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<document id="B397CA546404A47E9895489836F02133" ID-CLB-Dataset="2910" ID-DOI="10.11646/zootaxa.5082.3.2" ID-GBIF-Dataset="3997d831-ae29-42d0-ba99-cd20cc8b7fc3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5788224" ID-ZooBank="EFB0E4A5-6B60-4B39-A651-1C6470FD7222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639730480321" checkinUser="plazi" docAuthor="Li, Yan-Da, Tihelka, Erik, Dahan, Loïc, Huang, Di-Ying &amp; Cai, Chen-Yang" docDate="2021" docId="03D787A2FFE4EE70FF52F986891B6D77" docLanguage="en" docName="zootaxa.5082.3.2.pdf" docOrigin="Zootaxa 5082 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaenosodendron explanatum Li &amp; Cai 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="230" masterDocId="FFEEFFDAFFE0EE77FFC5FFAA8B796C12" masterDocTitle="On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)" masterLastPageNumber="244" masterPageNumber="223" pageNumber="227" updateTime="1699472730423" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6FF2BC0BC5B423FCCD531C87AB1E3D16">On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)</mods:title>
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<mods:namePart id="CB4655F00EB3548E7645787A28025792">Li, Yan-Da</mods:namePart>
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<mods:namePart id="5234E6A1E3679A071E26F5BCF7811000">Tihelka, Erik</mods:namePart>
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<mods:affiliation id="363720F98525D6FFB46649044F78C8CD">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK &amp; wn 20250 @ bristol. ac. uk; https: // orcid. org / 0000 - 0002 - 5048 - 5355</mods:affiliation>
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<mods:namePart id="24231380CA05126D7B64AA6B0F77F895">Dahan, Loïc</mods:namePart>
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<mods:affiliation id="D33107847BEB2A55AB804558CB8A436B">O. D. Taxonomy and Phylogeny - Entomology, Royal Belgian Institute of Natural Sciences, Vautierstreet 29, B- 1000 Brussels, Belgium ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X &amp; loicdahan @ hotmail. com; https: // orcid. org / 0000 - 0001 - 5680 - 0304</mods:affiliation>
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<mods:namePart id="6A0F53CDAF200FE4CBC61495F4182663">Huang, Di-Ying</mods:namePart>
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<mods:affiliation id="15D3E7071C16B7C1B30DA5A6367B98E0">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867</mods:affiliation>
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<mods:namePart id="8E4EAC6B0046340134B671A3933C7CF2">Cai, Chen-Yang</mods:namePart>
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<mods:affiliation id="9D56AA60CD5AF975776412AF44B086D1">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK &amp; cycai @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 9283 - 8323</mods:affiliation>
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<treatment id="03D787A2FFE4EE70FF52F986891B6D77" ID-DOI="http://doi.org/10.5281/zenodo.5788230" ID-GBIF-Taxon="191654784" ID-Zenodo-Dep="5788230" LSID="urn:lsid:plazi:treatment:03D787A2FFE4EE70FF52F986891B6D77" httpUri="http://treatment.plazi.org/id/03D787A2FFE4EE70FF52F986891B6D77" lastPageId="7" lastPageNumber="230" pageId="4" pageNumber="227">
<subSubSection id="C364653FFFE4EE73FF52F98689866A55" box="[151,767,1580,1607]" pageId="4" pageNumber="227" type="nomenclature">
<paragraph id="8BC136B4FFE4EE73FF52F98689866A55" blockId="4.[151,767,1580,1644]" box="[151,767,1580,1607]" pageId="4" pageNumber="227">
<heading id="D08981D8FFE4EE73FF52F98689866A55" bold="true" box="[151,767,1580,1607]" fontSize="11" level="1" pageId="4" pageNumber="227" reason="1">
<emphasis id="B90AEAA6FFE4EE73FF52F98689866A55" bold="true" box="[151,767,1580,1607]" pageId="4" pageNumber="227">
<taxonomicName id="4C7E4D37FFE4EE73FF52F98689E36A55" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2021" box="[151,666,1580,1607]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="227" phylum="Arthropoda" rank="species" species="explanatum" status="sp. nov.">
<emphasis id="B90AEAA6FFE4EE73FF52F986895D6A55" bold="true" box="[151,548,1580,1607]" italics="true" pageId="4" pageNumber="227">Archaenosodendron explanatum</emphasis>
Li &amp; Cai
</taxonomicName>
<taxonomicNameLabel id="A23957DDFFE4EE73FD67F98789866A55" box="[674,767,1581,1607]" pageId="4" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C364653FFFE4EE73FF52F9FB8A726A7E" box="[151,267,1617,1644]" pageId="4" pageNumber="227" type="description">
<paragraph id="8BC136B4FFE4EE73FF52F9FB8A726A7E" blockId="4.[151,767,1580,1644]" box="[151,267,1617,1644]" pageId="4" pageNumber="227">
(
<figureCitation id="13452A31FFE4EE73FF5AF9FB8A7A6A7E" box="[159,259,1617,1644]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="6.[151,250,1483,1507]" captionStartId-1="7.[151,250,1784,1808]" captionStartId-2="8.[151,250,1327,1351]" captionStartId-3="9.[151,250,1847,1871]" captionTargetBox-0="[151,1436,181,1459]" captionTargetBox-1="[151,1436,381,1761]" captionTargetBox-2="[151,1436,181,1303]" captionTargetBox-3="[151,1436,525,1824]" captionTargetId-0="figure-22@6.[151,1436,181,1459]" captionTargetId-1="figure-85@7.[151,1436,381,1761]" captionTargetId-2="figure-22@8.[151,1436,181,1303]" captionTargetId-3="figure-162@9.[151,1436,525,1824]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIGURE 5. General habitus of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 1 mm." captionText-1="FIGURE 6. General habitus of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613, under widefield fluorescence. A, Dorsal view. B, Ventral view. Scale bars: 1 mm." captionText-2="FIGURE 7. X-ray microtomographic reconstruction of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613. A, Dorsal view. B, Ventral view. C, Lateral view. Scale bar: 1 mm." captionText-3="FIGURE 8. Details of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Head, lateral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum. EG, Fore legs. H, Mid leg. I, Hind leg. Abbreviations: an1, antennomere 1; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; v1,2, ventrites 1,2. Scale bars: 200 μm." figureDoi-0="http://doi.org/10.5281/zenodo.5788241" figureDoi-1="http://doi.org/10.5281/zenodo.5788243" figureDoi-2="http://doi.org/10.5281/zenodo.5788249" figureDoi-3="http://doi.org/10.5281/zenodo.5788251" httpUri-0="https://zenodo.org/record/5788241/files/figure.png" httpUri-1="https://zenodo.org/record/5788243/files/figure.png" httpUri-2="https://zenodo.org/record/5788249/files/figure.png" httpUri-3="https://zenodo.org/record/5788251/files/figure.png" pageId="4" pageNumber="227">Figs 58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFE4EE73FF52F9338F3A6AC5" pageId="4" pageNumber="227" type="materials_examined">
<paragraph id="8BC136B4FFE4EE73FF52F9338F3A6AC5" blockId="4.[151,1437,1689,2039]" pageId="4" pageNumber="227">
<emphasis id="B90AEAA6FFE4EE73FF52F9338A7F6AA1" bold="true" box="[151,262,1689,1715]" pageId="4" pageNumber="227">Material.</emphasis>
<materialsCitation id="3B163CE9FFE4EE73FECAF9338F3A6AC5" ID-GBIF-Occurrence="3419363303" collectionCode="NIGP" pageId="4" pageNumber="227" specimenCode="NIGP177613" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C58816FFE4EE73FECAF9338A036AA1" box="[271,378,1689,1715]" pageId="4" pageNumber="227" type="holotype">Holotype</typeStatus>
,
<specimenCode id="DBD89ECFFFE4EE73FE43F93389666AA1" box="[390,543,1689,1715]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="4" pageNumber="227">NIGP177613</specimenCode>
. The specimen seems to have been damaged before entrapment in amber, with many appendages (mouthparts, antennae, and tarsi) detached and head outstretched.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFE4EE73FF02F94B8F9B6B51" pageId="4" pageNumber="227" type="etymology">
<paragraph id="8BC136B4FFE4EE73FF02F94B882D6AE9" blockId="4.[151,1437,1689,2039]" box="[199,852,1761,1787]" pageId="4" pageNumber="227">
<emphasis id="B90AEAA6FFE4EE73FF02F94B8A376AE9" bold="true" box="[199,334,1761,1787]" pageId="4" pageNumber="227">Etymology.</emphasis>
The specific name refers to its flattened body.
</paragraph>
<paragraph id="8BC136B4FFE4EE73FF02F8AF8F9B6B51" blockId="4.[151,1437,1689,2039]" pageId="4" pageNumber="227">
<emphasis id="B90AEAA6FFE4EE73FF02F8AF8AB66B0D" bold="true" box="[199,463,1797,1823]" pageId="4" pageNumber="227">Locality and horizon.</emphasis>
Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="49BAF856FFE4EE73FF52F8838A536B51" box="[151,298,1833,1859]" country="Myanmar" name="Kachin" pageId="4" pageNumber="227">Kachin State</collectingRegion>
,
<collectingCountry id="F3697624FFE4EE73FEF0F8838ADF6B51" box="[309,422,1833,1859]" name="Myanmar" pageId="4" pageNumber="227">Myanmar</collectingCountry>
; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFE4EE73FF02F8E78A3E6BE5" pageId="4" pageNumber="227" type="diagnosis">
<paragraph id="8BC136B4FFE4EE73FF02F8E78A3E6BE5" blockId="4.[151,1437,1689,2039]" pageId="4" pageNumber="227">
<emphasis id="B90AEAA6FFE4EE73FF02F8E78A3B6B75" bold="true" box="[199,322,1869,1895]" pageId="4" pageNumber="227">Diagnosis.</emphasis>
Body flattened and relatively elongate (
<figureCitation id="13452A31FFE4EE73FCCAF8E7882C6B75" box="[783,853,1869,1895]" captionStart="FIGURE 7" captionStartId="8.[151,250,1327,1351]" captionTargetBox="[151,1436,181,1303]" captionTargetId="figure-22@8.[151,1436,181,1303]" captionTargetPageId="8" captionText="FIGURE 7. X-ray microtomographic reconstruction of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613. A, Dorsal view. B, Ventral view. C, Lateral view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5788249" httpUri="https://zenodo.org/record/5788249/files/figure.png" pageId="4" pageNumber="227">Fig. 7</figureCitation>
). Lateral regions of mentum with somewhat longitudinally arranged larger punctures (
<figureCitation id="13452A31FFE4EE73FDF6F8DB89F56B99" box="[563,652,1905,1931]" captionStart="FIGURE 8" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,525,1824]" captionTargetId="figure-162@9.[151,1436,525,1824]" captionTargetPageId="9" captionText="FIGURE 8. Details of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Head, lateral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum. EG, Fore legs. H, Mid leg. I, Hind leg. Abbreviations: an1, antennomere 1; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; v1,2, ventrites 1,2. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788251" httpUri="https://zenodo.org/record/5788251/files/figure.png" pageId="4" pageNumber="227">Fig. 8B</figureCitation>
). Anterior angles of mentum produced anteriorly (
<figureCitation id="13452A31FFE4EE73FB0DF8DB8E586B99" box="[1224,1313,1905,1931]" captionStart="FIGURE 8" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,525,1824]" captionTargetId="figure-162@9.[151,1436,525,1824]" captionTargetPageId="9" captionText="FIGURE 8. Details of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Head, lateral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum. EG, Fore legs. H, Mid leg. I, Hind leg. Abbreviations: an1, antennomere 1; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; v1,2, ventrites 1,2. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788251" httpUri="https://zenodo.org/record/5788251/files/figure.png" pageId="4" pageNumber="227">Fig. 8B</figureCitation>
). Antennal grooves on prosternum longer than half-length of prosternum in front of coxae (
<figureCitation id="13452A31FFE4EE73FBE8F83F8FF06BBD" box="[1069,1161,1941,1967]" captionStart="FIGURE 8" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,525,1824]" captionTargetId="figure-162@9.[151,1436,525,1824]" captionTargetPageId="9" captionText="FIGURE 8. Details of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Head, lateral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum. EG, Fore legs. H, Mid leg. I, Hind leg. Abbreviations: an1, antennomere 1; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; v1,2, ventrites 1,2. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788251" httpUri="https://zenodo.org/record/5788251/files/figure.png" pageId="4" pageNumber="227">Fig. 8D</figureCitation>
). Outer edge of protibia with about 2123 denticles/stout setae (
<figureCitation id="13452A31FFE4EE73FD9FF81389AC6BC1" box="[602,725,1977,2003]" captionStart="FIGURE 8" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,525,1824]" captionTargetId="figure-162@9.[151,1436,525,1824]" captionTargetPageId="9" captionText="FIGURE 8. Details of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Head, lateral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum. EG, Fore legs. H, Mid leg. I, Hind leg. Abbreviations: an1, antennomere 1; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; v1,2, ventrites 1,2. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788251" httpUri="https://zenodo.org/record/5788251/files/figure.png" pageId="4" pageNumber="227">Fig. 8EG</figureCitation>
). Medial and lateral portions of prosternum not separated by a ridge (
<figureCitation id="13452A31FFE4EE73FF25F8778A436BE5" box="[224,314,2013,2039]" captionStart="FIGURE 8" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,525,1824]" captionTargetId="figure-162@9.[151,1436,525,1824]" captionTargetPageId="9" captionText="FIGURE 8. Details of Archaenosodendron explanatum Li &amp; Cai sp. nov., holotype, NIGP177613, under confocal microscopy. A, Head, dorsal view. B, Head, ventral view. C, Head, lateral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum. EG, Fore legs. H, Mid leg. I, Hind leg. Abbreviations: an1, antennomere 1; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; v1,2, ventrites 1,2. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788251" httpUri="https://zenodo.org/record/5788251/files/figure.png" pageId="4" pageNumber="227">Fig. 8D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF01663CFFE5EE72FF52F94689BB6BCF" ID-DOI="http://doi.org/10.5281/zenodo.5788234" ID-Zenodo-Dep="5788234" httpUri="https://zenodo.org/record/5788234/files/figure.png" pageId="5" pageNumber="228" startId="5.[151,250,1772,1796]" targetBox="[151,1436,181,1748]" targetPageId="5">
<paragraph id="8BC136B4FFE5EE72FF52F94689BB6BCF" blockId="5.[151,1437,1772,2013]" pageId="5" pageNumber="228">
<emphasis id="B90AEAA6FFE5EE72FF52F9468A6D6B16" bold="true" box="[151,276,1772,1796]" pageId="5" pageNumber="228">FIGURE 4.</emphasis>
Details of
<taxonomicName id="4C7E4D37FFE5EE72FE40F94688F06B16" authority="(Deng et al., 2018)" authorityName="Li &amp; Tihelka &amp; Dahan &amp; Huang &amp; Cai" authorityYear="2021" baseAuthorityName="Li &amp; Tihelka &amp; Dahan &amp; Huang &amp; Cai" baseAuthorityYear="2018" box="[389,905,1772,1796]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="228" phylum="Arthropoda" rank="species" species="cretaceum" status="comb. nov.">
<emphasis id="B90AEAA6FFE5EE72FE40F94689B86B16" box="[389,705,1772,1796]" italics="true" pageId="5" pageNumber="228">Archaenosodendron cretaceum</emphasis>
(Deng
<emphasis id="B90AEAA6FFE5EE72FCCBF947883B6B16" box="[782,834,1772,1796]" italics="true" pageId="5" pageNumber="228">et al.</emphasis>
, 2018)
</taxonomicName>
<emphasis id="B90AEAA6FFE5EE72FC55F9468F7C6B16" bold="true" box="[912,1029,1772,1796]" pageId="5" pageNumber="228">
<taxonomicNameLabel id="A23957DDFFE5EE72FC55F9468F7C6B16" box="[912,1029,1772,1796]" pageId="5" pageNumber="228" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, holotype, CNU-COL-MA-0321, under confocal microscopy.
<emphasis id="B90AEAA6FFE5EE72FEBDF8BA8AF36B3A" bold="true" box="[376,394,1808,1832]" pageId="5" pageNumber="228">A</emphasis>
, Mouthparts, ventral view.
<emphasis id="B90AEAA6FFE5EE72FD62F8BA89C16B3A" bold="true" box="[679,696,1808,1832]" pageId="5" pageNumber="228">B</emphasis>
, Antenna, ventral view.
<emphasis id="B90AEAA6FFE5EE72FC76F8BA88BC6B3A" bold="true" box="[947,965,1808,1832]" pageId="5" pageNumber="228">C</emphasis>
, Prothorax, ventral view, showing the incomplete boundary between the lateral medial portions of prosternum (arrowhead).
<emphasis id="B90AEAA6FFE5EE72FC1CF89E88926B5E" bold="true" box="[985,1003,1844,1868]" pageId="5" pageNumber="228">D</emphasis>
, Abdominal base, ventral view.
<emphasis id="B90AEAA6FFE5EE72FA85F89E8E286B5E" bold="true" box="[1344,1361,1844,1868]" pageId="5" pageNumber="228">E</emphasis>
, Head, anterodorsal view.
<emphasis id="B90AEAA6FFE5EE72FE92F8F28A1F6B62" bold="true" box="[343,358,1880,1904]" pageId="5" pageNumber="228">F</emphasis>
, Scutellar shield, dorsal view.
<emphasis id="B90AEAA6FFE5EE72FD59F8F289B36B62" bold="true" box="[668,714,1880,1904]" pageId="5" pageNumber="228">GJ</emphasis>
, Fore legs.
<emphasis id="B90AEAA6FFE5EE72FC85F8F2882D6B62" bold="true" box="[832,852,1880,1904]" pageId="5" pageNumber="228">K</emphasis>
, Mesotarsus.
<emphasis id="B90AEAA6FFE5EE72FC25F8F288886B62" bold="true" box="[992,1009,1880,1904]" pageId="5" pageNumber="228">L</emphasis>
, Metatibia. Abbreviations: an811, antennomeres 811; el, elytron; hd, head; lbp, labial palp; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtv, metaventrite; mxp, maxillary palp; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum; v13, ventrites 13. Scale bars: 200 μm.
</paragraph>
</caption>
<caption id="DF01663CFFE6EE71FF52FA6189E56A15" ID-DOI="http://doi.org/10.5281/zenodo.5788241" ID-Zenodo-Dep="5788241" httpUri="https://zenodo.org/record/5788241/files/figure.png" pageId="6" pageNumber="229" startId="6.[151,250,1483,1507]" targetBox="[151,1436,181,1459]" targetPageId="6">
<paragraph id="8BC136B4FFE6EE71FF52FA6189E56A15" blockId="6.[151,1437,1483,1543]" pageId="6" pageNumber="229">
<emphasis id="B90AEAA6FFE6EE71FF52FA618A6B69F1" bold="true" box="[151,274,1483,1507]" pageId="6" pageNumber="229">FIGURE 5.</emphasis>
General habitus of
<taxonomicName id="4C7E4D37FFE6EE71FE13FA61880369F1" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2021" box="[470,890,1483,1507]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="229" phylum="Arthropoda" rank="species" species="explanatum" status="sp. nov.">
<emphasis id="B90AEAA6FFE6EE71FE13FA61886569F1" box="[470,796,1483,1507]" italics="true" pageId="6" pageNumber="229">Archaenosodendron explanatum</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="B90AEAA6FFE6EE71FCBAFA6188B669F1" bold="true" box="[895,975,1483,1507]" pageId="6" pageNumber="229">
<taxonomicNameLabel id="A23957DDFFE6EE71FCBAFA6188B669F1" box="[895,975,1483,1507]" pageId="6" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP177613, under incident light.
<emphasis id="B90AEAA6FFE6EE71FF52FA458BD06A15" bold="true" box="[151,169,1519,1543]" pageId="6" pageNumber="229">A</emphasis>
, Dorsal view.
<emphasis id="B90AEAA6FFE6EE71FEF9FA458A346A15" bold="true" box="[316,333,1519,1543]" pageId="6" pageNumber="229">B</emphasis>
, Ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="C364653FFFE6EE70FF02F99F891B6D77" lastPageId="7" lastPageNumber="230" pageId="6" pageNumber="229" type="description">
<paragraph id="8BC136B4FFE6EE71FF02F99F8FDA6A5D" blockId="6.[151,1437,1589,2012]" box="[199,1187,1589,1616]" pageId="6" pageNumber="229">
<emphasis id="B90AEAA6FFE6EE71FF02F99F8A206A5D" bold="true" box="[199,345,1589,1615]" pageId="6" pageNumber="229">Description.</emphasis>
Body flattened and relatively elongate, about
<quantity id="4C869B51FFE6EE71FC9AF99F88BA6A42" box="[863,963,1589,1616]" metricMagnitude="-3" metricUnit="m" metricValue="3.67" pageId="6" pageNumber="229" unit="mm" value="3.67">3.67 mm</quantity>
long,
<quantity id="4C869B51FFE6EE71FBCCF99F8F266A42" box="[1033,1119,1589,1616]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="229" unit="mm" value="1.4">1.4 mm</quantity>
wide.
</paragraph>
<paragraph id="8BC136B4FFE6EE71FF02F9F38F4D6ACD" blockId="6.[151,1437,1589,2012]" pageId="6" pageNumber="229">
Head prognathous. Eyes not strongly protuberant. Antennal insertions widely separated and concealed from above. Antennal grooves on head well developed. Frontoclypeal suture absent. Mandibles broad, unidentate. Maxillae with setose lacinia (galea and palp detached).
<collectingRegion id="49BAF856FFE6EE71FD79F90888626AAE" box="[700,795,1698,1724]" country="Myanmar" name="Chin" pageId="6" pageNumber="229">Mentum</collectingRegion>
subtrapezoidal, widest at base; lateral regions with somewhat longitudinally arranged larger punctures; anterior angles produced anteriorly.
</paragraph>
<paragraph id="8BC136B4FFE6EE71FF02F9438AC96B81" blockId="6.[151,1437,1589,2012]" pageId="6" pageNumber="229">Pronotal disc subtrapezoidal, widest at base; anterior margin with very shallow emarginations laterally, slightly protruding medially. Prosternum in front coxae transverse, not extremely shortened; medial and lateral portions of prosternum not separated by a ridge; antennal grooves on prosternum longer than half-length of prosternum in front of coxae; prosternal process relatively slender. Procoxae strongly transverse, well separated. Procoxal cavities broadly open posteriorly.</paragraph>
<paragraph id="8BC136B4FFE6EE70FF02F837885A6CC7" blockId="6.[151,1437,1589,2012]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="230" pageId="6" pageNumber="229">Scutellar shield triangular, posteriorly acute. Elytra elongate, subparallel in the anterior half, tapering apically; anterior corners almost right. Mesoventrite short, anteriorly with well-developed mesoventral cavity and procoxal rests. Mesocoxal cavities widely separated. Metaventrite transverse, flattened; anterolateral portion with curved line delimiting the impression. Metacoxae narrowly separated.</paragraph>
<paragraph id="8BC136B4FFE7EE70FF02FF7589336D0F" blockId="7.[151,1436,151,357]" pageId="7" pageNumber="230">Femora with groove for receiving tibia. Tibiae expanded distally, with two apical spurs; outer edge of protibia with about 2123 denticles/stout setae.</paragraph>
<paragraph id="8BC136B4FFE7EE70FF02FE8D891B6D77" blockId="7.[151,1436,151,357]" pageId="7" pageNumber="230">Abdomen with five ventrites. Ventrites 1 and 2 with a pair of impressions for receiving hind legs. Anterior margin of ventrites smooth, without notches.</paragraph>
</subSubSection>
</treatment>
</document>