA8FA18B79EFA09FED7" box="[1429,1534,249,278]" pageId="1" pageNumber="2" type="female">females</specimenCount>
(
<specimenCount id="9D4BC0B8FF99FFA8FFE0B678FF72FEFA" box="[109,133,287,315]" pageId="1" pageNumber="2" type="female"></specimenCount>
#1-4) were collected at
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,
<collectingRegion id="4989C5D3FF99FFA8FFEDB623FEAAFEA1" box="[96,349,323,352]" country="Australia" name="Western Australia" pageId="1" pageNumber="2">Western Australia</collectingRegion>
(~
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,
<geoCoordinate id="EE796DF6FF99FFA8FDD4B623FCD9FEA1" box="[601,814,323,352]" degrees="116.033074" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="1" value="116.03307">E 116.033074°</geoCoordinate>
, 6 or
<date id="FFF32DF1FF99FFA8FC08B623FBB6FEA0" box="[901,1089,324,353]" pageId="1" pageNumber="2" value="2019-09-11">
<collectingDate id="EFB7D419FF99FFA8FC08B623FBB6FEA0" box="[901,1089,324,353]" pageId="1" pageNumber="2" value="2019-09-11">11 SEP 2019</collectingDate>
</date>
, coll.
<collectingMunicipality id="6B96914BFF99FFA8FB1EB624FF4EFE47" pageId="1" pageNumber="2">David Knowles and Kerry Stuart</collectingMunicipality>
). All types will be deposited in the
<location id="8E925DEAFF99FFA8FD1AB60DFBECFE47" LSID="urn:lsid:plazi:treatment:03E4BA27FF99FFBDFDC6B7CBFBC0FA57:8E925DEAFF99FFA8FD1AB60DFBECFE47" box="[663,1051,361,390]" latitude="-32.06286" longLatPrecision="1" longitude="116.03307" municipality="David Knowles and Kerry Stuart" name="Western Australian Museum" pageId="1" pageNumber="2" stateProvince="Western Australia">Western Australian Museum</location>
,
<location id="8E925DEAFF99FFA8FBAAB60EFB83FE47" LSID="urn:lsid:plazi:treatment:03E4BA27FF99FFBDFDC6B7CBFBC0FA57:8E925DEAFF99FFA8FBAAB60EFB83FE47" box="[1063,1140,361,390]" latitude="-32.06286" longLatPrecision="1" longitude="116.03307" municipality="David Knowles and Kerry Stuart" name="Perth" pageId="1" pageNumber="2" stateProvince="Western Australia">Perth</location>
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.
</paragraph>
</subSubSection>
<subSubSection id="C35758BAFF99FFA8FFEDB6D4FF0EFD80" pageId="1" pageNumber="2" type="etymology">
<paragraph id="8BF20B31FF99FFA8FFEDB6D4FF0EFD80" blockId="1.[96,1535,435,577]" pageId="1" pageNumber="2">
<emphasis id="B939D723FF99FFA8FFEDB6D4FF1AFE10" box="[96,237,435,465]" italics="true" pageId="1" pageNumber="2">Etymology</emphasis>
. The species group name (
<taxonomicName id="4C4D70B2FF99FFA8FD03B6D4FCF8FE10" authorityName="Otto &amp; Hill" authorityYear="2019" box="[654,783,435,465]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF99FFA8FD03B6D4FCF8FE10" box="[654,783,435,465]" italics="true" pageId="1" pageNumber="2">banyowla</emphasis>
</taxonomicName>
, noun in apposition, pronounced &quot;ban-yow-la&quot;) is a reference to the discovery of this species in Banyowla Regional Park. Before 2008 this was known as Kelmscott-Martin Regional Park. Banyowla was a Nyoongar (Aboriginal) elder at the time of European settlement.
</paragraph>
</subSubSection>
<subSubSection id="C35758BAFF99FFA8FFEDB509FCA3FC53" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8BF20B31FF99FFA8FFEDB509FCA3FC53" blockId="1.[96,1536,622,915]" pageId="1" pageNumber="2">
<emphasis id="B939D723FF99FFA8FFEDB509FF16FD4D" box="[96,225,622,652]" italics="true" pageId="1" pageNumber="2">Diagnosis</emphasis>
.
<taxonomicName id="4C4D70B2FF99FFA8FF75B509FE07FD4D" authorityName="Otto &amp; Hill" authorityYear="2019" box="[248,496,622,652]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF99FFA8FF75B509FE07FD4D" box="[248,496,622,652]" italics="true" pageId="1" pageNumber="2">Maratus banyowla</emphasis>
</taxonomicName>
are closely related to
<taxonomicName id="4C4D70B2FF99FFA8FC90B509FB48FD4D" authority="Otto &amp; Hill 2015" authorityName="Otto &amp; Hill" authorityYear="2015" box="[797,1215,622,652]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B939D723FF99FFA8FC90B509FC20FD4D" box="[797,983,622,652]" italics="true" pageId="1" pageNumber="2">M. personatus</emphasis>
<bibRefCitation id="EFDC76C0FF99FFA8FC52B508FB48FD4D" author="Otto, J. C. &amp; D. E. Hill" box="[991,1215,623,652]" journalOrPublisher="Peckhamia 127.1" pageId="1" pageNumber="2" pagination="1 - 30" refId="ref2603" refString="Otto, J. C. and D. E. Hill. 2015. Maratus personatus, a masked peacock spider from Cape Riche, Western Australia (Araneae: Salticidae: Euophryinae). Peckhamia 127.1: 1 - 30." title="Maratus personatus, a masked peacock spider from Cape Riche, Western Australia (Araneae: Salticidae: Euophryinae)" type="book chapter" year="2015">Otto &amp; Hill 2015</bibRefCitation>
</taxonomicName>
, found to the southeast at Cape Riche. The detailed structures of the male pedipalps (tegulum, embolus, and RTA) of
<taxonomicName id="4C4D70B2FF99FFA8FAD5B5F4F9F7FD70" authorityName="Otto &amp; Hill" authorityYear="2019" box="[1368,1536,659,689]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF99FFA8FAD5B5F4F9F7FD70" box="[1368,1536,659,689]" italics="true" pageId="1" pageNumber="2">M. banyowla</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D70B2FF99FFA8FF13B5DEFEACFD16" authorityName="Otto &amp; Hill" authorityYear="2015" box="[158,347,697,727]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B939D723FF99FFA8FF13B5DEFEACFD16" box="[158,347,697,727]" italics="true" pageId="1" pageNumber="2">M. personatus</emphasis>
</taxonomicName>
are virtually the same, very much like those of
<taxonomicName id="4C4D70B2FF99FFA8FC71B5DEFB9BFD16" authorityName="Karsch" authorityYear="1878" box="[1020,1132,697,727]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B939D723FF99FFA8FC71B5DEFB9BFD16" box="[1020,1132,697,727]" italics="true" pageId="1" pageNumber="2">Maratus</emphasis>
</taxonomicName>
in the
<emphasis id="B939D723FF99FFA8FB5FB5DEFADAFD16" box="[1234,1325,697,727]" italics="true" pageId="1" pageNumber="2">linnaei</emphasis>
,
<emphasis id="B939D723FF99FFA8FAB3B5DEFA35FD16" box="[1342,1474,697,727]" italics="true" pageId="1" pageNumber="2">mungaich</emphasis>
and
<emphasis id="B939D723FF99FFA8FFEDB5B9FF5CFD3D" box="[96,171,734,764]" italics="true" pageId="1" pageNumber="2">vespa</emphasis>
groups from
<collectingRegion id="4989C5D3FF99FFA8FEF8B587FD82FD3C" box="[373,629,736,765]" country="Australia" name="Western Australia" pageId="1" pageNumber="2">Western Australia</collectingRegion>
. Males of both species have a blue &quot;mask&quot; (lighter blue in
<taxonomicName id="4C4D70B2FF99FFA8FA53B5B9FF16FCE3" authorityName="Otto &amp; Hill" authorityYear="2019" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF99FFA8FA53B5B9FF16FCE3" italics="true" pageId="1" pageNumber="2">M. banyowla</emphasis>
</taxonomicName>
) but the males of
<taxonomicName id="4C4D70B2FF99FFA8FE6FB463FD7BFCE3" authorityName="Otto &amp; Hill" authorityYear="2019" box="[482,652,772,802]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF99FFA8FE6FB463FD7BFCE3" box="[482,652,772,802]" italics="true" pageId="1" pageNumber="2">M. banyowla</emphasis>
</taxonomicName>
can readily be identified by the presence of bright yellow setae on legs I-II, femora of legs III, and pedipalps, and a distinctive zig-zag scale pattern across the carapace (
<figureCitation id="137617B4FF99FFA8FFE4B437FF16FCAC" box="[105,225,848,877]" captionStart="Figure 2" captionStartId="2.[197,277,1914,1938]" captionTargetBox="[198,1437,165,1903]" captionTargetId="figure@2.[813,1437,765,1322]" captionTargetPageId="2" captionText="Figure 2. Comparison of adult male Maratus banyowla (1-3) with adult male M. personatus (4-6). Note the bright yellow setae on the pedipalps, legs I-II, femora of legs III, and pedipalps, and the distinctive zig-zag line across the eye region of M. banyowla." figureDoi="http://doi.org/10.5281/zenodo.3732665" httpUri="https://zenodo.org/record/3732665/files/figure.png" pageId="1" pageNumber="2">Figure 2</figureCitation>
). Some female
<taxonomicName id="4C4D70B2FF99FFA8FE36B428FD93FCAC" authorityName="Otto &amp; Hill" authorityYear="2019" box="[443,612,847,877]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF99FFA8FE36B428FD93FCAC" box="[443,612,847,877]" italics="true" pageId="1" pageNumber="2">M. banyowla</emphasis>
</taxonomicName>
also exhibit a less distinct zig-zag pattern across the carapace. We place both species in a new clade, the
<taxonomicName id="4C4D70B2FF99FFA8FDEFB413FD02FC53" authorityName="Otto &amp; Hill" authorityYear="2015" box="[610,757,884,914]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B939D723FF99FFA8FDEFB413FD02FC53" box="[610,757,884,914]" italics="true" pageId="1" pageNumber="2">personatus</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="C35758BAFF99FFA5FFEDB4D8FBF5F89F" lastPageId="12" lastPageNumber="13" pageId="1" pageNumber="2" type="description">
<paragraph id="8BF20B31FF99FFA8FFEDB4D8FB82FB25" blockId="1.[96,1535,959,1252]" pageId="1" pageNumber="2">
<emphasis id="B939D723FF99FFA8FFEDB4D8FE93FC1C" box="[96,356,959,989]" italics="true" pageId="1" pageNumber="2">Description of male</emphasis>
(
<figureCitation id="137617B4FF99FFA8FEF9B4A6FE0DFC1F" box="[372,506,961,990]" captionStart="Figure 2" captionStartId="2.[197,277,1914,1938]" captionTargetBox="[198,1437,165,1903]" captionTargetId="figure@2.[813,1437,765,1322]" captionTargetPageId="2" captionText="Figure 2. Comparison of adult male Maratus banyowla (1-3) with adult male M. personatus (4-6). Note the bright yellow setae on the pedipalps, legs I-II, femora of legs III, and pedipalps, and the distinctive zig-zag line across the eye region of M. banyowla." figureDoi="http://doi.org/10.5281/zenodo.3732665" httpUri="https://zenodo.org/record/3732665/files/figure.png" pageId="1" pageNumber="2">Figures 2</figureCitation>
:1-3, 3-5). Males (n=7) ranged from
<quantity id="4CB5A6D4FF99FFA8FC6AB4A6FB54FC1C" box="[999,1187,961,990]" metricMagnitude="-3" metricUnit="m" metricValue="3.9499999999999997" metricValueMax="4.1" metricValueMin="3.8" pageId="1" pageNumber="2" unit="mm" value="3.9499999999999997" valueMax="4.1" valueMin="3.8">3.8 to 4.1 mm</quantity>
in length. The chelicerae are short, black and glabrous. Long white setae extend anteromedially from the clypeus over the proximal part of each paturon. Light blue scales surround the anterior eyes and extend to the rear over the eye region, separated from dark cuticle at the rear by a distinctive zig-zag line. The PME are closer to the PLE than to the ALE. A middorsal thoracic tract of white setae is present. There is a tract of white setae below the lateral eyes, extending to the rear along the top of the carapace. A distinct band of white marginal setae is also present on either side of the carapace. Otherwise the sides of the carapace, and the dorsal carapace behind the eye region, are dark brown to black and glabrous.
</paragraph>
<paragraph id="8BF20B31FF99FFA8FFEDB275FF2AFA5E" blockId="1.[96,1534,1298,1439]" pageId="1" pageNumber="2">The opisthosoma has a dorsal plate but no lateral flaps. The dorsal plate of the opisthosoma is dark brown and shiny, with white setae around the lateral margins, and a wide but short anterodorsal tract of white setae. The ventral opisthosoma, coxae, sternum, labium and endites are brown and mostly glabrous.</paragraph>
<paragraph id="8BF20B31FF99FFA8FFEDB2A9FA57F931" blockId="1.[96,1536,1485,1776]" pageId="1" pageNumber="2">
Legs I and II are shorter, legs III and IV longer, and legs III by far the longest. Legs I and II, and the anterior femora of legs IV, are covered with bright yellow setae. The patellae and tibiae of legs III bear black stripes on a background of white setae. The metatarsi of legs III are black, and the tarsi of legs III are white. Legs IV are ringed with alternating bands of black cuticle and white setae. The patella, tibia and proximal cymbium of each pedipalp are covered dorsally with bright yellow setae. Each distal cymbium is covered with long white setae. The distally serrated RTA, tegulum and apophysis of each pedipalp are typical of those seen in many western
<taxonomicName id="4C4D70B2FF99FFA8FC92B1CBFC78F90B" authorityName="Karsch" authorityYear="1878" box="[799,911,1708,1738]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B939D723FF99FFA8FC92B1CBFC78F90B" box="[799,911,1708,1738]" italics="true" pageId="1" pageNumber="2">Maratus</emphasis>
</taxonomicName>
species (Figure 5). The heavier outer apex of the apophysis bears a single serration, and below this is a smaller, sharply pointed inner apophysis.
</paragraph>
<caption id="DF325BB9FF9AFFABFF48B01DFD90F810" ID-DOI="http://doi.org/10.5281/zenodo.3732665" ID-Zenodo-Dep="3732665" httpUri="https://zenodo.org/record/3732665/files/figure.png" pageId="2" pageNumber="3" startId="2.[197,277,1914,1938]" targetBox="[198,1437,165,1903]" targetPageId="2">
<paragraph id="8BF20B31FF9AFFABFF48B01DFD90F810" blockId="2.[197,1437,1872,2002]" pageId="2" pageNumber="3">
<emphasis id="B939D723FF9AFFABFF48B01DFEC4F853" bold="true" box="[197,307,1914,1938]" pageId="2" pageNumber="3">Figure 2.</emphasis>
Comparison of adult male
<taxonomicName id="4C4D70B2FF9AFFABFDFCB01EFCB6F853" authorityName="Otto &amp; Hill" authorityYear="2019" box="[625,833,1913,1938]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF9AFFABFDFCB01EFCB6F853" box="[625,833,1913,1938]" italics="true" pageId="2" pageNumber="3">Maratus banyowla</emphasis>
</taxonomicName>
(1-3) with adult male
<taxonomicName id="4C4D70B2FF9AFFABFBCAB01EFB13F853" authorityName="Otto &amp; Hill" authorityYear="2015" box="[1095,1252,1913,1938]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B939D723FF9AFFABFBCAB01EFB13F853" box="[1095,1252,1913,1938]" italics="true" pageId="2" pageNumber="3">M. personatus</emphasis>
</taxonomicName>
(4-6). Note the bright yellow setae on the pedipalps, legs I-II, femora of legs III, and pedipalps, and the distinctive zig-zag line across the eye region of
<taxonomicName id="4C4D70B2FF9AFFABFE5BB0DFFD96F810" authorityName="Otto &amp; Hill" authorityYear="2019" box="[470,609,1976,2001]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF9AFFABFE5BB0DFFD96F810" box="[470,609,1976,2001]" italics="true" pageId="2" pageNumber="3">M. banyowla</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF325BB9FF9BFFAAFFEDB047FC7DF8F6" box="[96,906,1823,1848]" pageId="3" pageNumber="4">
<paragraph id="8BF20B31FF9BFFAAFFEDB047FC7DF8F6" blockId="3.[96,906,1782,1848]" box="[96,906,1823,1848]" pageId="3" pageNumber="4">
<emphasis id="B939D723FF9BFFAAFFEDB047FDF4F8F9" bold="true" box="[96,515,1824,1848]" pageId="3" pageNumber="4">Figure 3 (continued on next page).</emphasis>
Male types for
<taxonomicName id="4C4D70B2FF9BFFAAFD3BB078FC73F8F9" authorityName="Otto &amp; Hill" authorityYear="2019" box="[694,900,1823,1848]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF9BFFAAFD3BB078FC73F8F9" box="[694,900,1823,1848]" italics="true" pageId="3" pageNumber="4">Maratus banyowla</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF325BB9FF9CFFADFFEDB051FAF3F88C" ID-DOI="http://doi.org/10.5281/zenodo.3732667" ID-Zenodo-Dep="3732667" box="[96,1284,1845,1870]" httpUri="https://zenodo.org/record/3732667/files/figure.png" pageId="4" pageNumber="5" startId="4.[96,175,1846,1870]" targetBox="[96,1537,165,1833]" targetPageId="4">
<paragraph id="8BF20B31FF9CFFADFFEDB051FAF3F88C" blockId="4.[96,1284,1803,1870]" box="[96,1284,1845,1870]" pageId="4" pageNumber="5">
<emphasis id="B939D723FF9CFFADFFEDB051FC8AF88F" bold="true" box="[96,893,1846,1870]" pageId="4" pageNumber="5">Figure 3 (continued from previous page, continued on next page).</emphasis>
Male types for
<taxonomicName id="4C4D70B2FF9CFFADFBBCB052FB08F88F" authorityName="Otto &amp; Hill" authorityYear="2019" box="[1073,1279,1845,1870]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF9CFFADFBBCB052FB08F88F" box="[1073,1279,1845,1870]" italics="true" pageId="4" pageNumber="5">Maratus banyowla</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF325BB9FF9DFFACFFEDB0D5FAF3F80B" ID-DOI="http://doi.org/10.5281/zenodo.3732669" ID-Zenodo-Dep="3732669" box="[96,1284,1969,1995]" httpUri="https://zenodo.org/record/3732669/files/figure.png" pageId="5" pageNumber="6" startId="5.[96,175,1970,1994]" targetBox="[96,1537,165,1961]" targetPageId="5">
<paragraph id="8BF20B31FF9DFFACFFEDB0D5FAF3F80B" blockId="5.[96,1284,1929,1995]" box="[96,1284,1969,1995]" pageId="5" pageNumber="6">
<emphasis id="B939D723FF9DFFACFFEDB0D5FC8AF80B" bold="true" box="[96,893,1970,1995]" pageId="5" pageNumber="6">Figure 3 (continued from previous page, continued on next page).</emphasis>
Male types for
<taxonomicName id="4C4D70B2FF9DFFACFBBCB0D6FB08F80B" authorityName="Otto &amp; Hill" authorityYear="2019" box="[1073,1279,1969,1994]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF9DFFACFBBCB0D6FB08F80B" box="[1073,1279,1969,1994]" italics="true" pageId="5" pageNumber="6">Maratus banyowla</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF325BB9FF9EFFAFFFEDB03FFC2BF8B1" ID-DOI="http://doi.org/10.5281/zenodo.3732671" ID-Zenodo-Dep="3732671" box="[96,988,1879,1905]" httpUri="https://zenodo.org/record/3732671/files/figure.png" pageId="6" pageNumber="7" startId="6.[96,175,1880,1904]" targetBox="[96,1536,165,1868]" targetPageId="6">
<paragraph id="8BF20B31FF9EFFAFFFEDB03FFC2BF8B1" blockId="6.[96,988,1838,1905]" box="[96,988,1879,1905]" pageId="6" pageNumber="7">
<emphasis id="B939D723FF9EFFAFFFEDB03FFDA2F8B1" bold="true" box="[96,597,1880,1904]" pageId="6" pageNumber="7">Figure 3 (continued from previous page).</emphasis>
Male types for
<taxonomicName id="4C4D70B2FF9EFFAFFC84B030FC20F8B1" authorityName="Otto &amp; Hill" authorityYear="2019" box="[777,983,1879,1904]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF9EFFAFFC84B030FC20F8B1" box="[777,983,1879,1904]" italics="true" pageId="6" pageNumber="7">Maratus banyowla</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF20B31FF9FFFAEFFEDB1C4FBF4F97A" blockId="7.[96,1027,1699,1724]" box="[96,1027,1698,1724]" pageId="7" pageNumber="8">
<emphasis id="B939D723FF9FFFAEFFEDB1C4FDF4F97A" bold="true" box="[96,515,1699,1723]" pageId="7" pageNumber="8">
<figureCitation id="137617B4FF9FFFAEFFEDB1C4FF32F97A" box="[96,197,1699,1723]" captionStart="Figure 4" captionStartId="8.[106,185,1983,2007]" captionTargetBox="[106,1527,165,1974]" captionTargetId="figure@8.[426,893,1019,1505]" captionTargetPageId="8" captionText="Figure 4 (continued from previous page, continued on next page). Male types for Maratus banyowla in ethanol." figureDoi="http://doi.org/10.5281/zenodo.3732673" httpUri="https://zenodo.org/record/3732673/files/figure.png" pageId="7" pageNumber="8">Figure 4</figureCitation>
(continued on next page).
</emphasis>
Male
<typeStatus id="54F6B593FF9FFFAEFDC1B1C3FD7DF97D" box="[588,650,1700,1724]" pageId="7" pageNumber="8">types</typeStatus>
for
<taxonomicName id="4C4D70B2FF9FFFAEFD3BB1C5FC73F97A" authorityName="Otto &amp; Hill" authorityYear="2019" box="[694,900,1698,1723]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF9FFFAEFD3BB1C5FC73F97A" box="[694,900,1698,1723]" italics="true" pageId="7" pageNumber="8">Maratus banyowla</emphasis>
</taxonomicName>
in ethanol.
</paragraph>
<caption id="DF325BB9FF90FFA1FFE7B0D8FA70F816" ID-DOI="http://doi.org/10.5281/zenodo.3732673" ID-Zenodo-Dep="3732673" box="[106,1415,1982,2008]" httpUri="https://zenodo.org/record/3732673/files/figure.png" pageId="8" pageNumber="9" startId="8.[106,185,1983,2007]" targetBox="[106,1527,165,1974]" targetPageId="8">
<paragraph id="8BF20B31FF90FFA1FFE7B0D8FA70F816" blockId="8.[106,1415,1941,2008]" box="[106,1415,1982,2008]" pageId="8" pageNumber="9">
<emphasis id="B939D723FF90FFA1FFE7B0D8FC71F816" bold="true" box="[106,902,1983,2007]" pageId="8" pageNumber="9">Figure 4 (continued from previous page, continued on next page).</emphasis>
Male types for
<taxonomicName id="4C4D70B2FF90FFA1FBB7B0D9FAFFF816" authorityName="Otto &amp; Hill" authorityYear="2019" box="[1082,1288,1982,2007]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF90FFA1FBB7B0D9FAFFF816" box="[1082,1288,1982,2007]" italics="true" pageId="8" pageNumber="9">Maratus banyowla</emphasis>
</taxonomicName>
in ethanol.
</paragraph>
</caption>
<caption id="DF325BB9FF91FFA0FFE7B0A5FA70F81B" ID-DOI="http://doi.org/10.5281/zenodo.3732675" ID-Zenodo-Dep="3732675" box="[106,1415,1985,2011]" httpUri="https://zenodo.org/record/3732675/files/figure.png" pageId="9" pageNumber="10" startId="9.[106,185,1986,2010]" targetBox="[106,1527,165,1976]" targetPageId="9">
<paragraph id="8BF20B31FF91FFA0FFE7B0A5FA70F81B" blockId="9.[106,1415,1985,2011]" box="[106,1415,1985,2011]" pageId="9" pageNumber="10">
<emphasis id="B939D723FF91FFA0FFE7B0A5FC71F81B" bold="true" box="[106,902,1986,2010]" pageId="9" pageNumber="10">Figure 4 (continued from previous page, continued on next page).</emphasis>
Male types for
<taxonomicName id="4C4D70B2FF91FFA0FBB7B0A6FAFFF81B" authorityName="Otto &amp; Hill" authorityYear="2019" box="[1082,1288,1985,2010]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF91FFA0FBB7B0A6FAFFF81B" box="[1082,1288,1985,2010]" italics="true" pageId="9" pageNumber="10">Maratus banyowla</emphasis>
</taxonomicName>
in ethanol.
</paragraph>
</caption>
<caption id="DF325BB9FF92FFA3FFD2B000FBA3F841" ID-DOI="http://doi.org/10.5281/zenodo.3732677" ID-Zenodo-Dep="3732677" box="[95,1108,1894,1920]" httpUri="https://zenodo.org/record/3732677/files/figure.png" pageId="10" pageNumber="11" startId="10.[95,174,1895,1919]" targetBox="[96,1536,165,1885]" targetPageId="10">
<paragraph id="8BF20B31FF92FFA3FFD2B000FBA3F841" blockId="10.[95,1108,1894,1920]" box="[95,1108,1894,1920]" pageId="10" pageNumber="11">
<emphasis id="B939D723FF92FFA3FFD2B000FDA4F8BE" bold="true" box="[95,595,1895,1920]" pageId="10" pageNumber="11">Figure 4 (continued from previous page).</emphasis>
Male types for
<taxonomicName id="4C4D70B2FF92FFA3FC8AB001FC22F8BE" authorityName="Otto &amp; Hill" authorityYear="2019" box="[775,981,1894,1919]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF92FFA3FC8AB001FC22F8BE" box="[775,981,1894,1919]" italics="true" pageId="10" pageNumber="11">Maratus banyowla</emphasis>
</taxonomicName>
in ethanol.
</paragraph>
</caption>
<paragraph id="8BF20B31FF94FFA5FFEDB34BFF4FFAED" blockId="12.[96,1534,1068,1324]" pageId="12" pageNumber="13">
<emphasis id="B939D723FF94FFA5FFEDB34BFE89FB8B" box="[96,382,1068,1098]" italics="true" pageId="12" pageNumber="13">Description of female</emphasis>
(
<figureCitation id="137617B4FF94FFA5FE1CB349FDC3FB8A" box="[401,564,1070,1099]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStart-2="Figure 8" captionStartId-0="13.[96,175,1864,1888]" captionStartId-1="16.[96,175,1836,1860]" captionStartId-2="17.[96,175,1499,1523]" captionTargetBox-0="[96,1537,165,1855]" captionTargetBox-1="[96,1537,165,1823]" captionTargetBox-2="[96,1536,1146,1489]" captionTargetId-0="figure@13.[587,1048,545,1060]" captionTargetId-1="figure@16.[785,1119,701,1275]" captionTargetId-2="figure@17.[486,822,1146,1488]" captionTargetPageId-0="13" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Figure 6 (continued on next page). Female types for Maratus banyowla." captionText-1="Figure 7 (continued on next page). Female types for Maratus banyowla in ethanol." captionText-2="Figure 8. Ventral view of the epigynum of the female types for Maratus banyowla in ethanol. The anterior direction is toward the top of this page." figureDoi-0="http://doi.org/10.5281/zenodo.3732679" figureDoi-1="http://doi.org/10.5281/zenodo.3732685" figureDoi-2="http://doi.org/10.5281/zenodo.3732689" httpUri-0="https://zenodo.org/record/3732679/files/figure.png" httpUri-1="https://zenodo.org/record/3732685/files/figure.png" httpUri-2="https://zenodo.org/record/3732689/files/figure.png" pageId="12" pageNumber="13">Figures 6-8</figureCitation>
). Females (n=4) ranged from
<quantity id="4CB5A6D4FF94FFA5FC54B349FB6DFB8A" box="[985,1178,1070,1099]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.1" metricValueMin="5.6" pageId="12" pageNumber="13" unit="mm" value="5.85" valueMax="6.1" valueMin="5.6">5.6 to 6.1 mm</quantity>
in length. The chelicerae are light brown and glabrous. Long off-white setae cover the clypeus. The anterior part of the eye region is covered with a tract of light brown to red-brown setae. At the rear this tract is separated from the dark brown cuticle of the dorsal carapace by a jagged transverse line, less distinct than that of the male. The PME are equidistant from the ALE and PLE. Light brown to red-brown setae also comprise a middorsal thoracic tract, and partly cover the sides of the carapace. The carapace lacks a distinct marginal band of scales.
</paragraph>
<paragraph id="8BF20B31FF94FFA5FFEDB23DFC14F9F3" blockId="12.[96,1534,1370,1586]" pageId="12" pageNumber="13">The cuticle of the dorsal opisthosoma is dark brown with a variable cover of light brown setae, bounded on either side by a broad marginal band of off-white to light brown setae. A very small, triangular tuft of off-white anal setae is present above the spinnerets. Below the opisthosoma is off-white to light brown with a uniform cover of short off-white setae; several irregular lines of small brown spots may be present. From below the coxae, sternum, labium and endites are mostly glabrous, light brown and translucent, with some off-white setae projecting from the rear of the sternum.</paragraph>
<paragraph id="8BF20B31FF94FFA5FFEDB107FD3DF909" blockId="12.[96,1533,1632,1736]" pageId="12" pageNumber="13">The legs and pedipalps are uniform light-brown in colour, mostly translucent with indistinct bands of offwhite setae. Legs I and II are shorter and of similar length, legs III and IV longer and of similar length. The pedipalps have a cover of off-white setae.</paragraph>
<paragraph id="8BF20B31FF94FFA5FFEDB191FBF5F89F" blockId="12.[96,1535,1780,1886]" pageId="12" pageNumber="13">
The epigynum (
<figureCitation id="137617B4FF94FFA5FEB0B191FE42F8D2" box="[317,437,1782,1811]" captionStart="Figure 8" captionStartId="17.[96,175,1499,1523]" captionTargetBox="[96,1536,1146,1489]" captionTargetId="figure@17.[486,822,1146,1488]" captionTargetPageId="17" captionText="Figure 8. Ventral view of the epigynum of the female types for Maratus banyowla in ethanol. The anterior direction is toward the top of this page." figureDoi="http://doi.org/10.5281/zenodo.3732689" httpUri="https://zenodo.org/record/3732689/files/figure.png" pageId="12" pageNumber="13">Figure 8</figureCitation>
) is typical for
<taxonomicName id="4C4D70B2FF94FFA5FD05B193FD0FF8D3" authorityName="Karsch" authorityYear="1878" box="[648,760,1780,1810]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B939D723FF94FFA5FD05B193FD0FF8D3" box="[648,760,1780,1810]" italics="true" pageId="12" pageNumber="13">Maratus</emphasis>
</taxonomicName>
, with a pair of rounded anterior fossae separated by a septum of variable width. Behind each fossa is a larger, rounded posterior spermatheca. Variably sclerotized ducts are visible through the posterior part of each fossa.
</paragraph>
</subSubSection>
<caption id="DF325BB9FF95FFA4FFEDB02FFC52F8A1" ID-DOI="http://doi.org/10.5281/zenodo.3732679" ID-Zenodo-Dep="3732679" box="[96,933,1863,1889]" httpUri="https://zenodo.org/record/3732679/files/figure.png" pageId="13" pageNumber="14" startId="13.[96,175,1864,1888]" targetBox="[96,1537,165,1855]" targetPageId="13">
<paragraph id="8BF20B31FF95FFA4FFEDB02FFC52F8A1" blockId="13.[96,933,1823,1889]" box="[96,933,1863,1889]" pageId="13" pageNumber="14">
<emphasis id="B939D723FF95FFA4FFEDB02FFDF4F8A1" bold="true" box="[96,515,1864,1889]" pageId="13" pageNumber="14">Figure 6 (continued on next page).</emphasis>
Female types for
<taxonomicName id="4C4D70B2FF95FFA4FD5CB020FC68F8A1" authorityName="Otto &amp; Hill" authorityYear="2019" box="[721,927,1863,1888]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF95FFA4FD5CB020FC68F8A1" box="[721,927,1863,1888]" italics="true" pageId="13" pageNumber="14">Maratus banyowla</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF325BB9FF96FFA7FFEDB000FAE8F8BE" ID-DOI="http://doi.org/10.5281/zenodo.3732681" ID-Zenodo-Dep="3732681" box="[96,1311,1894,1920]" httpUri="https://zenodo.org/record/3732681/files/figure.png" pageId="14" pageNumber="15" startId="14.[96,175,1895,1919]" targetBox="[96,1537,165,1884]" targetPageId="14">
<paragraph id="8BF20B31FF96FFA7FFEDB000FAE8F8BE" blockId="14.[96,1311,1855,1920]" box="[96,1311,1894,1920]" pageId="14" pageNumber="15">
<emphasis id="B939D723FF96FFA7FFEDB000FC8AF8BE" bold="true" box="[96,893,1895,1920]" pageId="14" pageNumber="15">Figure 6 (continued from previous page, continued on next page).</emphasis>
Female types for
<taxonomicName id="4C4D70B2FF96FFA7FBC6B001FAEDF8BE" authorityName="Otto &amp; Hill" authorityYear="2019" box="[1099,1306,1894,1919]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF96FFA7FBC6B001FAEDF8BE" box="[1099,1306,1894,1919]" italics="true" pageId="14" pageNumber="15">Maratus banyowla</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF325BB9FF97FFA6FFEDB0F7FC00F869" ID-DOI="http://doi.org/10.5281/zenodo.3732683" ID-Zenodo-Dep="3732683" box="[96,1015,1935,1961]" httpUri="https://zenodo.org/record/3732683/files/figure.png" pageId="15" pageNumber="16" startId="15.[96,175,1936,1960]" targetBox="[96,1537,165,1927]" targetPageId="15">
<paragraph id="8BF20B31FF97FFA6FFEDB0F7FC00F869" blockId="15.[96,1064,1896,1961]" box="[96,1015,1935,1961]" pageId="15" pageNumber="16">
<emphasis id="B939D723FF97FFA6FFEDB0F7FDA2F869" bold="true" box="[96,597,1936,1960]" pageId="15" pageNumber="16">Figure 6 (continued from previous page).</emphasis>
Female types for
<taxonomicName id="4C4D70B2FF97FFA6FCAEB0E8FC06F869" authorityName="Otto &amp; Hill" authorityYear="2019" box="[803,1009,1935,1960]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF97FFA6FCAEB0E8FC06F869" box="[803,1009,1935,1960]" italics="true" pageId="15" pageNumber="16">Maratus banyowla</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF325BB9FF88FFB9FFEDB04BFBE9F885" ID-DOI="http://doi.org/10.5281/zenodo.3732685" ID-Zenodo-Dep="3732685" box="[96,1054,1835,1861]" httpUri="https://zenodo.org/record/3732685/files/figure.png" pageId="16" pageNumber="17" startId="16.[96,175,1836,1860]" targetBox="[96,1537,165,1823]" targetPageId="16">
<paragraph id="8BF20B31FF88FFB9FFEDB04BFBE9F885" blockId="16.[96,1054,1792,1861]" box="[96,1054,1835,1861]" pageId="16" pageNumber="17">
<emphasis id="B939D723FF88FFB9FFEDB04BFDF4F885" bold="true" box="[96,515,1836,1861]" pageId="16" pageNumber="17">Figure 7 (continued on next page).</emphasis>
Female types for
<taxonomicName id="4C4D70B2FF88FFB9FD5CB04CFC68F885" authorityName="Otto &amp; Hill" authorityYear="2019" box="[721,927,1835,1860]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF88FFB9FD5CB04CFC68F885" box="[721,927,1835,1860]" italics="true" pageId="16" pageNumber="17">Maratus banyowla</emphasis>
</taxonomicName>
in ethanol.
</paragraph>
</caption>
<caption id="DF325BB9FF89FFB8FFEDB344FB87FBFA" ID-DOI="http://doi.org/10.5281/zenodo.3732687" ID-Zenodo-Dep="3732687" box="[96,1136,1058,1084]" httpUri="https://zenodo.org/record/3732687/files/figure.png" pageId="17" pageNumber="18" startId="17.[96,175,1059,1083]" targetBox="[96,1536,165,1048]" targetPageId="17">
<paragraph id="8BF20B31FF89FFB8FFEDB344FB87FBFA" blockId="17.[96,1136,1058,1084]" box="[96,1136,1058,1084]" pageId="17" pageNumber="18">
<emphasis id="B939D723FF89FFB8FFEDB344FDA2FBFA" bold="true" box="[96,597,1059,1083]" pageId="17" pageNumber="18">Figure 7 (continued from previous page).</emphasis>
Female types for
<taxonomicName id="4C4D70B2FF89FFB8FCAEB345FC06FBFA" authorityName="Otto &amp; Hill" authorityYear="2019" box="[803,1009,1058,1083]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF89FFB8FCAEB345FC06FBFA" box="[803,1009,1058,1083]" italics="true" pageId="17" pageNumber="18">Maratus banyowla</emphasis>
</taxonomicName>
in ethanol.
</paragraph>
</caption>
<caption id="DF325BB9FF89FFB8FFEDB2BCFEC8F9D2" ID-DOI="http://doi.org/10.5281/zenodo.3732689" ID-Zenodo-Dep="3732689" httpUri="https://zenodo.org/record/3732689/files/figure.png" pageId="17" pageNumber="18" startId="17.[96,175,1499,1523]" targetBox="[96,1536,1146,1489]" targetPageId="17">
<paragraph id="8BF20B31FF89FFB8FFEDB2BCFEC8F9D2" blockId="17.[96,1535,1451,1555]" pageId="17" pageNumber="18">
<emphasis id="B939D723FF89FFB8FFEDB2BCFF3BFA32" bold="true" box="[96,204,1499,1523]" pageId="17" pageNumber="18">Figure 8.</emphasis>
Ventral view of the epigynum of the female types for
<taxonomicName id="4C4D70B2FF89FFB8FCBBB2BDFBF2FA32" authorityName="Otto &amp; Hill" authorityYear="2019" box="[822,1029,1498,1523]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF89FFB8FCBBB2BDFBF2FA32" box="[822,1029,1498,1523]" italics="true" pageId="17" pageNumber="18">Maratus banyowla</emphasis>
</taxonomicName>
in ethanol. The anterior direction is toward the top of this page.
</paragraph>
</caption>
<subSubSection id="C35758BAFF89FFBDFFEDB127FBC0FA57" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18" type="biology_ecology">
<paragraph id="8BF20B31FF89FFB8FFEDB127FE96F935" blockId="17.[96,1536,1600,1780]" pageId="17" pageNumber="18">
<emphasis id="B939D723FF89FFB8FFEDB127FF31F99F" box="[96,198,1600,1630]" italics="true" pageId="17" pageNumber="18">Habitat</emphasis>
.
<taxonomicName id="4C4D70B2FF89FFB8FF52B127FE2FF99F" authorityName="Otto &amp; Hill" authorityYear="2019" box="[223,472,1600,1630]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF89FFB8FF52B127FE2FF99F" box="[223,472,1600,1630]" italics="true" pageId="17" pageNumber="18">Maratus banyowla</emphasis>
</taxonomicName>
were collected on or near the ground in the Banyowla Regional Park, an area with many quartzite dykes exposed at the surface and a complex plant community (
<emphasis id="B939D723FF89FFB8FB71B102FA00F942" box="[1276,1527,1637,1667]" italics="true" pageId="17" pageNumber="18">Wandoo woodland</emphasis>
;
<figureCitation id="137617B4FF89FFB8FFEDB1EBFF2BF968" box="[96,220,1676,1705]" captionStart="Figure 9" captionStartId="18.[96,175,653,677]" captionTargetBox="[96,1537,165,643]" captionTargetPageId="18" captionText="Figure 9. Habitat of Maratus banyowla at Banyowla Regional Park near Perth. 1, Exposed quartize dyke in Wandoo woodland. 2, Note the many broken rock fragments littering the ground at this site. Photographs © by David Knowles, used with permission." figureDoi="http://doi.org/10.5281/zenodo.3732691" httpUri="https://zenodo.org/record/3732691/files/figure.png" pageId="17" pageNumber="18">Figure 9</figureCitation>
;
<bibRefCitation id="EFDC76C0FF89FFB8FF60B1EAFE31F968" author="Ecoscape" box="[237,454,1676,1705]" pageId="17" pageNumber="18" pagination="1 - 68" publicationUrl="https://www.gosnells.wa.gov.au/files/sharedassets/public/pdfs/environment/ellis_brook_valley_management_plan.pdf" refId="ref2439" refString="Ecoscape. 2016. Ellis Brook Valley Reserve Strategic Management Plan 2016 - 2026 (9689 - 3216 - 14 R final rev 1). City of Gosnells, Ecoscape (Australia) Pty Ltd: i-v, 1 - 68, online at https: // www. gosnells. wa. gov. au / files / sharedassets / public / pdfs / environment / ellis _ brook _ valley _ management _ plan. pdf" title="Ellis Brook Valley Reserve Strategic Management Plan 2016 - 2026 (9689 - 3216 - 14 R final rev 1). City of Gosnells, Ecoscape (Australia) Pty Ltd: i-v" type="book chapter" year="2016">Ecoscape 2016</bibRefCitation>
). Two large quarries are situated near the western boundary of this park.
<figureCitation id="137617B4FF89FFB8FFEDB1D6FF10F90E" box="[96,231,1713,1743]" captionStart="Figure 10" captionStartId="18.[96,175,1712,1736]" captionTargetBox="[96,1537,804,1701]" captionTargetId="figure@18.[96,834,1171,1698]" captionTargetPageId="18" captionText="Figure 10 (continued on next page). Maratus banyowla in their complex microhabitat at Banyowla Regional Park. 1-5, Adult males. Photographs © by Kerry Stuart, used with permission." figureDoi="http://doi.org/10.5281/zenodo.3732693" httpUri="https://zenodo.org/record/3732693/files/figure.png" pageId="17" pageNumber="18">Figure 10</figureCitation>
shows live individuals in their natural surroundings and illustrates some of the complexity of their microhabitat.
</paragraph>
<caption id="DF325BB9FF8AFFBBFFEDB5EAFF11FD25" ID-DOI="http://doi.org/10.5281/zenodo.3732691" ID-Zenodo-Dep="3732691" httpUri="https://zenodo.org/record/3732691/files/figure.png" pageId="18" pageNumber="19" startId="18.[96,175,653,677]" targetBox="[96,1537,165,643]" targetPageId="18">
<paragraph id="8BF20B31FF8AFFBBFFEDB5EAFF11FD25" blockId="18.[96,1536,614,740]" pageId="18" pageNumber="19">
<emphasis id="B939D723FF8AFFBBFFEDB5EAFF3CFD64" bold="true" box="[96,203,653,677]" pageId="18" pageNumber="19">Figure 9.</emphasis>
Habitat of
<taxonomicName id="4C4D70B2FF8AFFBBFEC2B5EBFDEAFD64" authorityName="Otto &amp; Hill" authorityYear="2019" box="[335,541,652,677]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF8AFFBBFEC2B5EBFDEAFD64" box="[335,541,652,677]" italics="true" pageId="18" pageNumber="19">Maratus banyowla</emphasis>
</taxonomicName>
at Banyowla Regional Park near Perth.
<emphasis id="B939D723FF8AFFBBFC69B5EAFC0DFD64" bold="true" box="[996,1018,653,677]" pageId="18" pageNumber="19">1,</emphasis>
Exposed quartize dyke in Wandoo woodland.
<emphasis id="B939D723FF8AFFBBFFEDB5CBFF81FD05" bold="true" box="[96,118,684,708]" pageId="18" pageNumber="19">2,</emphasis>
Note the many broken rock fragments littering the ground at this site. Photographs © by David Knowles, used with permission.
</paragraph>
</caption>
<caption id="DF325BB9FF8AFFBBFFEDB1D7FCEBF926" ID-DOI="http://doi.org/10.5281/zenodo.3732693" ID-Zenodo-Dep="3732693" httpUri="https://zenodo.org/record/3732693/files/figure.png" pageId="18" pageNumber="19" startId="18.[96,175,1712,1736]" targetBox="[96,1537,804,1701]" targetPageId="18">
<paragraph id="8BF20B31FF8AFFBBFFEDB1D7FCEBF926" blockId="18.[96,1536,1668,1768]" pageId="18" pageNumber="19">
<emphasis id="B939D723FF8AFFBBFFEDB1D7FDE3F909" bold="true" box="[96,532,1712,1736]" pageId="18" pageNumber="19">Figure 10 (continued on next page).</emphasis>
<taxonomicName id="4C4D70B2FF8AFFBBFD91B1C8FD1DF909" authorityName="Otto &amp; Hill" authorityYear="2019" box="[540,746,1711,1736]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF8AFFBBFD91B1C8FD1DF909" box="[540,746,1711,1736]" italics="true" pageId="18" pageNumber="19">Maratus banyowla</emphasis>
</taxonomicName>
in their complex microhabitat at Banyowla Regional Park.
<emphasis id="B939D723FF8AFFBBFA00B1D7FA4BF909" bold="true" box="[1421,1468,1712,1736]" pageId="18" pageNumber="19">1-5,</emphasis>
Adult males. Photographs © by Kerry Stuart, used with permission.
</paragraph>
</caption>
<caption id="DF325BB9FF8BFFBAFFEDB047FA9CF857" ID-DOI="http://doi.org/10.5281/zenodo.3732695" ID-Zenodo-Dep="3732695" httpUri="https://zenodo.org/record/3732695/files/figure.png" pageId="19" pageNumber="20" startId="19.[96,176,1824,1848]" targetBox="[96,1536,165,1815]" targetPageId="19">
<paragraph id="8BF20B31FF8BFFBAFFEDB047FA9CF857" blockId="19.[96,1536,1784,1943]" pageId="19" pageNumber="20">
<emphasis id="B939D723FF8BFFBAFFEDB047FC5FF8F9" bold="true" box="[96,936,1824,1848]" pageId="19" pageNumber="20">Figure 10 (continued from previous page, continued on next page).</emphasis>
<taxonomicName id="4C4D70B2FF8BFFBAFC23B078FB88F8F9" authorityName="Otto &amp; Hill" authorityYear="2019" box="[942,1151,1823,1848]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF8BFFBAFC23B078FB88F8F9" box="[942,1151,1823,1848]" italics="true" pageId="19" pageNumber="20">Maratus banyowla</emphasis>
</taxonomicName>
in their complex microhabitat at Banyowla Regional Park.
<emphasis id="B939D723FF8BFFBAFE04B058FE13F896" bold="true" box="[393,484,1855,1879]" pageId="19" pageNumber="20">6-7, 16,</emphasis>
Adult males.
<emphasis id="B939D723FF8BFFBAFD03B058FD0EF896" bold="true" box="[654,761,1855,1879]" pageId="19" pageNumber="20">8-10, 15,</emphasis>
Males advertising or displaying with extended legs III.
<emphasis id="B939D723FF8BFFBAFAF4B058FA3FF896" bold="true" box="[1401,1480,1855,1879]" pageId="19" pageNumber="20">11-14,</emphasis>
Two composite images (11-12, 13-14) showing a male (at left) displaying to a nearby female. Like
<taxonomicName id="4C4D70B2FF8BFFBAFB0BB03AFAD6F8B7" authorityName="Otto &amp; Hill" authorityYear="2015" box="[1158,1313,1885,1910]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B939D723FF8BFFBAFB0BB03AFAD6F8B7" box="[1158,1313,1885,1910]" italics="true" pageId="19" pageNumber="20">M. personatus</emphasis>
</taxonomicName>
, we suspect that
<taxonomicName id="4C4D70B2FF8BFFBAFA6EB03AFF3CF857" authorityName="Otto &amp; Hill" authorityYear="2019" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF8BFFBAFA6EB03AFF3CF857" italics="true" pageId="19" pageNumber="20">M. banyowla</emphasis>
</taxonomicName>
do not raise their opisthosoma during courtship. Photographs © by Kerry Stuart, used with permission.
</paragraph>
</caption>
<caption id="DF325BB9FF8CFFBDFFEDB410FB80FC6E" ID-DOI="http://doi.org/10.5281/zenodo.3732697" ID-Zenodo-Dep="3732697" httpUri="https://zenodo.org/record/3732697/files/figure.png" pageId="20" pageNumber="21" startId="20.[96,176,887,911]" targetBox="[96,1537,165,876]" targetPageId="20">
<paragraph id="8BF20B31FF8CFFBDFFEDB410FB80FC6E" blockId="20.[96,1536,846,943]" pageId="20" pageNumber="21">
<emphasis id="B939D723FF8CFFBDFFEDB410FD83FC4E" bold="true" box="[96,628,887,911]" pageId="20" pageNumber="21">Figure 10 (continued from previous page).</emphasis>
<taxonomicName id="4C4D70B2FF8CFFBDFD0BB411FCA0FC4E" authorityName="Otto &amp; Hill" authorityYear="2019" box="[646,855,886,911]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF8CFFBDFD0BB411FCA0FC4E" box="[646,855,886,911]" italics="true" pageId="20" pageNumber="21">Maratus banyowla</emphasis>
</taxonomicName>
in their complex microhabitat at Banyowla Regional Park.
<emphasis id="B939D723FF8CFFBDFFEDB4F1FF59FC6F" bold="true" box="[96,174,918,942]" pageId="20" pageNumber="21">17-19,</emphasis>
Adult males.
<emphasis id="B939D723FF8CFFBDFEC0B4F1FE6CFC6F" bold="true" box="[333,411,918,942]" pageId="20" pageNumber="21">20-22,</emphasis>
Females. Photographs © by Kerry Stuart, used with permission.
</paragraph>
</caption>
<paragraph id="8BF20B31FF8CFFBDFFEDB4BCFA79FB51" blockId="20.[96,1535,987,1168]" pageId="20" pageNumber="21">
<emphasis id="B939D723FF8CFFBDFFEDB4BCFE25FC38" box="[96,466,987,1017]" italics="true" pageId="20" pageNumber="21">Host to endoparasitoid flies</emphasis>
. One female
<taxonomicName id="4C4D70B2FF8CFFBDFD1FB4BCFC7BFC38" authorityName="Otto &amp; Hill" authorityYear="2019" box="[658,908,987,1017]" class="Arachnida" family="Salticidae" genus="Maratus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="banyowla">
<emphasis id="B939D723FF8CFFBDFD1FB4BCFC7BFC38" box="[658,908,987,1017]" italics="true" pageId="20" pageNumber="21">Maratus banyowla</emphasis>
</taxonomicName>
(
<specimenCount id="9D4BC0B8FF8CFFBDFC2FB4B9FC4DFC3B" box="[930,954,990,1018]" pageId="20" pageNumber="21"></specimenCount>
#5) was collected as a penultimate at the
<typeStatus id="54F6B593FF8CFFBDFFEDB364FF6BFBDE" box="[96,156,1027,1055]" pageId="20" pageNumber="21">type</typeStatus>
locality and reared in the laboratory. This spider was photographed on
<date id="FFF32DF1FF8CFFBDFBFFB365FAE8FBE1" box="[1138,1311,1026,1056]" pageId="20" pageNumber="21" value="2019-09-26">26 SEP 2019</date>
(
<figureCitation id="137617B4FF8CFFBDFABDB365FA4FFBE1" box="[1328,1464,1026,1056]" captionStart="Figure 11" captionStartId="20.[96,175,1927,1951]" captionTargetBox="[96,1536,1494,1917]" captionTargetId="figure@20.[565,1120,1494,1917]" captionTargetPageId="20" captionText="Figure 11 (continued on next page). 1-2, Penultimate female Maratus banyowla collected at the type locality. 3, Remains of this spider after it was devoured by a larval acrocerid." figureDoi="http://doi.org/10.5281/zenodo.3732699" httpUri="https://zenodo.org/record/3732699/files/figure.png" pageId="20" pageNumber="21">Figure 11</figureCitation>
:1-2), and was enclosed in what appeared to be a moulting sac several days later. On
<date id="FFF32DF1FF8CFFBDFB26B34FFAA7FB84" box="[1195,1360,1064,1093]" pageId="20" pageNumber="21" value="2019-10-08">8 OCT 2019</date>
this sac was opened to reveal the remains of this female (
<figureCitation id="137617B4FF8CFFBDFD37B32AFCB5FBAA" box="[698,834,1101,1131]" captionStart="Figure 11" captionStartId="20.[96,175,1927,1951]" captionTargetBox="[96,1536,1494,1917]" captionTargetId="figure@20.[565,1120,1494,1917]" captionTargetPageId="20" captionText="Figure 11 (continued on next page). 1-2, Penultimate female Maratus banyowla collected at the type locality. 3, Remains of this spider after it was devoured by a larval acrocerid." figureDoi="http://doi.org/10.5281/zenodo.3732699" httpUri="https://zenodo.org/record/3732699/files/figure.png" pageId="20" pageNumber="21">Figure 11</figureCitation>
: 3-4) and the pupa of an acrocerid fly (
<figureCitation id="137617B4FF8CFFBDFAC3B32AFA21FBAA" box="[1358,1494,1101,1131]" captionStart="Figure 11" captionStartId="20.[96,175,1927,1951]" captionTargetBox="[96,1536,1494,1917]" captionTargetId="figure@20.[565,1120,1494,1917]" captionTargetPageId="20" captionText="Figure 11 (continued on next page). 1-2, Penultimate female Maratus banyowla collected at the type locality. 3, Remains of this spider after it was devoured by a larval acrocerid." figureDoi="http://doi.org/10.5281/zenodo.3732699" httpUri="https://zenodo.org/record/3732699/files/figure.png" pageId="20" pageNumber="21">Figure 11</figureCitation>
: 5- 6). By
<date id="FFF32DF1FF8CFFBDFF30B314FE87FB51" box="[189,368,1139,1168]" pageId="20" pageNumber="21" value="2019-10-15">15 OCT 2019</date>
the mature acrocerid fly (
<taxonomicName id="4C4D70B2FF8CFFBDFD43B314FCCCFB51" box="[718,827,1139,1168]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
:
<taxonomicName id="4C4D70B2FF8CFFBDFCCBB314FC19FB51" box="[838,1006,1139,1168]" class="Insecta" family="Acroceridae" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Acroceridae</taxonomicName>
) had emerged (
<figureCitation id="137617B4FF8CFFBDFB4DB314FAB0FB51" box="[1216,1351,1139,1168]" captionStart="Figure 11" captionStartId="20.[96,175,1927,1951]" captionTargetBox="[96,1536,1494,1917]" captionTargetId="figure@20.[565,1120,1494,1917]" captionTargetPageId="20" captionText="Figure 11 (continued on next page). 1-2, Penultimate female Maratus banyowla collected at the type locality. 3, Remains of this spider after it was devoured by a larval acrocerid." figureDoi="http://doi.org/10.5281/zenodo.3732699" httpUri="https://zenodo.org/record/3732699/files/figure.png" pageId="20" pageNumber="21">Figure 11</figureCitation>
:7-8).
</paragraph>
<paragraph id="8BF20B31FF8CFFBDFFEDB3D9FBC0FA57" blockId="20.[96,1534,1214,1430]" pageId="20" pageNumber="21">
Acrocerids are commonly known as spider flies, small-headed flies, or hunch-backed flies. Almost all known acrocerid larvae are endoparasitoids of spiders (
<bibRefCitation id="EFDC76C0FF8CFFBDFCF9B384FBB9FAC1" author="Schlinger, E. I." box="[884,1102,1251,1280]" editor="W. Nentwig" journalOrPublisher="Berlin, Springer Verlag" pageId="20" pageNumber="21" pagination="319 - 327" refId="ref2689" refString="Schlinger, E. I. 1987. The biology of Acroceridae (Diptera): true endoparasitoids of spiders. In W. Nentwig, ed., Ecophysiology of spiders. Berlin, Springer Verlag: 319 - 327." title="The biology of Acroceridae (Diptera): true endoparasitoids of spiders" type="book chapter" volumeTitle="Ecophysiology of spiders" year="1987">Schlinger 1987</bibRefCitation>
;
<bibRefCitation id="EFDC76C0FF8CFFBDFBD1B384FAB6FAC1" author="Winterton, S. L." box="[1116,1345,1251,1280]" journalOrPublisher="ZooKeys" pageId="20" pageNumber="21" pagination="7 - 75" part="172" refId="ref2731" refString="Winterton, S. L. 2012. Review of the Australasian spider flies (Diptera, Acroceridae) with a revision of Panops Lamarck. ZooKeys 172: 7 - 75." title="Review of the Australasian spider flies (Diptera, Acroceridae) with a revision of Panops Lamarck" type="journal article" year="2012">Winterton 2012</bibRefCitation>
;
<bibRefCitation id="EFDC76C0FF8CFFBDFAC2B384FEABFAE7" author="Winterton, S. L. &amp; D. A. Barraclough" editor="A. H. Kirk-Spriggs &amp; B. J. Sinclair" journalOrPublisher="South African National Biodiversity Institute. Pretoria. Suricata" pageId="20" pageNumber="21" pagination="981 - 993" part="5" refId="ref2764" refString="Winterton, S. L. and D. A. Barraclough. 2017. Acroceridae (Small-headed Flies or Spider Flies). In A. H. Kirk-Spriggs and B. J. Sinclair, eds., Manual of Afrotropical Diptera, Volume 2. Nematocerous Diptera and lower Brachycera. South African National Biodiversity Institute. Pretoria. Suricata 5: 981 - 993." title="Acroceridae (Small-headed Flies or Spider Flies)" type="journal article" volumeTitle="Manual of Afrotropical Diptera, Volume 2. Nematocerous Diptera and lower Brachycera" year="2017">Winterton &amp; Barraclough 2017</bibRefCitation>
). Adult acrocerids deposit numbers of minute eggs on plants and the emergent larvae burrow into spiders where they grow and moult but do not kill their host until late in their development. Prior to this time they may not be detectable. Several North American species are endoparasitoids of the salticids
<taxonomicName id="4C4D70B2FF8CFFBDFF55B21FFE73FA57" baseAuthorityName="Hentz" baseAuthorityYear="1845" box="[216,388,1400,1430]" class="Arachnida" family="Salticidae" genus="Eris" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="militaris">
<emphasis id="B939D723FF8CFFBDFF55B21FFE73FA57" box="[216,388,1400,1430]" italics="true" pageId="20" pageNumber="21">Eris militaris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D70B2FF8CFFBDFE48B21FFBC4FA57" authority="(Larrivee &amp; Borkent 2009)" baseAuthorityName="Larrivee &amp; Borkent" baseAuthorityYear="2009" box="[453,1075,1400,1430]" class="Arachnida" family="Salticidae" genus="Pelegrina" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="proterva">
<emphasis id="B939D723FF8CFFBDFE48B21FFD48FA57" box="[453,703,1400,1430]" italics="true" pageId="20" pageNumber="21">Pelegrina proterva</emphasis>
(
<bibRefCitation id="EFDC76C0FF8CFFBDFD5DB21EFBDEFA57" author="Larrivee, M. &amp; C. J. Borkent" box="[720,1065,1401,1430]" journalOrPublisher="The Journal of Arachnology" pageId="20" pageNumber="21" pagination="241 - 242" part="37" refId="ref2565" refString="Larrivee, M. and C. J. Borkent. 2009. New spider host associations for three acrocerid fly species (Diptera, Acroceridae). The Journal of Arachnology 37: 241 - 242." title="New spider host associations for three acrocerid fly species (Diptera, Acroceridae)" type="journal article" year="2009">Larrivée &amp; Borkent 2009</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>