174 lines
19 KiB
XML
174 lines
19 KiB
XML
<document id="C58EA416FC16FF9864E0FA82A8DF00A5" ID-DOI="10.11646/zootaxa.2618.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5304087" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629699783393" checkinUser="felipe" docAuthor="Oláh, János & Johanson, Kjell Arne" docDate="2010" docId="03E487BBFFC32B1EFF3CFF0C3C241DFD" docLanguage="en" docName="zt02618p035.pdf" docOrigin="Zootaxa 2618 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2618.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nyctiophylax (Nyctiophylax) tapan Oláh & Johanson 2010, new species" docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="FFDDFFC3FFD32B0FFFABFF9B39571F41" masterDocTitle="Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia" masterLastPageNumber="35" masterPageNumber="1" pageNumber="17" updateTime="1699097725960" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="393A185A7F4867D83912CC4268BFE6A8">
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<mods:title id="52EAF3BD46323E97565F657B683154E1">Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name id="F505B300FAE554F8962440CA3CBF151F" type="personal">
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<mods:roleTerm id="37DEE2D53EFFB3F24502830C89E5DC05">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5E70AAF6526C9E29B96BE8CC4C0AE235">Oláh, János</mods:namePart>
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<mods:affiliation id="5123188CC5A49AFE23B54A568E49A6E1">Szent István University, Gödöll, Centre of Environmental Health. Residence postal address: Tarján u. 28, H- 4032 Debrecen, Hungary. E-mail: profolah @ gmail. com</mods:affiliation>
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<mods:roleTerm id="0723C9874D2F7DFE35D150BFA91EC78E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1C3937A3C715B197316427738576C757">Johanson, Kjell Arne</mods:namePart>
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<mods:affiliation id="4937DC8BC76C5B2AF6C9DABA9494AC31">Swedish Museum of Natural History, Entomology Department, Box 50007, S- 10405 Stockholm, Sweden.</mods:affiliation>
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<mods:title id="99EDE25367B669202995482C7F9DEA70">Zootaxa</mods:title>
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<mods:part id="EF489AFBB0BC4DC132BCB93865A7AE9D">
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<mods:date id="5096F04EFC82949825FBD74E827D7C82">2010</mods:date>
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<mods:number id="33D3842C887FF60C6556FA8E8669A7F1">2010-09-20</mods:number>
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<mods:detail id="B1508A9CF07DFC2419302ED3FA0DE5A8" type="volume">
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<mods:number id="07404E4CD346D7E0D37DCF0BF73C2999">2618</mods:number>
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<mods:url id="070237BA2AC03EABED803DD236E26FF0">https://biotaxa.org/Zootaxa/article/view/zootaxa.2618.1.1</mods:url>
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</mods:location>
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<mods:classification id="5EA01279C8E1163DB1EDA382AD33C2E5">journal article</mods:classification>
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<mods:identifier id="68AEA2EFE5661FAE2D557CCBD420556E" type="DOI">10.11646/zootaxa.2618.1.1</mods:identifier>
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<mods:identifier id="1B818CA89615BA832BA8248D007D53DE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E487BBFFC32B1EFF3CFF0C3C241DFD" ID-DOI="http://doi.org/10.5281/zenodo.5311084" ID-Zenodo-Dep="5311084" LSID="urn:lsid:plazi:treatment:03E487BBFFC32B1EFF3CFF0C3C241DFD" httpUri="http://treatment.plazi.org/id/03E487BBFFC32B1EFF3CFF0C3C241DFD" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection id="C3576526FFC32B1FFF3CFF0C3B8C1FF0" box="[151,731,151,177]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8BF236ADFFC32B1FFF3CFF0C3B8C1FF0" blockId="16.[151,731,151,212]" box="[151,731,151,177]" pageId="16" pageNumber="17">
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<heading id="D0BA81C1FFC32B1FFF3CFF0C3B8C1FF0" bold="true" box="[151,731,151,177]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<emphasis id="B939EABFFFC32B1FFF3CFF0C3B8C1FF0" bold="true" box="[151,731,151,177]" pageId="16" pageNumber="17">
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<taxonomicName id="4C4D4D2EFFC32B1FFF3CFF0C3B6B1FF0" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,572,151,177]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tapan" status="sp. nov." subGenus="Nyctiophylax">
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<emphasis id="B939EABFFFC32B1FFF3CFF0C38601FF0" bold="true" box="[151,311,151,177]" italics="true" pageId="16" pageNumber="17">Nyctiophylax</emphasis>
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(
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<emphasis id="B939EABFFFC32B1FFEECFF0C38B11FF0" bold="true" box="[327,486,151,177]" italics="true" pageId="16" pageNumber="17">Nyctiophylax</emphasis>
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)
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<emphasis id="B939EABFFFC32B1FFE53FF0C3B6B1FF0" bold="true" box="[504,572,151,177]" italics="true" pageId="16" pageNumber="17">tapan</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20A57C4FFC32B1FFDE0FF0C3B8C1FF0" box="[587,731,151,177]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576526FFC32B1EFF3CFF2139B21D66" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
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<paragraph id="8BF236ADFFC32B1FFF3CFF21384D1F95" blockId="16.[151,731,151,212]" box="[151,282,186,212]" pageId="16" pageNumber="17">
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<figureCitation id="13762A28FFC32B1FFF3CFF21384D1F95" box="[151,282,186,212]" captionStart="FIGURES 27–30" captionStartId="16.[151,270,1481,1505]" captionTargetBox="[177,1361,517,1394]" captionTargetId="figure-293@16.[151,1436,490,1459]" captionTargetPageId="16" captionText="FIGURES 27–30. Nyctiophylax (Nyctiophylax) tapan, new species, holotype. 27—genitalia, left lateral; 28—genitalia, dorsal; 29—right gonocoxite, ventral; 30—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5304115" httpUri="https://zenodo.org/record/5304115/files/figure.png" pageId="16" pageNumber="17">Figs 27–30</figureCitation>
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</paragraph>
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<paragraph id="8BF236ADFFC32B1FFF3CFE9E38FF1EF5" blockId="16.[151,1436,261,436]" pageId="16" pageNumber="17">
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The genitalia of this species of subgenus
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<taxonomicName id="4C4D4D2EFFC32B1FFD2BFE9E3A4C1E5F" authorityName="Brauer" authorityYear="1865" box="[640,795,261,286]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis id="B939EABFFFC32B1FFD2BFE9E3A4C1E5F" box="[640,795,261,286]" italics="true" pageId="16" pageNumber="17">Nyctiophylax</emphasis>
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</taxonomicName>
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resemble those of
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<taxonomicName id="4C4D4D2EFFC32B1FFC57FE9E3DE01E5F" authorityName="Malicky & Chantaramongkol" authorityYear="1993" box="[1020,1207,261,286]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="khaoyaiensis">
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<emphasis id="B939EABFFFC32B1FFC57FE9E3DE01E5F" box="[1020,1207,261,286]" italics="true" pageId="16" pageNumber="17">N. khaoyaiensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4D4D2EFFC32B1FFB59FE9E3C0C1E5F" authorityName="Malicky & Chantaramongkol" authorityYear="1993" box="[1266,1371,261,286]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="salma">
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<emphasis id="B939EABFFFC32B1FFB59FE9E3C0C1E5F" box="[1266,1371,261,286]" italics="true" pageId="16" pageNumber="17">N. salma</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="F35A763DFFC32B1FFF3CFEB138511E05" box="[151,262,298,324]" name="Thailand" pageId="16" pageNumber="17">Thailand</collectingCountry>
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.
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<taxonomicName id="4C4D4D2EFFC32B1FFEB9FEB138AA1E02" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[274,509,298,323]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tapan" status="sp. nov.">
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<emphasis id="B939EABFFFC32B1FFEB9FEB138AA1E02" box="[274,509,298,323]" italics="true" pageId="16" pageNumber="17">Nyctiophylax tapan</emphasis>
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</taxonomicName>
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,
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<emphasis id="B939EABFFFC32B1FFDA5FEB13BF51E05" bold="true" box="[526,674,298,324]" pageId="16" pageNumber="17">
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<taxonomicNameLabel id="A20A57C4FFC32B1FFDA5FEB13BF51E05" box="[526,674,298,324]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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is distinguished from the 2 similar species by the shape of the ventroapical keels of the gonopods and the ventral paraproctal plate of sternite IX; the shape of sternite IX in lateral view; and the presence of a pronounced antecostal suture running around the entire sternite IX except at the posterior margin.
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</paragraph>
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<caption id="DF326625FFC32B1FFF3CFA523ACB1941" ID-DOI="http://doi.org/10.5281/zenodo.5304115" ID-Zenodo-Dep="5304115" httpUri="https://zenodo.org/record/5304115/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,270,1481,1505]" targetBox="[177,1361,517,1394]" targetPageId="16">
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||
<paragraph id="8BF236ADFFC32B1FFF3CFA523ACB1941" blockId="16.[151,1436,1481,1536]" pageId="16" pageNumber="17">
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<emphasis id="B939EABFFFC32B1FFF3CFA5238091AA0" bold="true" box="[151,350,1481,1505]" pageId="16" pageNumber="17">FIGURES 27–30.</emphasis>
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<taxonomicName id="4C4D4D2EFFC32B1FFECEFA523B8A1AA1" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[357,733,1481,1504]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="tapan" status="sp. nov." subGenus="Nyctiophylax">
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<emphasis id="B939EABFFFC32B1FFECEFA523B8A1AA1" box="[357,733,1481,1504]" italics="true" pageId="16" pageNumber="17">Nyctiophylax (Nyctiophylax) tapan</emphasis>
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</taxonomicName>
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,
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<emphasis id="B939EABFFFC32B1FFD40FA523A391AA0" bold="true" box="[747,878,1481,1505]" pageId="16" pageNumber="17">
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<taxonomicNameLabel id="A20A57C4FFC32B1FFD40FA523A391AA0" box="[747,878,1481,1505]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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, holotype. 27—genitalia, left lateral; 28—genitalia, dorsal; 29—right gonocoxite, ventral; 30—phallic apparatus, left lateral.
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</paragraph>
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</caption>
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<paragraph id="8BF236ADFFC32B1FFF6EF9B538C3180F" blockId="16.[151,1436,1582,2019]" pageId="16" pageNumber="17">
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Male (in alcohol). Body uniformly brown. Maxillary palp formula II-I-IV-III-V, with segment III originating from mesal face before apex of segment II. Anterior tentorial pits located laterally, near eyes. Frontal and clypeal areas wide; frontogenal and clypeogenal areas narrow. Pair of large, triangular, frontal lateral compact setose warts filling dark frontal area, with white alveoli; warts almost tangential mesally; clypeal sclerites light brown, covered by large, single, clypeal diffuse setose wart; few anterior diffuse setae present on labrum. Forewing length 4.5 mm; discoidal cell and median cell closed; apical forks 2, 3, 4 and 5 present; anal veins
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<emphasis id="B939EABFFFC32B1FFE2BF89438F61869" box="[384,417,1807,1832]" italics="true" pageId="16" pageNumber="17">A1</emphasis>
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,
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<emphasis id="B939EABFFFC32B1FFE1AF89438851869" box="[433,466,1807,1832]" italics="true" pageId="16" pageNumber="17">A2</emphasis>
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, and
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<emphasis id="B939EABFFFC32B1FFDBDF8943B601869" box="[534,567,1807,1832]" italics="true" pageId="16" pageNumber="17">A3</emphasis>
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without loop. Hind wing discoidal cell closed, median cell open; apical forks 2 and 5 present.
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</paragraph>
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<paragraph id="8BF236ADFFC32B1FFF6EF8C238FE1832" blockId="16.[151,1436,1582,2019]" box="[197,425,1881,1907]" pageId="16" pageNumber="17">Spur formula 3,4,4.</paragraph>
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<paragraph id="8BF236ADFFC32B1EFF6EF8E539B21D66" blockId="16.[151,1436,1582,2019]" lastBlockId="17.[151,1436,152,700]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">Male genitalia. Sternite IX robust, sclerotized, quadrangular in lateral view; delimited by pronounced antecostal suture except on posterior margin; ventroapical keel developed into slightly S-shaped, tapering process in lateral view. Tergum IX membranous, with semisclerotized integument above sternite IX and behind paraproctal complex visible, confluent with paraprocts. Segment X membranous, elongate, nearly quadrangular in lateral view. Cerci elongate, club-shaped, originating from paraproctal bodies. Paraproctal complex with 2 pairs of paraproctal processes: dorsal paraproctal processes forming arching spines originating anteriorly before running dorsad and turning posterad; ventral paraproctal processes forming pair of large, horizontally elongated plates, each with capitate apex and short, dorsal process with setae on terminal area. Plates of paraproctal complex meeting ventrally, forming sclerous subphallic bridge and phallic guide. Gonopods elongate, almost parallel-sided, each with bifid apex in lateral view. Phallic apparatus located dorsally in genitalia, fixed, guided by paraproctal processes; phallotheca short, sclerotized, with apical rim and basidorsal plate. Phallobase with wide anterior foramen; membranous endotheca (or phallicata) with lesssclerotic lobe apicoventrally, spine-shaped in lateral view; pair of obscure phallotremal sclerites present inside.</paragraph>
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</subSubSection>
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||
<subSubSection id="C3576526FFC22B1EFF6EFDA93B1A1D33" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="8BF236ADFFC22B1EFF6EFDA93B1A1D33" blockId="17.[151,1436,152,700]" pageId="17" pageNumber="18">
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||
<materialsCitation id="3B253CF0FFC22B1EFF6EFDA93B1A1D33" collectingDate="2008-07-10" collectorName="J. Olah" country="Indonesia" elevation="1000" location="Tapan Road" municipality="Kerinci National Park" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
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<emphasis id="B939EABFFFC22B1EFF6EFDA9382D1D0D" bold="true" box="[197,378,562,588]" pageId="17" pageNumber="18">
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<typeStatus id="54F6880FFFC22B1EFF6EFDA938601D0D" box="[197,311,562,588]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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:
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<emphasis id="B939EABFFFC22B1EFE20FDA93B661D0D" bold="true" box="[395,561,562,588]" pageId="17" pageNumber="18">
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||
<collectingCountry id="F35A763DFFC22B1EFE20FDA93B661D0D" box="[395,561,562,588]" name="Indonesia" pageId="17" pageNumber="18">INDONESIA</collectingCountry>
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</emphasis>
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||
(
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<collectingRegion id="4989F84FFFC22B1EFDEBFDA93BFE1D0D" box="[576,681,562,588]" country="Indonesia" name="Sumatera Utara" pageId="17" pageNumber="18">Sumatra</collectingRegion>
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):
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<collectingMunicipality id="6B96ACD7FFC22B1EFD16FDA93A951D0D" box="[701,962,562,588]" pageId="17" pageNumber="18">Kerinci National Park</collectingMunicipality>
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||
,
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<location id="8E926076FFC22B1EFC65FDA93D091D0D" LSID="urn:lsid:plazi:treatment:03E487BBFFC32B1EFF3CFF0C3C241DFD:8E926076FFC22B1EFC65FDA93D091D0D" box="[974,1118,562,588]" country="Indonesia" municipality="Kerinci National Park" name="Tapan Road" pageId="17" pageNumber="18" stateProvince="Sumatera Utara">Tapan Road</location>
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||
,
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<quantity id="4CB59B48FFC22B1EFBC1FDA93D921D0D" box="[1130,1221,562,588]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="1000.0">
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||
<elevation id="0060D19EFFC22B1EFBC1FDA93D921D0D" box="[1130,1221,562,588]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="1000.0">1000 m</elevation>
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</quantity>
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||
,
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<date id="FFF3106DFFC22B1EFB79FDA93C0E1D0D" box="[1234,1369,562,588]" pageId="17" pageNumber="18" value="2008-07-10">
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||
<collectingDate id="EFB7E985FFC22B1EFB79FDA93C0E1D0D" box="[1234,1369,562,588]" pageId="17" pageNumber="18" value="2008-07-10">10.vii.2008</collectingDate>
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</date>
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,
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<collectingMethod id="520C4EBAFFC22B1EFACDFDA939921D33" pageId="17" pageNumber="18">light trap</collectingMethod>
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[
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<collectorName id="26B8537BFFC22B1EFF7FFDC3387C1D33" box="[212,299,600,626]" pageId="17" pageNumber="18">J. Oláh</collectorName>
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, Jr.]—(OPC, in alcohol).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576526FFC22B1EFF6EFDE63B321DD6" box="[197,613,637,663]" pageId="17" pageNumber="18" type="distribution">
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||
<paragraph id="8BF236ADFFC22B1EFF6EFDE63B321DD6" blockId="17.[151,1436,152,700]" box="[197,613,637,663]" pageId="17" pageNumber="18">
|
||
<emphasis id="B939EABFFFC22B1EFF6EFDE638301DD6" bold="true" box="[197,359,637,663]" pageId="17" pageNumber="18">Distribution:</emphasis>
|
||
<collectingCountry id="F35A763DFFC22B1EFEC4FDE638B61DD6" box="[367,481,637,663]" name="Indonesia" pageId="17" pageNumber="18">Indonesia</collectingCountry>
|
||
(
|
||
<collectingRegion id="4989F84FFFC22B1EFE5AFDE63B0E1DD6" box="[497,601,637,663]" country="Indonesia" name="Sumatera Utara" pageId="17" pageNumber="18">Sumatra</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576526FFC22B1EFF6EFD393C241DFD" box="[197,1395,674,700]" pageId="17" pageNumber="18" type="etymology">
|
||
<paragraph id="8BF236ADFFC22B1EFF6EFD393C241DFD" blockId="17.[151,1436,152,700]" box="[197,1395,674,700]" pageId="17" pageNumber="18">
|
||
<emphasis id="B939EABFFFC22B1EFF6EFD3938021DFD" bold="true" box="[197,341,674,700]" pageId="17" pageNumber="18">Etymology:</emphasis>
|
||
<emphasis id="B939EABFFFC22B1EFEF6FD3938F21DFA" box="[349,421,674,699]" italics="true" pageId="17" pageNumber="18">Tapan</emphasis>
|
||
, named after the
|
||
<typeStatus id="54F6880FFFC22B1EFDDAFD393BF41DFD" box="[625,675,674,700]" pageId="17" pageNumber="18">type</typeStatus>
|
||
locality of the species, Tapan Road in Kerinci National Park.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |