273 lines
35 KiB
XML
273 lines
35 KiB
XML
<document id="04219B7D6B38798843C1F2AA7C6238F3" ID-DOI="10.5281/zenodo.276689" ID-GBIF-Dataset="b6781430-d288-4cb4-b178-db6192a61dde" ID-ISSN="1175-5326" ID-Zenodo-Dep="276689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460230557710" checkinUser="plazi" docAuthor="Enghoff, Henrik" docDate="2011" docId="03BC87EAFFB93176FF22FCFD0639FA81" docLanguage="en" docName="zt02753p041.pdf" docOrigin="Zootaxa 2753" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hyperbolus morogoroensis Kraus 1958, n. comb." docType="treatment" docVersion="9" lastPageNumber="23" masterDocId="FF85FF92FFAD3160FFB5FFF00761FFF9" masterDocTitle="East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="21" updateTime="1698238933756" updateUser="plazi">
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<mods:titleInfo id="F03A1939EC0C26D0BEED0CC5CB122796">
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<mods:title id="1A40874E460EF1E001CCB7B6C0A58A26">East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="9445DA942A73A4FCE02F8E6D8B64096F" type="personal">
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<mods:role id="711D402956522C51240A1828937683EB">
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<mods:roleTerm id="7C7CA5CC7B20ADB7E79762FF62F5541C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0C30AB54A8150A2FC4194F77B5EAAE68">Enghoff, Henrik</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="846CED834FB99461863B560CB108DFFA">text</mods:typeOfResource>
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<mods:titleInfo id="3B4F48D12AA77DBE35DEEA63A85CCDFC">
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<mods:title id="A4D830BA909ECAC1C04FF966ACCD836F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="624662A672E12A92F39858A9D440CA9F">
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<mods:date id="8C7C065916EF7B9670A8D474E752BBF8">2011</mods:date>
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<mods:detail id="A15EB96B0CACD6174AA29E753DCF3F0B" type="volume">
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<mods:number id="8DC76A94788F194B28007FAE656B57B2">2753</mods:number>
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</mods:detail>
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<mods:extent id="D073C7E2DC46E95FDC30C6E8A66A4150" unit="page">
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<mods:start id="C82CEC2B68B80031179DD328F5E771DB">1</mods:start>
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<mods:end id="70DA705297918496D15CD5D58C6254C9">41</mods:end>
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<mods:classification id="EC4F4523A5C5F6557AFA6C79A6BCCC0A">journal article</mods:classification>
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<mods:identifier id="34E8C2F38BAA1F2AE427FE62EED54BA7" type="DOI">10.5281/zenodo.276689</mods:identifier>
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<mods:identifier id="B24D789E3B78BFD4EC940E3405C1DA9D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="DB999EE943C0B99C4903A229F7DA3434" type="Zenodo-Dep">276689</mods:identifier>
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<treatment id="03BC87EAFFB93176FF22FCFD0639FA81" ID-DOI="http://doi.org/10.5281/zenodo.5612658" ID-GBIF-Taxon="119387235" ID-Zenodo-Dep="5612658" LSID="urn:lsid:plazi:treatment:03BC87EAFFB93176FF22FCFD0639FA81" httpUri="http://treatment.plazi.org/id/03BC87EAFFB93176FF22FCFD0639FA81" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<subSubSection id="C30F6577FFB93174FF22FCFD065FFCB1" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BAA36FCFFB93174FF22FCFD0469FCDE" blockId="20.[151,776,781,840]" box="[151,776,781,807]" pageId="20" pageNumber="21">
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<heading id="D0E28190FFB93174FF22FCFD0469FCDE" bold="true" box="[151,776,781,807]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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||
<emphasis id="B961EAEEFFB93174FF22FCFD0469FCDE" bold="true" box="[151,776,781,807]" pageId="20" pageNumber="21">
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||
<taxonomicName id="4C154D7FFFB93174FF22FCFD05F1FCDE" ID-CoL="3NNX5" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[151,656,781,807]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="morogoroensis" status="comb. nov.">
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<emphasis id="B961EAEEFFB93174FF22FCFD06BCFCDE" bold="true" box="[151,477,781,807]" italics="true" pageId="20" pageNumber="21">Hyperbolus morogoroensis</emphasis>
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||
(
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||
<bibRefCitation id="EF844B0DFFB93174FE58FCFD05E9FCDE" author="Kraus" box="[493,648,781,807]" pageId="20" pageNumber="40" refString="Kraus, O. (1958) Myriapoden aus Ostafrika (Tanganyika Territory). Veroffentlichungen des Uberseemuseums Bremen, Reihe A, 3 (1), 1 - 16." type="journal article" year="1958">Kraus, 1958</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="A2525795FFB93174FD2BFCFD0469FCDE" box="[670,776,781,807]" pageId="20" pageNumber="21">n. comb.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BAA36FCFFB93174FF22FCC0065FFCB1" blockId="20.[151,776,781,840]" box="[151,318,816,840]" pageId="20" pageNumber="21">
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||
<figureCitation id="132E2A79FFB93174FF22FCC00676FCB1" box="[151,279,816,840]" captionStart-0="FIGURES 38 – 41. 38" captionStart-1="FIGURES 42 – 43" captionStartId-0="21.[151,264,1827,1850]" captionStartId-1="22.[151,264,1042,1065]" captionTargetBox-0="[189,1397,342,1801]" captionTargetBox-1="[209,1377,182,1024]" captionTargetId-0="figure@21.[184,1403,337,1806]" captionTargetId-1="figure@22.[208,1381,181,1031]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 38 – 41. 38: Hyperbolus morogoroensis, male from Ngambaula L. A. Forest Reserve, gonopod block, anterior view, G. Brovad phot., 39: do., posterior view, G. Brovad phot. s, 40: do., right anterior gonopod coxa, anterior view, SEM, 41: right anterior gonopod telopodite, anterior view (i. e., from inside of anterior gonopod cavity), SEM. Scales 1 cm. cx: coxa, fp: finger-shaped process, mp: mesal, thumb-like process of telopodite, st: sternum, tlp: telopodite." captionText-1="FIGURES 42 – 43. Hyperbolus morogoroensis. 42: male from Mkungwe Forest Reserve, tip of left posterior gonopod, oral view. 43: female from Mkungwe Forest Reserve, left vulva, posterior-lateral view. Scales 1 mm. ap: apical appendix, lav: subrectangular lobe of aboral valve, lov: subtriangular lobe of oral valve, op: operculum, slm: solenomerite." httpUri-0="https://zenodo.org/record/276704/files/figure.png" httpUri-1="https://zenodo.org/record/276705/files/figure.png" pageId="20" pageNumber="21">Figs 38–43</figureCitation>
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,
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<figureCitation id="132E2A79FFB93174FE97FCC0065FFCB1" box="[290,318,816,840]" captionStart="FIGURE 84" captionStartId="36.[151,249,1479,1502]" captionTargetBox="[456,1120,410,1444]" captionTargetId="figure@36.[456,1122,407,1445]" captionTargetPageId="36" captionText="FIGURE 84. Known distribution of East African Pachybolini, except Hadrobolus crassicollis and Epibolus pulchripes. dots (●): Crurifarcimen vagans, filled diamonds (♦): Hyperbolus apicomplexus, open diamonds (♢): Hyperbolus morogoroensis, filled triangle (˔): Parabolus dimorphus, inverted open triangle (): Parabolus calceus, inverted filled triangle (ˑ): Pachybolus tectus,, filled square (․): Pachybolini indet. Based on Yahoo maps." httpUri="https://zenodo.org/record/276715/files/figure.png" pageId="20" pageNumber="21">84</figureCitation>
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C30F6577FFB93174FF22FC8305FDFC5E" pageId="20" pageNumber="21" type="reference_group">
|
||
<paragraph id="8BAA36FCFFB93174FF22FC83054DFC70" blockId="20.[151,668,883,936]" box="[151,556,883,905]" pageId="20" pageNumber="21">
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||
<taxonomicName id="4C154D7FFFB93174FF22FC83054DFC70" ID-CoL="4BQ4R" authority="Kraus, 1958" authorityName="Kraus" authorityYear="1958" box="[151,556,883,905]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="morogoroensis">
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<emphasis id="B961EAEEFFB93174FF22FC8306C8FC70" box="[151,425,883,905]" italics="true" pageId="20" pageNumber="21">Pachybolus morogoroensis</emphasis>
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||
<bibRefCitation id="EF844B0DFFB93174FE05FC84054DFC70" author="Kraus" box="[432,556,883,905]" pageId="20" pageNumber="40" refString="Kraus, O. (1958) Myriapoden aus Ostafrika (Tanganyika Territory). Veroffentlichungen des Uberseemuseums Bremen, Reihe A, 3 (1), 1 - 16." type="journal article" year="1958">Kraus, 1958</bibRefCitation>
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||
</taxonomicName>
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||
</paragraph>
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||
<paragraph id="8BAA36FCFFB93174FF22FC6205FDFC5E" blockId="20.[151,668,883,936]" box="[151,668,914,936]" pageId="20" pageNumber="21">
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||
‘
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<emphasis id="B961EAEEFFB93174FF2AFC6206A7FC51" box="[159,454,914,936]" italics="true" pageId="20" pageNumber="21">
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||
Pachybolous’
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<taxonomicName id="4C154D7FFFB93174FE85FC6306A7FC51" ID-CoL="4BQ4R" authority="Wesener et al. 2008" authorityName="Wesener et al." authorityYear="2008" box="[304,454,915,936]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="morogoroensis">morogoroensis</taxonomicName>
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||
</emphasis>
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||
:
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<bibRefCitation id="EF844B0DFFB93174FE61FC6305FDFC5E" author="Wesener" box="[468,668,914,936]" pageId="20" pageNumber="41" refString="Wesener, T., Enghoff, H. & Wagele, J. - W. (2008) Pachybolini - a tribe of giant Afrotropical millipedes. With arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics, 22, 37 - 53." type="journal article" year="2008">
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||
Wesener
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<emphasis id="B961EAEEFFB93174FD84FC630505FC51" box="[561,612,914,936]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
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2008
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</bibRefCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30F6577FFB93174FF22FC230414FADA" pageId="20" pageNumber="21" type="materials_examined">
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||
<paragraph id="8BAA36FCFFB93174FF22FC2306BEFBC9" blockId="20.[151,1436,979,1870]" pageId="20" pageNumber="21">
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<typeStatus id="54AE885EFFB93174FF22FC23064AFC12" box="[151,299,979,1003]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
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3,
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<collectingCountry id="F302766CFFB93174FEE9FC23068FFC12" box="[348,494,979,1003]" name="Tanzania" pageId="20" pageNumber="21">TANZANIA</collectingCountry>
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, Morogoro District, Uluguru Mountains, between Kimboza and Kibungu,
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<geoCoordinate id="EE21503BFFB93174FAFBFC2302FDFC12" box="[1358,1436,979,1003]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-7.0333333">7°02’S</geoCoordinate>
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||
<geoCoordinate id="EE21503BFFB93174FF22FC060796FBF7" box="[151,247,1014,1038]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="37.733334">37°44’E</geoCoordinate>
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,
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<quantity id="4CED9B19FFB93174FEB6FC06062CFBF7" box="[259,333,1014,1038]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="m" value="500.0">500 m</quantity>
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, in rain forest under leaf litter and in rotten wood,
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<date id="FFAB103CFFB93174FC38FC06036FFBF7" box="[909,1038,1014,1038]" pageId="20" pageNumber="21" value="1952-07-16">16.vii.1952</date>
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, H. Knipper leg. (
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<collectionCode id="ED04AE39FFB93174FB6EFC060240FBF7" box="[1243,1313,1014,1038]" httpUri="http://grbio.org/cool/9tx7-8fc2" name="Uebersee-Museum" pageId="20" pageNumber="21">UMB</collectionCode>
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). – PARA-
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<typeStatus id="54AE885EFFB93174FF22FBE907BCFBC9" box="[151,221,1049,1072]" pageId="20" pageNumber="21">
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||
<collectionCode id="ED04AE39FFB93174FF22FBE907BCFBC9" box="[151,221,1049,1072]" pageId="20" pageNumber="21">TYPE</collectionCode>
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</typeStatus>
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1 Ƥ, same data (SNF).
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</paragraph>
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<paragraph id="8BAA36FCFFB93174FF70FBCC0414FADA" blockId="20.[151,1436,979,1870]" pageId="20" pageNumber="21">
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||
<collectionCode id="ED04AE39FFB93174FF70FBCC0664FBAA" box="[197,261,1084,1107]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="20" pageNumber="21">NEW</collectionCode>
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<collectionCode id="ED04AE39FFB93174FEBBFBCB06FAFBAA" box="[270,411,1083,1107]" pageId="20" pageNumber="21">MATERIAL</collectionCode>
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from
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<collectingCountry id="F302766CFFB93174FE51FBCB0517FBAA" box="[484,630,1083,1107]" name="Tanzania" pageId="20" pageNumber="21">TANZANIA</collectingCountry>
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, Morogoro region, Morogoro district, N. Doggart, Uluguru Mountains Biodiversity Conservation Project, leg.: 1 3, 1 Ƥ,
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<specimenCount id="9D13FD75FFB93174FD7DFBAE0456FB8F" box="[712,823,1118,1142]" pageId="20" pageNumber="21" type="juvenile">1 juvenile</specimenCount>
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Ƥ Kasanga Forest Reserve,
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<geoCoordinate id="EE21503BFFB93174FBC3FBAE03A5FB8F" box="[1142,1220,1118,1142]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-7.1666665">7°10’S</geoCoordinate>
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||
<geoCoordinate id="EE21503BFFB93174FB7EFBAE024AFB8F" box="[1227,1323,1118,1142]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="37.75">37°45’E</geoCoordinate>
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, leaf litter and rotting logs,
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<date id="FFAB103CFFB93174FED4FB70055FFB61" box="[353,574,1152,1176]" pageId="20" pageNumber="21" value="2000-07-27" valueMax="2000-08-02" valueMin="2000-07-27">27.vii. –2.viii.2000</date>
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, (
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<collectionCode id="ED04AE39FFB93174FDE0FB7005C9FB61" box="[597,680,1152,1176]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="21">ZMUC</collectionCode>
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||
00101080). – 2 3, 1 Ƥ Mkungwe Forest Reserve,
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<geoCoordinate id="EE21503BFFB93174FA94FB7002FDFB61" box="[1313,1436,1152,1176]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="-6.875">6°52’30’’S</geoCoordinate>
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||
<geoCoordinate id="EE21503BFFB93174FF22FB530642FB42" box="[151,291,1187,1211]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="15" value="37.89167">37°53’30’’E</geoCoordinate>
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||
, lowland submontane forest, 400–1000 masl,
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<date id="FFAB103CFFB93174FC9BFB5304F4FB42" box="[814,917,1187,1211]" pageId="20" pageNumber="21" value="2000-08">viii.2000</date>
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, (
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<collectionCode id="ED04AE39FFB93174FC1CFB53049DFB42" box="[937,1020,1187,1211]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="21">ZMUC</collectionCode>
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||
00101082). – 1 3, 1 Ƥ Ruvu Forest Reserve,
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||
<geoCoordinate id="EE21503BFFB93174FF4AFB36062FFB27" box="[255,334,1222,1246]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-7.0">7°00’S</geoCoordinate>
|
||
<geoCoordinate id="EE21503BFFB93174FEE1FB3606D5FB27" box="[340,436,1222,1246]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="37.85">37°51’E</geoCoordinate>
|
||
, 200 masl, 2000 (
|
||
<collectionCode id="ED04AE39FFB93174FDCEFB3605AFFB27" box="[635,718,1222,1246]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="21">ZMUC</collectionCode>
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||
00101083). - 1 3, 1 Ƥ Ngambaula L.A. Forest Reserve,
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<geoCoordinate id="EE21503BFFB93174FAFBFB3602FDFB27" box="[1358,1436,1222,1246]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-6.9833336">6°59’S</geoCoordinate>
|
||
<geoCoordinate id="EE21503BFFB93174FF22FB180796FAF9" box="[151,247,1256,1280]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="37.75">37°45’E</geoCoordinate>
|
||
, 250 masl, 2000 (
|
||
<collectionCode id="ED04AE39FFB93174FE6EFB18054FFAF9" box="[475,558,1256,1280]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="21">ZMUC</collectionCode>
|
||
00101084). – 1 Ƥ Chamanyani Forest Reserve, Mvuha, lowland forest,
|
||
<geoCoordinate id="EE21503BFFB93174FF22FAFB0670FADA" box="[151,273,1291,1315]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="15" value="-7.1749997">7°10’30’’S</geoCoordinate>
|
||
<geoCoordinate id="EE21503BFFB93174FEADFAFB0618FADA" box="[280,377,1291,1315]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="37.816666">37°49’E</geoCoordinate>
|
||
, 140–400 masl,
|
||
<date id="FFAB103CFFB93174FD85FAFB05EDFADA" box="[560,652,1291,1315]" pageId="20" pageNumber="21" value="2000-09">ix. 2000</date>
|
||
(
|
||
<collectionCode id="ED04AE39FFB93174FD2FFAFB058CFADA" box="[666,749,1291,1315]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="21">ZMUC</collectionCode>
|
||
00101086).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F6577FFB93174FF70FADE02E2FA29" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="8BAA36FCFFB93174FF70FADE02E2FA29" blockId="20.[151,1436,979,1870]" pageId="20" pageNumber="21">
|
||
MORE NEW MATERIAL from
|
||
<collectingCountry id="F302766CFFB93174FD8CFADE05ADFABF" box="[569,716,1326,1350]" name="Tanzania" pageId="20" pageNumber="21">TANZANIA</collectingCountry>
|
||
: 2 3, Uluguru Mountains, Vituri,
|
||
<geoCoordinate id="EE21503BFFB93174FBD7FADE03D1FABF" box="[1122,1200,1326,1350]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-7.1666665">7°10’S</geoCoordinate>
|
||
<geoCoordinate id="EE21503BFFB93174FB0DFADE0279FABF" box="[1208,1304,1326,1350]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="37.75">37°45’E</geoCoordinate>
|
||
,
|
||
<date id="FFAB103CFFB93174FA90FADE02F6FABF" box="[1317,1431,1326,1350]" pageId="20" pageNumber="21" value="1926-10-30">30.x.1926</date>
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, A. Loveridge leg. (MCZ no. 67914). – 1 Ƥ Uluguru Mountains, Kikosi near Bagilo.
|
||
<geoCoordinate id="EE21503BFFB93174FBFDFAA003F7FA91" box="[1096,1174,1360,1384]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-6.9166665">6°55’S</geoCoordinate>
|
||
<geoCoordinate id="EE21503BFFB93174FB28FAA0039CFA91" box="[1181,1277,1360,1384]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="37.666668">37°40’E</geoCoordinate>
|
||
,
|
||
<date id="FFAB103CFFB93174FABCFAA00214FA91" box="[1289,1397,1360,1384]" pageId="20" pageNumber="21" value="1926-09-09">9.ix.1926</date>
|
||
, A. Loveridge leg. (MCZ no. 43311). – 1 Ƥ Morogoro Region, Uluguru South Forest Reserve, forest above Ukwama Village,
|
||
<geoCoordinate id="EE21503BFFB93174FF4DFA660624FA57" box="[248,325,1430,1454]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-7.1833334">7°11’S</geoCoordinate>
|
||
<geoCoordinate id="EE21503BFFB93174FEFAFA6606CEFA57" box="[335,431,1430,1454]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="36.7">36°42’E</geoCoordinate>
|
||
, 1500–1600 masl,
|
||
<date id="FFAB103CFFB93174FD3FFA6605BBFA57" box="[650,730,1430,1454]" pageId="20" pageNumber="21" value="1993-10">x.1993</date>
|
||
, L. Sørensen leg. (ZMUC 00101085). – 1 Ƥ Uluguru Mountains, Kimboza Forest,
|
||
<geoCoordinate id="EE21503BFFB93174FE2EFA48068BFA29" box="[411,490,1464,1488]" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-7.0333333">7°02’S</geoCoordinate>
|
||
<geoCoordinate id="EE21503BFFB93174FE45FA480530FA29" box="[496,593,1464,1488]" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="36.783333">36°47’E</geoCoordinate>
|
||
, 250 masl,
|
||
<date id="FFAB103CFFB93174FD65FA480430FA29" box="[720,849,1464,1488]" pageId="20" pageNumber="21" value="1981-07-18">18.vii.1981</date>
|
||
, M. Stoltze & N. Scharff leg. (ZMUC 00101087).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F6577FFB93174FF72FA2A07B7F9A2" pageId="20" pageNumber="21" type="diagnosis">
|
||
<paragraph id="8BAA36FCFFB93174FF72FA2A07B7F9A2" blockId="20.[151,1436,979,1870]" pageId="20" pageNumber="21">
|
||
<emphasis id="B961EAEEFFB93174FF72FA2A0620FA0A" bold="true" box="[199,321,1498,1523]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
|
||
A species of
|
||
<taxonomicName id="4C154D7FFFB93174FE6CFA2B053DFA0A" box="[473,604,1499,1523]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B961EAEEFFB93174FE6CFA2B053DFA0A" box="[473,604,1499,1523]" italics="true" pageId="20" pageNumber="21">Hyperbolus</emphasis>
|
||
</taxonomicName>
|
||
in which the anterior gonopod coxite has a single mesal process, the anterior gonopod telopodite has an approximately semicircular mesal process directed straight mesad, and the apical lobe of the multiridged ‘mushroom-like’ structure of the posterior gonopod telopodite drawn out in to a smooth hook.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F6577FFB93176FF72F9950639FA81" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="8BAA36FCFFB93174FF72F9950429F91F" blockId="20.[151,1436,979,1870]" pageId="20" pageNumber="21">
|
||
<emphasis id="B961EAEEFFB93174FF72F9950638F987" bold="true" box="[199,345,1637,1662]" pageId="20" pageNumber="21">Description.</emphasis>
|
||
Adults mostly with 56 (9 out of 15 entire specimens), sometimes with 55 (
|
||
<specimenCount id="9D13FD75FFB93174FB12F9960255F987" box="[1191,1332,1638,1662]" pageId="20" pageNumber="21" type="generic">3 specimens</specimenCount>
|
||
) or 57 (
|
||
<specimenCount id="9D13FD75FFB93174FA3BF996066FF959" pageId="20" pageNumber="21" type="generic">3 specimens</specimenCount>
|
||
) podous rings, no apodous rings. Length
|
||
<quantity id="4CED9B19FFB93174FD68F9780431F959" box="[733,848,1672,1696]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="20" pageNumber="21" unit="cm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 cm</quantity>
|
||
. Width: 11.3–12.2 mm (3) / 11.9–13.9 mm (ƤƤ). Head, including eyes, antennae, mandibles, and gnathochilarium as in
|
||
<taxonomicName id="4C154D7FFFB93174FC13F95C0303F93A" box="[934,1122,1707,1731]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="apicomplexus">
|
||
<emphasis id="B961EAEEFFB93174FC13F95C0303F93A" box="[934,1122,1707,1731]" italics="true" pageId="20" pageNumber="21">H. apicomplexus</emphasis>
|
||
</taxonomicName>
|
||
. Each gnathochilarial stipes with 3 apical setae; each lamella lingualis with 4 apical setae.
|
||
</paragraph>
|
||
<paragraph id="8BAA36FCFFB93174FF70F9000436F8F1" blockId="20.[151,1436,979,1870]" box="[197,855,1776,1800]" pageId="20" pageNumber="21">
|
||
Collum, body rings, telson and legs as in
|
||
<taxonomicName id="4C154D7FFFB93174FD21F9010431F8F1" box="[660,848,1776,1800]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="apicomplexus">
|
||
<emphasis id="B961EAEEFFB93174FD21F9010431F8F1" box="[660,848,1776,1800]" italics="true" pageId="20" pageNumber="21">H. apicomplexus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAA36FCFFB93174FF72F8E303E7F8D2" blockId="20.[151,1436,979,1870]" box="[199,1158,1811,1835]" pageId="20" pageNumber="21">
|
||
<emphasis id="B961EAEEFFB93174FF72F8E30680F8D2" bold="true" box="[199,481,1811,1835]" pageId="20" pageNumber="21">Male sexual characters.</emphasis>
|
||
Ring VII and leg modifications as in
|
||
<taxonomicName id="4C154D7FFFB93174FC3DF8E3031EF8D2" box="[904,1151,1811,1835]" class="Diplopoda" family="Pachybolidae" genus="Crurifarcimen" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="vagans">
|
||
<emphasis id="B961EAEEFFB93174FC3DF8E3031EF8D2" box="[904,1151,1811,1835]" italics="true" pageId="20" pageNumber="21">Crurifarcimen vagans</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAA36FCFFB93174FF70F8C60350F8B7" blockId="20.[151,1436,979,1870]" box="[197,1073,1846,1870]" pageId="20" pageNumber="21">
|
||
Gonopods (
|
||
<figureCitation id="132E2A79FFB93174FEF2F8C606A6F8B7" box="[327,455,1846,1870]" captionStart-0="FIGURES 38 – 41. 38" captionStart-1="FIGURES 42 – 43" captionStartId-0="21.[151,264,1827,1850]" captionStartId-1="22.[151,264,1042,1065]" captionTargetBox-0="[189,1397,342,1801]" captionTargetBox-1="[209,1377,182,1024]" captionTargetId-0="figure@21.[184,1403,337,1806]" captionTargetId-1="figure@22.[208,1381,181,1031]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 38 – 41. 38: Hyperbolus morogoroensis, male from Ngambaula L. A. Forest Reserve, gonopod block, anterior view, G. Brovad phot., 39: do., posterior view, G. Brovad phot. s, 40: do., right anterior gonopod coxa, anterior view, SEM, 41: right anterior gonopod telopodite, anterior view (i. e., from inside of anterior gonopod cavity), SEM. Scales 1 cm. cx: coxa, fp: finger-shaped process, mp: mesal, thumb-like process of telopodite, st: sternum, tlp: telopodite." captionText-1="FIGURES 42 – 43. Hyperbolus morogoroensis. 42: male from Mkungwe Forest Reserve, tip of left posterior gonopod, oral view. 43: female from Mkungwe Forest Reserve, left vulva, posterior-lateral view. Scales 1 mm. ap: apical appendix, lav: subrectangular lobe of aboral valve, lov: subtriangular lobe of oral valve, op: operculum, slm: solenomerite." httpUri-0="https://zenodo.org/record/276704/files/figure.png" httpUri-1="https://zenodo.org/record/276705/files/figure.png" pageId="20" pageNumber="21">Figs 38–42</figureCitation>
|
||
) as in
|
||
<taxonomicName id="4C154D7FFFB93174FDA4F8C705ACF8B7" box="[529,717,1846,1870]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="apicomplexus">
|
||
<emphasis id="B961EAEEFFB93174FDA4F8C705ACF8B7" box="[529,717,1846,1870]" italics="true" pageId="20" pageNumber="21">H. apicomplexus</emphasis>
|
||
</taxonomicName>
|
||
, with the following exceptions:
|
||
</paragraph>
|
||
<paragraph id="8BAA36FCFFB93174FF22F88F03AAF827" blockId="20.[151,1436,1918,2014]" pageId="20" pageNumber="21">
|
||
- Anterior gonopod coxa (
|
||
<figureCitation id="132E2A79FFB93174FE6CF88E0538F86F" box="[473,601,1918,1942]" captionStart="FIGURES 38 – 41. 38" captionStartId="21.[151,264,1827,1850]" captionTargetBox="[189,1397,342,1801]" captionTargetId="figure@21.[184,1403,337,1806]" captionTargetPageId="21" captionText="FIGURES 38 – 41. 38: Hyperbolus morogoroensis, male from Ngambaula L. A. Forest Reserve, gonopod block, anterior view, G. Brovad phot., 39: do., posterior view, G. Brovad phot. s, 40: do., right anterior gonopod coxa, anterior view, SEM, 41: right anterior gonopod telopodite, anterior view (i. e., from inside of anterior gonopod cavity), SEM. Scales 1 cm. cx: coxa, fp: finger-shaped process, mp: mesal, thumb-like process of telopodite, st: sternum, tlp: telopodite." httpUri="https://zenodo.org/record/276704/files/figure.png" pageId="20" pageNumber="21">Figs 38, 40</figureCitation>
|
||
) with apical margin not sloping, almost straight, and with one rounded-triangular process at mesal margin just distal to sternum (
|
||
<emphasis id="B961EAEEFFB93174FC9FF8530413F843" box="[810,882,1955,1978]" italics="true" pageId="20" pageNumber="21">versus</emphasis>
|
||
two finger-shaped processes in
|
||
<taxonomicName id="4C154D7FFFB93174FB62F85302F2F843" box="[1239,1427,1954,1978]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="apicomplexus">
|
||
<emphasis id="B961EAEEFFB93174FB62F85302F2F843" box="[1239,1427,1954,1978]" italics="true" pageId="20" pageNumber="21">H. apicomplexus</emphasis>
|
||
</taxonomicName>
|
||
) - Overlapping lobes of anterior gonopod telopodites much smaller, sub-semicircular (
|
||
<figureCitation id="132E2A79FFB93174FBC4F83603A3F827" box="[1137,1218,1990,2014]" captionStart="FIGURES 38 – 41. 38" captionStartId="21.[151,264,1827,1850]" captionTargetBox="[189,1397,342,1801]" captionTargetId="figure@21.[184,1403,337,1806]" captionTargetPageId="21" captionText="FIGURES 38 – 41. 38: Hyperbolus morogoroensis, male from Ngambaula L. A. Forest Reserve, gonopod block, anterior view, G. Brovad phot., 39: do., posterior view, G. Brovad phot. s, 40: do., right anterior gonopod coxa, anterior view, SEM, 41: right anterior gonopod telopodite, anterior view (i. e., from inside of anterior gonopod cavity), SEM. Scales 1 cm. cx: coxa, fp: finger-shaped process, mp: mesal, thumb-like process of telopodite, st: sternum, tlp: telopodite." httpUri="https://zenodo.org/record/276704/files/figure.png" pageId="20" pageNumber="21">Fig. 39</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BAA36FCFFB83175FF22FF680682FEE5" blockId="21.[151,1436,151,284]" pageId="21" pageNumber="22">
|
||
- Terminal segment of posterior gonopods (
|
||
<figureCitation id="132E2A79FFB83175FD2FFF67058BFF56" box="[666,746,151,175]" captionStart="FIGURES 42 – 43" captionStartId="22.[151,264,1042,1065]" captionTargetBox="[209,1377,182,1024]" captionTargetId="figure@22.[208,1381,181,1031]" captionTargetPageId="22" captionText="FIGURES 42 – 43. Hyperbolus morogoroensis. 42: male from Mkungwe Forest Reserve, tip of left posterior gonopod, oral view. 43: female from Mkungwe Forest Reserve, left vulva, posterior-lateral view. Scales 1 mm. ap: apical appendix, lav: subrectangular lobe of aboral valve, lov: subtriangular lobe of oral valve, op: operculum, slm: solenomerite." httpUri="https://zenodo.org/record/276705/files/figure.png" pageId="21" pageNumber="22">Fig. 42</figureCitation>
|
||
) less mushroom-like; distal lobe of “hat” drawn out into slen- der hook; oral side of “hat” with parallel, strongly sclerotized ridges; field of ridges detached from surface basally and hiding basal lobe of “hat” from view; ridges diminishing in size and eventually disappearing towards tip of distal lobe.
|
||
</paragraph>
|
||
<caption id="DF6A6674FFB83175FF22F8D30321F86F" httpUri="https://zenodo.org/record/276704/files/figure.png" pageId="21" pageNumber="22" targetBox="[189,1397,342,1801]" targetPageId="21">
|
||
<paragraph id="8BAA36FCFFB83175FF22F8D30321F86F" blockId="21.[151,1436,1827,1942]" pageId="21" pageNumber="22">
|
||
<emphasis id="B961EAEEFFB83175FF22F8D30634F8C0" bold="true" box="[151,341,1827,1850]" pageId="21" pageNumber="22">FIGURES 38–41.</emphasis>
|
||
38:
|
||
<taxonomicName id="4C154D7FFFB83175FE36F8D405F7F8C3" box="[387,662,1828,1850]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="morogoroensis">
|
||
<emphasis id="B961EAEEFFB83175FE36F8D405F7F8C3" box="[387,662,1828,1850]" italics="true" pageId="21" pageNumber="22">Hyperbolus morogoroensis</emphasis>
|
||
</taxonomicName>
|
||
, male from Ngambaula L.A. Forest Reserve, gonopod block, anterior view, G. Brovad phot., 39: do., posterior view, G. Brovad phot.s, 40: do., right anterior gonopod coxa, anterior view, SEM, 41: right anterior gonopod telopodite, anterior view (i.e., from inside of anterior gonopod cavity), SEM. Scales 1 cm. cx: coxa, fp: finger-shaped process, mp: mesal, thumb-like process of telopodite, st: sternum, tlp: telopodite.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6A6674FFBB3176FF22FBE203DBFB9F" httpUri="https://zenodo.org/record/276705/files/figure.png" pageId="22" pageNumber="23" targetBox="[209,1377,182,1024]" targetPageId="22">
|
||
<paragraph id="8BAA36FCFFBB3176FF22FBE203DBFB9F" blockId="22.[151,1436,1042,1126]" pageId="22" pageNumber="23">
|
||
<emphasis id="B961EAEEFFBB3176FF22FBE20637FBD1" bold="true" box="[151,342,1042,1065]" pageId="22" pageNumber="23">FIGURES 42–43.</emphasis>
|
||
<emphasis id="B961EAEEFFBB3176FEEAFBE2051DFBD1" box="[351,636,1042,1064]" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="4C154D7FFFBB3176FEEAFBE20519FBD1" box="[351,632,1042,1064]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="morogoroensis">Hyperbolus morogoroensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
42: male from Mkungwe Forest Reserve, tip of left posterior gonopod, oral view. 43: female from Mkungwe Forest Reserve, left vulva, posterior-lateral view. Scales 1 mm. ap: apical appendix, lav: subrectangular lobe of aboral valve, lov: subtriangular lobe of oral valve, op: operculum, slm: solenomerite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAA36FCFFBB3176FF72FB7904D9FAF5" blockId="22.[151,1436,1160,1400]" pageId="22" pageNumber="23">
|
||
In some specimens, the posterior gonopod solonomerite has a row of small denticles.
|
||
<emphasis id="B961EAEEFFBB3176FF72FB5C0562FB3D" bold="true" box="[199,515,1196,1220]" pageId="22" pageNumber="23">Female sexual characters.</emphasis>
|
||
Distal margin of lateral coxosternal extensions projecting as right-angled lobe. Vulva (
|
||
<figureCitation id="132E2A79FFBB3176FF5EFB200657FB11" box="[235,310,1232,1256]" captionStart="FIGURES 42 – 43" captionStartId="22.[151,264,1042,1065]" captionTargetBox="[209,1377,182,1024]" captionTargetId="figure@22.[208,1381,181,1031]" captionTargetPageId="22" captionText="FIGURES 42 – 43. Hyperbolus morogoroensis. 42: male from Mkungwe Forest Reserve, tip of left posterior gonopod, oral view. 43: female from Mkungwe Forest Reserve, left vulva, posterior-lateral view. Scales 1 mm. ap: apical appendix, lav: subrectangular lobe of aboral valve, lov: subtriangular lobe of oral valve, op: operculum, slm: solenomerite." httpUri="https://zenodo.org/record/276705/files/figure.png" pageId="22" pageNumber="23">Fig.43</figureCitation>
|
||
) similar to those of
|
||
<taxonomicName id="4C154D7FFFBB3176FD94FB2105BEFB11" box="[545,735,1232,1256]" class="Diplopoda" family="Pachybolidae" genus="Hyperbolus" kingdom="Animalia" order="Spirobolida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="apicomplexus">
|
||
<emphasis id="B961EAEEFFBB3176FD94FB2105BEFB11" box="[545,735,1232,1256]" italics="true" pageId="22" pageNumber="23">H. apicomplexus</emphasis>
|
||
</taxonomicName>
|
||
but differing in having the subrectangular lobe on the aboral valve and the subtriangular lobe on the oral valve much less developed.
|
||
</paragraph>
|
||
<paragraph id="8BAA36FCFFBB3176FF72FAE70639FA81" blockId="22.[151,1436,1160,1400]" pageId="22" pageNumber="23">
|
||
<emphasis id="B961EAEEFFBB3176FF72FAE7064BFAC9" bold="true" box="[199,298,1303,1328]" pageId="22" pageNumber="23">Habitat.</emphasis>
|
||
Lowland forest and lowland submontane forest, found in litter and rotten logs.
|
||
<emphasis id="B961EAEEFFBB3176FF72FACB0601FAAD" bold="true" box="[199,352,1339,1364]" pageId="22" pageNumber="23">Distribution.</emphasis>
|
||
Only known from the Uluguru Mountains (
|
||
<figureCitation id="132E2A79FFBB3176FCE6FACC04C4FAAD" box="[851,933,1340,1364]" captionStart="FIGURE 84" captionStartId="36.[151,249,1479,1502]" captionTargetBox="[456,1120,410,1444]" captionTargetId="figure@36.[456,1122,407,1445]" captionTargetPageId="36" captionText="FIGURE 84. Known distribution of East African Pachybolini, except Hadrobolus crassicollis and Epibolus pulchripes. dots (●): Crurifarcimen vagans, filled diamonds (♦): Hyperbolus apicomplexus, open diamonds (♢): Hyperbolus morogoroensis, filled triangle (˔): Parabolus dimorphus, inverted open triangle (): Parabolus calceus, inverted filled triangle (ˑ): Pachybolus tectus,, filled square (․): Pachybolini indet. Based on Yahoo maps." httpUri="https://zenodo.org/record/276715/files/figure.png" pageId="22" pageNumber="23">Fig. 84</figureCitation>
|
||
) where it occurs from 250 (possibly 140) to 1500–1600 masl.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |