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<document id="ADA3FC85F7BA5AEA8E0859FE6271AF42" ID-CLB-Dataset="116156" ID-DOI="10.5281/zenodo.5039012" ID-GBIF-Dataset="fd4e444b-9dd5-4dc6-86a2-4b609136931d" ID-ISSN="0253-116X" ID-Zenodo-Dep="5039012" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624912002508" checkinUser="carolina" docAuthor="Assing, Volker" docDate="2021" docId="03BC87CCF35EFF8FFF40FCCD7267FDBF" docLanguage="en" docName="LinzerbiolBeitr.52.2.869-889.pdf" docOrigin="Linzer biologische Beiträge 52 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Cephennium vitsiense Assing 2021, nov.sp." docType="treatment" docVersion="8" lastPageNumber="876" masterDocId="FF85FFB4F358FF88FFD7FFA0730BFFA0" masterDocTitle="On the Cephennium fauna of Greece (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="889" masterPageNumber="869" pageNumber="875" updateTime="1698977987753" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="90FC4C7253350D63FF5DA53BC0761C48">On the Cephennium fauna of Greece (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="D6B28E553CFD413170FC02D8250F8BCC">Assing, Volker</mods:namePart>
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<mods:title id="E141AAD01DE177F094FB530BA7348B36">Linzer biologische Beiträge</mods:title>
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<mods:date id="E161096C9791C06E51AB4C886C7C706B">2021</mods:date>
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<mods:number id="38D79A49EA4D45869E17D5C4BD38C3B1">2021-02-28</mods:number>
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<mods:identifier id="5E0A7B3A53E84B2D1959C5D2D48F1581" type="DOI">10.5281/zenodo.5039012</mods:identifier>
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<treatment id="03BC87CCF35EFF8FFF40FCCD7267FDBF" ID-DOI="http://doi.org/10.5281/zenodo.6975185" ID-GBIF-Taxon="198858042" ID-Zenodo-Dep="6975185" LSID="urn:lsid:plazi:treatment:03BC87CCF35EFF8FFF40FCCD7267FDBF" httpUri="http://treatment.plazi.org/id/03BC87CCF35EFF8FFF40FCCD7267FDBF" lastPageId="7" lastPageNumber="876" pageId="6" pageNumber="875">
<subSubSection id="C30F6551F35EFF8EFF40FCCD7166FC23" box="[151,621,877,899]" pageId="6" pageNumber="875" type="nomenclature">
<paragraph id="8BAA36DAF35EFF8EFF40FCCD7166FC23" blockId="6.[151,621,877,899]" box="[151,621,877,899]" pageId="6" pageNumber="875">
<heading id="D0E281B6F35EFF8EFF40FCCD7166FC23" box="[151,621,877,899]" fontSize="8" level="2" pageId="6" pageNumber="875" reason="2">
<emphasis id="B961EAC8F35EFF8EFF40FCCD72DBFC22" bold="true" box="[151,464,877,899]" pageId="6" pageNumber="875">
<taxonomicName id="4C154D59F35EFF8EFF40FCCD7275FC22" ID-CoL="8XWYH" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" box="[151,382,877,899]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="875" phylum="Arthropoda" rank="species" species="vitsiense" status="sp. nov.">
<emphasis id="B961EAC8F35EFF8EFF40FCCD7275FC22" bold="true" box="[151,382,877,899]" italics="true" pageId="6" pageNumber="875">Cephennium vitsiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257B3F35EFF8EFE53FCCD72DBFC22" box="[388,464,877,898]" pageId="6" pageNumber="875" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132E2A5FF35EFF8EFE0AFCCD7114FC23" box="[477,543,877,899]" captionStart="Figs 6-10" captionStartId="14.[151,191,1499,1521]" captionTargetBox="[161,1059,203,1470]" captionTargetId="figure-38@14.[153,1059,203,1487]" captionTargetPageId="14" captionText="Figs 6-10: Male habitus of Cephennium vitsiense (6), C. catax (7), C. dramanum (8), C. fibuliferum (9), and C. cornutum (10). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5039018" httpUri="https://zenodo.org/record/5039018/files/figure.png" pageId="6" pageNumber="875">Figs 6</figureCitation>
,
<figureCitation id="132E2A5FF35EFF8EFDFDFCCD716DFC23" box="[554,614,877,899]" captionStart="Figs 20-29" captionStartId="16.[151,191,1499,1521]" captionTargetBox="[150,1054,207,1481]" captionTargetId="figure-62@16.[150,1057,201,1486]" captionTargetPageId="16" captionText="Figs 20-29: Cephennium pellucidum (20-22), C. brachati (23), C. vitsiense (24-25), C. vellicans (26-27), and C. catax (28-29): (20-24, 26, 28-29) aedeagus in lateral and in ventral view; (25, 27) internal structures of aedeagus. Scale bars: 20-24, 26, 28-29: 0.1 mm; 25, 27: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5039020" httpUri="https://zenodo.org/record/5039020/files/figure.png" pageId="6" pageNumber="875">24-25</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F6551F35EFF8EFF40FC3872D8FC7B" pageId="6" pageNumber="875" type="materials_examined">
<paragraph id="8BAA36DAF35EFF8EFF40FC3872D8FC7B" blockId="6.[151,1059,918,987]" pageId="6" pageNumber="875">
Type material:
<materialsCitation id="3B7D3C87F35EFF8EFE5BFC3772D8FC7B" ID-GBIF-Occurrence="3881968343" collectingDate="2005-05-22" collectionCode="cAss" collectorName="V. Assing" country="Greece" determinerName="V. Assing" elevation="1875" latitude="40.642223" location="Florina" longLatPrecision="19" longitude="21.379444" municipality="Florina" pageId="6" pageNumber="875" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54AE8878F35EFF8EFE5BFC3772EAFC0D" box="[396,481,919,941]" pageId="6" pageNumber="875" type="holotype">Holotype</typeStatus>
<specimenCount id="9D13FD53F35EFF8EFE31FC3872F1FC0D" box="[486,506,920,941]" pageId="6" pageNumber="875" type="male">♂</specimenCount>
: &quot;
<collectingCountry id="F302764AF35EFF8EFDC6FC37713FFC0C" box="[529,564,919,940]" pageId="6" pageNumber="875">GR</collectingCountry>
. [1]
<collectingMunicipality id="6BCEACA0F35EFF8EFD8AFC3771AAFC0D" box="[605,673,919,941]" pageId="6" pageNumber="875">Flórina</collectingMunicipality>
,
<locationDeviation id="9A182FF8F35EFF8EFD7CFC377012FC0C" bearing="202.5" box="[683,793,919,941]" distance="20000.0" pageId="6" pageNumber="875">
<quantity id="4CED9B3FF35EFF8EFD7CFC3771EFFC0D" box="[683,740,919,941]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="875" unit="km" value="20.0">20 km</quantity>
SSW
</locationDeviation>
<location id="8ECA6001F35EFF8EFCC9FC377069FC0D" LSID="urn:lsid:plazi:treatment:03BC87CCF35EFF8FFF40FCCD7267FDBF:8ECA6001F35EFF8EFCC9FC377069FC0D" box="[798,866,919,941]" country="GR" latitude="40.642223" longLatPrecision="19" longitude="21.379444" municipality="Florina" name="Florina" pageId="6" pageNumber="875">Flórina</location>
,
<location id="8ECA6001F35EFF8EFCBCFC3770C0FC0D" LSID="urn:lsid:plazi:treatment:03BC87CCF35EFF8FFF40FCCD7267FDBF:8ECA6001F35EFF8EFCBCFC3770C0FC0D" box="[875,971,919,941]" country="GR" latitude="40.642223" longLatPrecision="19" longitude="21.379444" municipality="Florina" name="Oros Vitsi" pageId="6" pageNumber="875">Oros Vitsi</location>
, N-slope,
<quantity id="4CED9B3FF35EFF8EFF40FC0E721DFC63" box="[151,278,942,964]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="1.9" metricValueMin="1.85" pageId="6" pageNumber="875" unit="m" value="1875.0" valueMax="1900.0" valueMin="1850.0">
<elevation id="0038D1E9F35EFF8EFF40FC0E721DFC63" box="[151,278,942,964]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="1.9" metricValueMin="1.85" pageId="6" pageNumber="875" unit="m" value="1875.0" valueMax="1900.0" valueMin="1850.0">1850-1900 m</elevation>
</quantity>
,
<geoCoordinate id="EE21501DF35EFF8EFEF6FC0E728EFC63" box="[289,389,942,963]" degrees="40" direction="north" minutes="38" orientation="latitude" pageId="6" pageNumber="875" precision="15" seconds="32" value="40.642223">40°38'32N</geoCoordinate>
,
<geoCoordinate id="EE21501DF35EFF8EFE46FC0E72FAFC64" box="[401,497,942,964]" degrees="21" direction="east" minutes="22" orientation="longitude" pageId="6" pageNumber="875" precision="15" seconds="46" value="21.379444">21°22'46E</geoCoordinate>
,
<date id="FFAB101AF35EFF8EFE2AFC0E7155FC64" box="[509,606,942,964]" pageId="6" pageNumber="875" value="2005-05-22">
<collectingDate id="EFEFE9F2F35EFF8EFE2AFC0E7155FC64" box="[509,606,942,964]" pageId="6" pageNumber="875" value="2005-05-22">22.V.2005</collectingDate>
</date>
<collectorName id="26E0530CF35EFF8EFDB2FC0E71C8FC64" box="[613,707,942,964]" pageId="6" pageNumber="875">V. Assing</collectorName>
/ Holotypus ♂
<originalDetermination id="A80B8E26F35EFF8EFC8EFC0E73C3FC7A" pageId="6" pageNumber="875">
<taxonomicName id="4C154D59F35EFF8EFC8EFC0E7729FC64" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" box="[857,1058,942,964]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="875" phylum="Arthropoda" rank="species" species="vitsiense" status="sp. nov.">
<emphasis id="B961EAC8F35EFF8EFC8EFC0E7729FC64" box="[857,1058,942,964]" italics="true" pageId="6" pageNumber="875">Cephennium vitsiense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257B3F35EFF8EFF40FC6573C3FC7A" box="[151,200,965,986]" pageId="6" pageNumber="875" rank="species">sp. n.</taxonomicNameLabel>
</originalDetermination>
det.
<determinerName id="7E76DA1EF35EFF8EFF23FC657244FC7B" box="[244,335,965,987]" pageId="6" pageNumber="875">V. Assing</determinerName>
2020&quot; (
<collectionCode id="ED04AE1FF35EFF8EFE4FFC6572C3FC7B" box="[408,456,965,987]" pageId="6" pageNumber="875">cAss</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30F6551F35EFF8EFF40FC497175FBBC" pageId="6" pageNumber="875" type="etymology">
<paragraph id="8BAA36DAF35EFF8EFF40FC497175FBBC" blockId="6.[151,1058,1001,1052]" pageId="6" pageNumber="875">
Etymology: The specific epithet is an adjective derived from Vitsi, the name of the mountain where the
<typeStatus id="54AE8878F35EFF8EFE5BFBA772B3FBBC" box="[396,440,1031,1052]" pageId="6" pageNumber="875">type</typeStatus>
locality is situated.
</paragraph>
</subSubSection>
<subSubSection id="C30F6551F35EFF8FFF40FB8B73F2FE84" lastPageId="7" lastPageNumber="876" pageId="6" pageNumber="875" type="description">
<paragraph id="8BAA36DAF35EFF8EFF40FB8B72FFFBFF" blockId="6.[151,1054,1067,1119]" pageId="6" pageNumber="875">
Description: Body length
<quantity id="4CED9B3FF35EFF8EFDDDFB8B7167FBE1" box="[522,620,1067,1089]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="875" unit="mm" value="1.25">1.25 mm</quantity>
; antenna
<quantity id="4CED9B3FF35EFF8EFD05FB8B703BFBE1" box="[722,816,1067,1089]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="875" unit="mm" value="0.55">0.55 mm</quantity>
long. Habitus (
<figureCitation id="132E2A5FF35EFF8EFC0EFB8B7713FBE1" box="[985,1048,1067,1089]" captionStart="Figs 6-10" captionStartId="14.[151,191,1499,1521]" captionTargetBox="[161,1059,203,1470]" captionTargetId="figure-38@14.[153,1059,203,1487]" captionTargetPageId="14" captionText="Figs 6-10: Male habitus of Cephennium vitsiense (6), C. catax (7), C. dramanum (8), C. fibuliferum (9), and C. cornutum (10). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5039018" httpUri="https://zenodo.org/record/5039018/files/figure.png" pageId="6" pageNumber="875">Fig. 6</figureCitation>
) moderately slender. Body reddish.
</paragraph>
<paragraph id="8BAA36DAF35EFF8EFF40FBCF71B4FB1F" blockId="6.[151,1058,1134,1215]" pageId="6" pageNumber="875">Eyes composed of approximately ten ommatidia. Antenna with antennomeres V approximately as long as broad, VII weakly transverse, IX and X barely 1.5 times as broad as long, and XI less than twice as long as broad.</paragraph>
<paragraph id="8BAA36DAF35EFF8EFF40FB6F73D3FABF" blockId="6.[151,1058,1230,1311]" pageId="6" pageNumber="875">Pronotum relatively weakly transverse, 1.12 times as broad as long, moderately convex in cross-section. Punctation rather sparse and very fine, approximately as fine as that of elytra.</paragraph>
<paragraph id="8BAA36DAF35EFF8EFF40FA8E72BEFADF" blockId="6.[151,1058,1326,1407]" pageId="6" pageNumber="875">Elytra broader than pronotum; humeral carinae weakly pronounced, nearly as long as the combined length of the basal four antennomeres; anterior impressions transverse and with tomentose pubescence.</paragraph>
<paragraph id="8BAA36DAF35EFF8EFF40FA2F7276FA5C" blockId="6.[151,1058,1422,1532]" pageId="6" pageNumber="875">
<specimenCount id="9D13FD53F35EFF8EFF40FA2F73A6FA04" box="[151,173,1423,1444]" pageId="6" pageNumber="875" type="male">♂</specimenCount>
: protibia apically distinctly bent and excavate; aedeagus (
<figureCitation id="132E2A5FF35EFF8EFCD0FA2E7072FA04" box="[775,889,1422,1444]" captionStart="Figs 20-29" captionStartId="16.[151,191,1499,1521]" captionTargetBox="[150,1054,207,1481]" captionTargetId="figure-62@16.[150,1057,201,1486]" captionTargetPageId="16" captionText="Figs 20-29: Cephennium pellucidum (20-22), C. brachati (23), C. vitsiense (24-25), C. vellicans (26-27), and C. catax (28-29): (20-24, 26, 28-29) aedeagus in lateral and in ventral view; (25, 27) internal structures of aedeagus. Scale bars: 20-24, 26, 28-29: 0.1 mm; 25, 27: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5039020" httpUri="https://zenodo.org/record/5039020/files/figure.png" pageId="6" pageNumber="875">Figs 24-25</figureCitation>
)
<quantity id="4CED9B3FF35EFF8EFC5EFA2E70EEFA04" box="[905,997,1422,1444]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="875" unit="mm" value="0.3">0.30 mm</quantity>
long; median lobe apically very acute in ventral view; internal sac with a pair of large clawshaped structures; parameres thin, straight, not reaching apex of median lobe, and with rather long apical seta.
</paragraph>
<paragraph id="8BAA36DAF35EFF8FFF40F9AB73F2FE84" blockId="6.[151,1058,1547,1599]" lastBlockId="7.[151,1058,211,292]" lastPageId="7" lastPageNumber="876" pageId="6" pageNumber="875">
Comparative notes: Based on the similar shape of the aedeagus and the similar shapes and arrangement of its internal structures,
<taxonomicName id="4C154D59F35EFF8EFD30F9897056F99E" authorityName="Assing" authorityYear="2021" box="[743,861,1577,1598]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="875" phylum="Arthropoda" rank="species" species="vitsiense">
<emphasis id="B961EAC8F35EFF8EFD30F9897056F99E" box="[743,861,1577,1598]" italics="true" pageId="6" pageNumber="875">C. vitsiense</emphasis>
</taxonomicName>
is closely allied to
<emphasis id="B961EAC8F35FFF8FFF40FF74721EFF48" box="[151,277,211,233]" italics="true" pageId="7" pageNumber="876">C. angelinii</emphasis>
. It is distinguished from this species by the apically much more acute median lobe, distinctly claw-shaped internal structures, and the modifications of the male protibiae.
</paragraph>
</subSubSection>
<subSubSection id="C30F6551F35FFF8FFF40FE937119FE01" pageId="7" pageNumber="876" type="distribution">
<paragraph id="8BAA36DAF35FFF8FFF40FE937119FE01" blockId="7.[151,1059,307,417]" pageId="7" pageNumber="876">
Distribution and natural history: The type locality is situated in Oros Vitsi, a mountain rich in endemics to the south-southwest of Flórina, Northwest
<collectingCountry id="F302764AF35FFF8FFF40FECE73E9FE23" box="[151,226,366,387]" name="Greece" pageId="7" pageNumber="876">Greece</collectingCountry>
. The
<typeStatus id="54AE8878F35FFF8FFEF6FECE7271FE24" box="[289,378,366,388]" pageId="7" pageNumber="876" type="holotype">holotype</typeStatus>
was sifted from grass and moss in the alpine zone of the peak region at an altitude of
<quantity id="4CED9B3FF35FFF8FFE54FE2B7105FE01" box="[387,526,395,417]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="1.9" metricValueMin="1.85" pageId="7" pageNumber="876" unit="m" value="1875.0" valueMax="1900.0" valueMin="1850.0">1850-1900 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F6551F35FFF8FFF40FE117267FDBF" pageId="7" pageNumber="876" type="discussion">
<paragraph id="8BAA36DAF35FFF8FFF40FE117267FDBF" blockId="7.[151,1059,433,543]" pageId="7" pageNumber="876">
Comment: The
<typeStatus id="54AE8878F35FFF8FFE5DFE1172E7FE67" box="[394,492,433,455]" pageId="7" pageNumber="876" type="holotype">holotypes</typeStatus>
of this and the following species were returned by Miroslav Stevanović to the second author with
<typeStatus id="54AE8878F35FFF8FFD7BFE6F7008FE44" box="[684,771,463,484]" pageId="7" pageNumber="876" type="paratype">paratype</typeStatus>
labels reading &quot;
<typeStatus id="54AE8878F35FFF8FFC6BFE6E7729FE43" box="[956,1058,462,483]" pageId="7" pageNumber="876" type="paratype">Paratypus</typeStatus>
<taxonomicName id="4C154D59F35FFF8FFF40FE4B721CFDA0" authorityName="MULLER &amp; KUNZE" authorityYear="1822" box="[151,279,491,512]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="889" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B961EAC8F35FFF8FFF40FE4B721CFDA0" box="[151,279,491,512]" italics="true" pageId="7" pageNumber="876">Cephennium</emphasis>
</taxonomicName>
(s.str.)
<emphasis id="B961EAC8F35FFF8FFEBCFE4C72E3FDA1" box="[363,488,492,513]" italics="true" pageId="7" pageNumber="876">acuminatum</emphasis>
m., det. M. Stevanović 2013&quot;. This species, however, was never described.
</paragraph>
</subSubSection>
</treatment>
</document>
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