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<document id="393E7F56EB5091598A0A3EE8FEDE4A75" ID-CLB-Dataset="6633" ID-DOI="10.5281/zenodo.4648766" ID-GBIF-Dataset="2f0e5269-a3b3-489e-a4c1-7e766cefb113" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648766" ID-ZooBank="A3C18398-5DB2-449B-B87C-B2E614AE2936" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617136404546" checkinUser="carolina" docAuthor="Sulakhe, Shauri, Deshpande, Shubhankar, Dandekar, Nikhil, Ketkar, Makarand, Gowande, Gaurang, Padhye, Anand &amp; Bastawade, Deshabhushan" docDate="2020" docId="03BCD52E417AE14EEDE4F9BCFD01F923" docLanguage="en" docName="Euscorpius.320.1-27.pdf" docOrigin="Euscorpius 320" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Chiromachetes Pocock 1899" docType="treatment" docVersion="9" lastPageNumber="4" masterDocId="FF85AD56417FE14BED3AFFA8FFDBFFED" masterDocTitle="Two new species of Chiromachetes (Scorpiones: Hormuridae) from the northern Western Ghats, India" masterLastPageNumber="27" masterPageNumber="1" pageNumber="4" updateTime="1698935505347" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="085B88ADC8687C94B0C970DAEE2FCBE4">Two new species of Chiromachetes (Scorpiones: Hormuridae) from the northern Western Ghats, India</mods:title>
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<mods:namePart id="469B0ED1C3ED17C814ACEF54949F3A70">Dandekar, Nikhil</mods:namePart>
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<mods:namePart id="C8066F58DCFCDBE2B91735DEB1566BCD">Ketkar, Makarand</mods:namePart>
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<mods:namePart id="66BD40117CEAB9435AA9A79A2E9B1FCD">Padhye, Anand</mods:namePart>
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<emphasis id="B961B82A417AE14EEDE4F9BCFEF5F9C3" bold="true" box="[222,302,1556,1582]" pageId="5" pageNumber="4">Genus</emphasis>
<taxonomicName id="4C151FBB417AE14EEC0FF9BBFD4CF9C3" ID-CoL="3NC2" authority="Pocock, 1899" authorityName="Pocock" authorityYear="1899" box="[309,663,1555,1582]" class="Arachnida" family="Hormuridae" genus="Chiromachetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="B961B82A417AE14EEC0FF9BBFE37F9C3" bold="true" box="[309,492,1555,1582]" italics="true" pageId="5" pageNumber="4">Chiromachetes</emphasis>
Pocock
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,
<emphasis id="B961B82A417AE14EEF66F9BCFD4CF9C3" bold="true" box="[604,663,1556,1582]" pageId="5" pageNumber="4">1899</emphasis>
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(
<figureCitation id="132E78BD417AE14EEC08F99EFE19F9A3" box="[306,450,1590,1614]" captionStart-0="Figures 1–6" captionStart-1="Figures 7–9" captionStart-10="Figures 49–57" captionStart-11="Figures 58–60" captionStart-12="Figures" captionStart-13="Figures 64–69" captionStart-14="Figures 70–73" captionStart-15="Figures 74–80" captionStart-16="Figures 81–82" captionStart-17="Figure 83" captionStart-2="Figures 10–13" captionStart-3="Figures 14–18" captionStart-4="Figures 19–27" captionStart-5="Figures 28–30" captionStart-6="Figure 31–36" captionStart-7="Figures 37–39" captionStart-8="Figures 40–43" captionStart-9="Figures 44–48" captionStartId-0="4.[115,192,1982,2004]" captionStartId-1="6.[113,190,968,990]" captionStartId-10="17.[112,189,1950,1972]" captionStartId-11="18.[113,190,1967,1989]" captionStartId-12="19.[116,193,1990,2012]" captionStartId-13="21.[116,193,1991,2013]" captionStartId-14="23.[115,192,2019,2041]" captionStartId-15="24.[113,190,1130,1152]" captionStartId-16="25.[115,192,1981,2003]" captionStartId-17="26.[116,184,1502,1524]" captionStartId-2="7.[116,193,1998,2020]" captionStartId-3="8.[115,192,1988,2010]" captionStartId-4="9.[116,193,1945,1967]" captionStartId-5="10.[113,190,1960,1982]" captionStartId-6="12.[113,181,1976,1998]" captionStartId-7="14.[115,192,769,791]" captionStartId-8="15.[113,190,1992,2014]" captionStartId-9="16.[116,193,1992,2014]" captionTargetBox-0="[113,1474,249,1973]" captionTargetBox-1="[113,1474,234,957]" captionTargetBox-10="[124,1448,226,1941]" captionTargetBox-11="[115,1476,229,1950]" captionTargetBox-12="[113,1474,234,1963]" captionTargetBox-13="[113,1467,239,1973]" captionTargetBox-14="[113,1474,227,2004]" captionTargetBox-15="[120,1441,238,1094]" captionTargetBox-16="[113,1474,234,1970]" captionTargetBox-17="[180,1451,290,1479]" captionTargetBox-2="[113,1474,230,1991]" captionTargetBox-3="[113,1474,272,1973]" captionTargetBox-4="[139,1474,233,1930]" captionTargetBox-5="[111,1472,232,1952]" captionTargetBox-6="[113,1474,259,1951]" captionTargetBox-7="[113,1474,233,763]" captionTargetBox-8="[113,1474,228,1975]" captionTargetBox-9="[113,1474,270,1973]" captionTargetId-0="figure-52@4.[113,1474,234,1973]" captionTargetId-1="figure-566@6.[113,1474,234,957]" captionTargetId-10="figure-92@17.[113,1474,223,1951]" captionTargetId-11="figure-13@18.[115,1476,226,1953]" captionTargetId-12="figure-9@19.[113,1474,234,1965]" captionTargetId-13="figure-8@21.[113,1474,234,1973]" captionTargetId-14="figure-7@23.[113,1474,227,2004]" captionTargetId-15="figure-396@24.[113,1474,234,1121]" captionTargetId-16="figure-7@25.[113,1474,234,1973]" captionTargetId-17="figure-266@26.[136,1476,253,1495]" captionTargetId-2="figure-42@7.[113,1474,227,1993]" captionTargetId-3="figure-49@8.[113,1474,234,1973]" captionTargetId-4="figure-93@9.[113,1474,229,1940]" captionTargetId-5="figure-65@10.[111,1472,230,1955]" captionTargetId-6="figure-53@12.[113,1474,255,1954]" captionTargetId-7="figure-746@14.[113,1474,230,766]" captionTargetId-8="figure-42@15.[113,1474,225,1975]" captionTargetId-9="figure-49@16.[113,1474,234,1973]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-10="17" captionTargetPageId-11="18" captionTargetPageId-12="19" captionTargetPageId-13="21" captionTargetPageId-14="23" captionTargetPageId-15="24" captionTargetPageId-16="25" captionTargetPageId-17="26" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="12" captionTargetPageId-7="14" captionTargetPageId-8="15" captionTargetPageId-9="16" captionText-0="Figures 1–6. Chiromachetes parakrami sp. n., male, holotype, dorsal (1) and ventral (2) views, sternopectinal area (3), metasomal segment V and telson in lateral view (4), chelicera in dorsal view (5) and tarsi IV (6)." captionText-1="Figures 7–9. Chiromachetes parakrami sp. n., male, holotype, carapace (7), anterior margin of carapace (8) and left lateral eyes in dorsal view (9)." captionText-10="Figures 49–57: Chiromachetes ramdasswamii sp. n. Figures 49–55. Male, holotype, pedipalp chela dorsal (49), external (50) and ventral (51) views, patella dorsal (52), ventral (53) and external (54) views and femur dorsal (55) view. Trichpobothrial pattern indicated by yellow circles. Figures 56–57. Male, paratype, INHER 254, right hemispermatophore in internal (56) and external (57) views. Abbreviations: p, pedicel; t, trunk; c, capsule; h, hook; dl, distal lamina." captionText-11="Figures 58–60. Type locality of Chiromachetes ramdasswamii sp. n., view of riparian forest with waterfall at the type locality (58), view of Kawla fort through which the Varandha Ghat road is carved (59) and basaltic bounder adjacent to the Varandha Ghat road from where the type series is collected (60)." captionText-12="Figures: 61–63. Figure 61. Distribution of Chiromachetes in India.Figures 62–63. Chiromachetes sahyadriensis, male, INHER 172, carapace under UV light (62) and metasomal segment V and telson in lateral view (63)." captionText-13="Figures 64–69. Chiromachetes fergusoni, male topotype, dorsal (64) and ventral (65) views, sternopectinal area (66), metasoma V and telson lateral view (67), chelicera dorsal (68) and tarsi IV (69)." captionText-14="Figures 70–73. Chiromachetes fergusoni, male topotype, dorsal (70) and ventral (71) views under UV lights, carapace under white (72) and UV (73) light." captionText-15="Figures 74–80. Chiromachetes fergusoni, male topotype, pedipalp chela dorsal (74), external (75) and ventral (76) views, patella dorsal (77), ventral (78) and external (79) views and femur dorsal (80) view. Trichobothrial pattern indicated by yellow circles." captionText-16="Figures 81–82. Figure 81. Discriminant function analyses projection on first two factor planes explaining 96.85% of variation among the five species.Abbreviations:Circlecells (males), Squarecells (females).Figure 82. Maximum Likelihood phylogenetic tree (ML) for Chiromachetes from northern Western Ghats of India. Values along the nodes are bootstraps for 1000 iterations." captionText-17="Figure 83. Bayesian phylogenetic tree for Chiromachetes from northern Western Ghats of India. Values along the nodes are Bayesian posterior probabilities for Bayesian Inference and Bayesian Poisson Tree Process (bPTP), respectively." captionText-2="Figures 10–13. Chiromachetes parakrami sp. n., male, holotype in dorsal (10) and ventral (11) views, carapace (12) and dentition on movable finger of chela (13) under UV fluorescence." captionText-3="Figures 14–18. Chiromachetes parakrami sp. n., female, paratype, BNHS SC 181, in dorsal (14) ventral (15) views, sternopectinal area (16), metasomal segment V and telson in lateral view (17) and carapace (18)." captionText-4="Figures 19–27: Chiromachetes parakrami sp. n. Figures 19–25. Male, holotype, pedipalp chela dorsal (19), external (20) and ventral (21) views, patella dorsal (22), ventral (23) and external (24) views and femur dorsal (25) view. Trichobothrial pattern indicated by yellow circles. Figures 26–27. Male, paratype, BNHS SC 180, right hemispermatophore in internal (26) and external (27) views.Abbreviations: p, pedicel; t, trunk; c, capsule; h, hook; dl, distal lamina." captionText-5="Figures 28–30. Type locality of Chiromachetes parakrami sp. n., view of the dense semi–evergreen forest and mountains of Amba Ghat (28), view of the Amba Ghat road with adjacent boulders (29) and columnar basaltic bounder at the type locality from where the type series is collected (30)." captionText-6="Figure 31–36. Chiromachetes ramdasswamii sp. n., male, holotype, dorsal (31) and ventral (32) views, sternopectinal area (33), metasomal segment V and telson in lateral view (34), chelicera in dorsal view (55) and tarsi IV (36)." captionText-7="Figures 37–39. Chiromachetes ramdasswamii sp. n., male, holotype, carapace (37), anterior margin of carapace (38) and right lateral eyes in dorsal view (39)." captionText-8="Figures 40–43. Chiromachetes ramdasswamii sp. n., male, holotype in dorsal (40) and ventral (41) views, carapace (42) and dentition on movable finger of chela (43) under UV fluorescence." captionText-9="Figures 44–48. Chiromachetes ramdasswamii sp. n., female, paratype, BNHS SC 184, in dorsal (44) ventral (45) views, sternopectinal area (46), metasomal segment V and telson in lateral view (47) and carapace (48)." figureDoi-0="http://doi.org/10.5281/zenodo.4648768" figureDoi-1="http://doi.org/10.5281/zenodo.4648770" figureDoi-10="http://doi.org/10.5281/zenodo.4648791" figureDoi-11="http://doi.org/10.5281/zenodo.4648793" figureDoi-12="http://doi.org/10.5281/zenodo.4648795" figureDoi-13="http://doi.org/10.5281/zenodo.4648797" figureDoi-14="http://doi.org/10.5281/zenodo.4648799" figureDoi-15="http://doi.org/10.5281/zenodo.4648801" figureDoi-16="http://doi.org/10.5281/zenodo.4648803" figureDoi-17="http://doi.org/10.5281/zenodo.4648805" figureDoi-2="http://doi.org/10.5281/zenodo.4648772" figureDoi-3="http://doi.org/10.5281/zenodo.4648774" figureDoi-4="http://doi.org/10.5281/zenodo.4648776" figureDoi-5="http://doi.org/10.5281/zenodo.4648778" figureDoi-6="http://doi.org/10.5281/zenodo.4648780" figureDoi-7="http://doi.org/10.5281/zenodo.4648785" figureDoi-8="http://doi.org/10.5281/zenodo.4648787" figureDoi-9="http://doi.org/10.5281/zenodo.4648789" httpUri-0="https://zenodo.org/record/4648768/files/figure.png" httpUri-1="https://zenodo.org/record/4648770/files/figure.png" httpUri-10="https://zenodo.org/record/4648791/files/figure.png" httpUri-11="https://zenodo.org/record/4648793/files/figure.png" httpUri-12="https://zenodo.org/record/4648795/files/figure.png" httpUri-13="https://zenodo.org/record/4648797/files/figure.png" httpUri-14="https://zenodo.org/record/4648799/files/figure.png" httpUri-15="https://zenodo.org/record/4648801/files/figure.png" httpUri-16="https://zenodo.org/record/4648803/files/figure.png" httpUri-17="https://zenodo.org/record/4648805/files/figure.png" httpUri-2="https://zenodo.org/record/4648772/files/figure.png" httpUri-3="https://zenodo.org/record/4648774/files/figure.png" httpUri-4="https://zenodo.org/record/4648776/files/figure.png" httpUri-5="https://zenodo.org/record/4648778/files/figure.png" httpUri-6="https://zenodo.org/record/4648780/files/figure.png" httpUri-7="https://zenodo.org/record/4648785/files/figure.png" httpUri-8="https://zenodo.org/record/4648787/files/figure.png" httpUri-9="https://zenodo.org/record/4648789/files/figure.png" pageId="5" pageNumber="4">Figures 1–83</figureCitation>
,
<tableCitation id="C6975183417AE14EECF7F99EFD9AF9A3" box="[461,577,1590,1614]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStart-4="Table 5" captionStart-5="Table 6" captionStart-6="Table 7" captionStartId-0="5.[113,169,1159,1180]" captionStartId-1="13.[113,169,1977,1998]" captionStartId-2="20.[115,171,1123,1144]" captionStartId-3="22.[111,167,745,766]" captionStartId-4="22.[110,166,1070,1091]" captionStartId-5="22.[115,171,1503,1524]" captionStartId-6="22.[116,172,1993,2014]" captionTargetBox-3="[122,1447,256,727]" captionTargetId-0="graphics-343@5.[110,1468,250,1133]" captionTargetId-1="graphics-41@13.[114,1474,253,1946]" captionTargetId-2="graphics-12@20.[116,1469,237,1093]" captionTargetId-3="graphics-34@22.[116,1467,238,729]" captionTargetId-4="graphics-156@22.[111,1469,823,1054]" captionTargetId-5="graphics-218@22.[111,1462,1155,1485]" captionTargetId-6="graphics-283@22.[113,1469,1565,1971]" captionTargetPageId-0="5" captionTargetPageId-1="13" captionTargetPageId-2="20" captionTargetPageId-4="22" captionTargetPageId-5="22" captionTargetPageId-6="22" captionText-0="Table 1. Morphometric data for Chiromachetes parakrami sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D), holotype (HT), paratype (PT)." captionText-1="Table 2. Morphometric data for Chiromachetes ramdasswamii sp. n. and C. fergusoni Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D), holotype (HT), paratype (PT)." captionText-2="Table 3. Morphometric data for Chiromachetes sahyadriensis. Abbreviations: length (L), width (W, in carapace it corresponds to median width), depth (D)." captionText-3="Table 4. Comparison among Chiromachetes species based upon selected morphometric ratios of adults. Abbreviations: length (L), width (W) depth (D)." captionText-4="Table 5. Pairwise uncorrected raw distances (%) expressed as minimum–maximum based on COI gene sequence for Chiromachetes species from northern Western Ghats, India. Values in brackets are intra–clade distances." captionText-5="Table 6: Primers used for PCR amplification and sequencing of (COI) mitochondrial gene." captionText-6="Table 7: Voucher numbers and GenBank accession numbers for the sequence data used for the phylogenetic analysis (* Indicate non-type specimens)." httpUri-0="http://table.plazi.org/id/DF6A34B0417AE14EED4BFB2FFD9DFB57" httpUri-1="http://table.plazi.org/id/DF6A34B04172E146ED4BF811FCFEF801" httpUri-2="http://table.plazi.org/id/DF6A34B0416BE15FED49FBCBFEFAFB7B" httpUri-3="http://table.plazi.org/id/DF6A34B04169E15DED55FD41FF09FCF1" httpUri-4="http://table.plazi.org/id/DF6A34B04169E15DED54FB86FCADFB8C" httpUri-5="http://table.plazi.org/id/DF6A34B04169E15DED49FA77FC00FA18" httpUri-6="http://table.plazi.org/id/DF6A34B04169E15DED4EF861FF3CF811" pageId="5" pageNumber="4" tableUuid-0="DF6A34B0417AE14EED4BFB2FFD9DFB57" tableUuid-1="DF6A34B04172E146ED4BF811FCFEF801" tableUuid-2="DF6A34B0416BE15FED49FBCBFEFAFB7B" tableUuid-3="DF6A34B04169E15DED55FD41FF09FCF1" tableUuid-4="DF6A34B04169E15DED54FB86FCADFB8C" tableUuid-5="DF6A34B04169E15DED49FA77FC00FA18" tableUuid-6="DF6A34B04169E15DED4EF861FF3CF811">Tables 1–7</tableCitation>
)
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<emphasis id="B961B82A417AE14EEF4EF9FEFD5EF963" bold="true" pageId="5" pageNumber="4">D1B455F7– E75B–4DF5–BAC7–31B0341D1784</emphasis>
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</subSubSection>
<subSubSection id="C30F37B3417AE14EED4BF91EFD01F923" box="[113,730,1718,1742]" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BAA6438417AE14EED4BF91EFD01F923" blockId="5.[113,730,1718,1742]" box="[113,730,1718,1742]" pageId="5" pageNumber="4">
<materialsCitation id="3B7D6E65417AE14EED4BF91EFD0DF923" ID-GBIF-Occurrence="3114122306" box="[113,726,1718,1742]" collectionCode="SPECIES" pageId="5" pageNumber="4" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AEDA9A417AE14EED4BF91EFF6FF920" box="[113,180,1718,1741]" pageId="5" pageNumber="4">TYPE</typeStatus>
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<taxonomicName id="4C151FBB417AE14EEC0AF91EFD0DF923" authority="Pocock, 1899" authorityName="Pocock" authorityYear="1899" box="[304,726,1718,1742]" class="Arachnida" family="Hormuridae" genus="Chiromachetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="fergusoni">
<emphasis id="B961B82A417AE14EEC0AF91EFDE0F923" box="[304,571,1718,1742]" italics="true" pageId="5" pageNumber="4">Chiromachetes fergusoni</emphasis>
Pocock, 1899
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.
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</treatment>
</document>
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