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<document id="C783E636E378C6FFA6D0B51FDBDE005B" ID-DOI="10.11646/zootaxa.4991.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5042656" ID-ZooBank="F0FAE07E-4A90-4602-BCC5-6D80E158BB6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624989159860" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. &amp; Sangpradub, Narumon" docDate="2021" docId="03BC87C1FFEF224E0CC07AB4FC6C54D4" docLanguage="en" docName="zootaxa.4991.3.8.pdf" docOrigin="Zootaxa 4991 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patrus apicalis" docType="treatment" docVersion="3" lastPageNumber="566" masterDocId="FF85FFB9FFEA224B0C577A23FFDF537D" masterDocTitle="Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)" masterLastPageNumber="591" masterPageNumber="561" pageNumber="566" updateTime="1698978743576" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="0ED5D90511B4BC3AFC0F379A18122BAA">
<mods:title id="25D750EFCB1FCECDA3A16E4030E24A9E">Review of the Patrus landaisi species group from Thailand with description of three new species (Coleoptera, Gyrinidae)</mods:title>
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<mods:namePart id="81724AC3C1BD997F2F77974105EF2317">Suksai, Benjamart</mods:namePart>
<mods:nameIdentifier id="084332DB3168A0EEC568D732809639CF" type="ORCID">0000-0002-2472-6100</mods:nameIdentifier>
<mods:nameIdentifier id="5572DD39D27756139F301B082F941DBF" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="DFC1FC7C6E470E0BC87EA0BD8A8A0168">Gustafson, Grey T.</mods:namePart>
<mods:affiliation id="2A1045FD5FF68691CB1BE675BE9BA8C0">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. &amp; Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
</mods:name>
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<mods:namePart id="C7D4B25BB86091C2436B0BE4C489B2A9">Sites, Robert W.</mods:namePart>
<mods:nameIdentifier id="4CBE527FEFB73EDE80B176EA2E1AEA7E" type="ORCID">0000-0002-3895-813X</mods:nameIdentifier>
<mods:affiliation id="93AAF2F0461127183E7991920F091528">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
</mods:name>
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<mods:role id="B89AD0B125988C678852E34B97C8FF13">
<mods:roleTerm id="BA23B9945688C67282451E6C17E69BC3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7AFBAA1D27634643AB1437B1E57BF077">Sangpradub, Narumon</mods:namePart>
<mods:nameIdentifier id="2E02E2C9B5B2139E854D206B4B85DB15" type="ORCID">0000-0002-2472-6100</mods:nameIdentifier>
<mods:affiliation id="392AEAFAB9B55E3BAC354D74D771A10E">Applied Taxonomic Research Center, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand &amp; Corresponding Author: narumon @ kku. ac. th; https: // orcid. org / 0000 - 0003 - 0228 - 8696</mods:affiliation>
<mods:nameIdentifier id="33D71A6D7DDBFA44A74F2A1AA2EB957C" type="email">benjamartbenjamart@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="A178DEAA5683199BD10AD11A608D9A60">text</mods:typeOfResource>
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<mods:title id="2CB750C9BE6431E7A34CB6E8E1650ED0">Zootaxa</mods:title>
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<mods:part id="9B78D439976FB53E0EBA9BD852A53ED2">
<mods:date id="E6F90A9F10972A20B57FA643D1596C79">2021</mods:date>
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<mods:number id="F9F0CCE3E862E8A0FC1CF9F295815D35">2021-06-25</mods:number>
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<mods:number id="DCBF9AFFB5402045731C9F8EA56E5853">4991</mods:number>
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<mods:number id="A84ADFEFA2F332BBA9EF9324EC512CE0">3</mods:number>
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<mods:identifier id="B07CD817FA75AE038089221A3B89ACB2" type="DOI">10.11646/zootaxa.4991.3.8</mods:identifier>
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<mods:identifier id="F99381D31F2B122BD6A5F90BB967A570" type="Zenodo-Dep">5042656</mods:identifier>
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<treatment id="03BC87C1FFEF224E0CC07AB4FC6C54D4" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4" httpUri="http://treatment.plazi.org/id/03BC87C1FFEF224E0CC07AB4FC6C54D4" lastPageNumber="566" pageId="5" pageNumber="566">
<subSubSection id="C30F655CFFEF224E0CC07AB4FDE953CC" box="[151,566,150,178]" pageId="5" pageNumber="566" type="nomenclature">
<paragraph id="8BAA36D7FFEF224E0CC07AB4FDE953CC" blockId="5.[151,566,150,214]" box="[151,566,150,178]" pageId="5" pageNumber="566">
<heading id="D0E281BBFFEF224E0CC07AB4FDE953CC" bold="true" box="[151,566,150,178]" fontSize="11" level="1" pageId="5" pageNumber="566" reason="1">
<taxonomicName id="4C154D54FFEF224E0CC07AB4FDE953CC" authority="(Regimbart, 1891)" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[151,566,150,178]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFEF224E0CC07AB4FDE953CC" bold="true" box="[151,566,150,178]" pageId="5" pageNumber="566">
<emphasis id="B961EAC5FFEF224E0CC07AB4FE9453CF" bold="true" box="[151,331,151,178]" italics="true" pageId="5" pageNumber="566">Patrus apicalis</emphasis>
(
<bibRefCitation id="EF844B26FFEF224E0D0C7AB5FDF153CC" author="Regimbart, M." box="[347,558,150,177]" pageId="5" pageNumber="566" pagination="537 - 554" refId="ref23003" refString="Regimbart, M. (1891) Viaggio di Leonardo Fea in Birmania e regioni vicine. XXXIII. Enumeration des Dytiscidae et Gyrinidae recueillis par M. Leonardo Fea, dans ses voyages en Birmanie et regions voisines. Annali del Museo Civico di Storia Naturale Giacomo Doria Genova, 10 (2), 537 - 554." type="journal article" year="1891">Régimbart, 1891</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFEF224E0CC07A98FDDE53A8" box="[151,513,187,214]" pageId="5" pageNumber="566" type="description">
<paragraph id="8BAA36D7FFEF224E0CC07A98FDDE53A8" blockId="5.[151,566,150,214]" box="[151,513,187,214]" pageId="5" pageNumber="566">
(
<figureCitation id="132E2A52FFEF224E0CC87A98FEDC53A8" box="[159,259,187,213]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="5" pageNumber="566">Figs. 2A</figureCitation>
,
<figureCitation id="132E2A52FFEF224E0D587A98FEEC53A8" box="[271,307,187,213]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="5" pageNumber="566">3A</figureCitation>
,
<figureCitation id="132E2A52FFEF224E0D687A98FEBB53A8" box="[319,356,187,213]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="5" pageNumber="566">4A</figureCitation>
,
<figureCitation id="132E2A52FFEF224E0D387A98FEA053AB" box="[367,383,187,214]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">5</figureCitation>
,
<figureCitation id="132E2A52FFEF224E0DDC7A98FE6353A8" box="[395,444,187,213]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="5" pageNumber="566">13B</figureCitation>
,
<figureCitation id="132E2A52FFEF224E0D9F7A98FE2653A8" box="[456,505,187,213]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="5" pageNumber="566">14A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFEF224E0CC07B23FCC3524B" pageId="5" pageNumber="566" type="reference_group">
<paragraph id="8BAA36D7FFEF224E0CC07B23FCEE5265" blockId="5.[151,817,255,310]" box="[151,817,255,280]" pageId="5" pageNumber="566">
<treatmentCitationGroup id="AB0511F9FFEF224E0CC07B23FCEE5265" box="[151,817,255,280]" pageId="5" pageNumber="566">
<treatmentCitation id="0AB410C6FFEF224E0CC07B23FD8B526A" author="Regimbart, M." box="[151,596,255,280]" page="548" pageId="5" pageNumber="566" year="1891">
<taxonomicName id="4C154D54FFEF224E0CC07B23FD8B526A" authority="Regimbart, 1891: 548" authorityName="Regimbart" authorityPageNumber="548" authorityYear="1891" box="[151,596,255,280]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFEF224E0CC07B23FEB05265" box="[151,367,256,280]" italics="true" pageId="5" pageNumber="566">Orectochilus apicalis</emphasis>
<bibRefCitation id="EF844B26FFEF224E0D217B23FD8B526A" author="Regimbart, M." box="[374,596,255,280]" pageId="5" pageNumber="566" pagination="537 - 554" refId="ref23003" refString="Regimbart, M. (1891) Viaggio di Leonardo Fea in Birmania e regioni vicine. XXXIII. Enumeration des Dytiscidae et Gyrinidae recueillis par M. Leonardo Fea, dans ses voyages en Birmanie et regions voisines. Annali del Museo Civico di Storia Naturale Giacomo Doria Genova, 10 (2), 537 - 554." type="journal article" year="1891">Régimbart, 1891: 548</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA36D7FFEF224E0CC07B3DFCC3524B" blockId="5.[151,817,255,310]" box="[151,796,286,310]" pageId="5" pageNumber="566">
<treatmentCitationGroup id="AB0511F9FFEF224E0CC07B3DFCC3524B" box="[151,796,286,310]" pageId="5" pageNumber="566">
<taxonomicName id="4C154D54FFEF224E0CC07B3DFDA1524B" authority="Regimbart, 1891" authorityName="Regimbart" authorityYear="1891" box="[151,638,286,310]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis" subGenus="Patrus">
<emphasis id="B961EAC5FFEF224E0CC07B3DFEC6524B" box="[151,281,286,310]" italics="true" pageId="5" pageNumber="566">Orectochilus</emphasis>
(
<emphasis id="B961EAC5FFEF224E0D7F7B3CFEB3524B" box="[296,364,287,310]" italics="true" pageId="5" pageNumber="566">Patrus</emphasis>
)
<emphasis id="B961EAC5FFEF224E0D2D7B3DFE15524B" box="[378,458,286,310]" italics="true" pageId="5" pageNumber="566">apicalis</emphasis>
<bibRefCitation id="EF844B26FFEF224E0D867B3DFDA1524B" author="Regimbart, M." box="[465,638,286,310]" pageId="5" pageNumber="566" pagination="537 - 554" refId="ref23003" refString="Regimbart, M. (1891) Viaggio di Leonardo Fea in Birmania e regioni vicine. XXXIII. Enumeration des Dytiscidae et Gyrinidae recueillis par M. Leonardo Fea, dans ses voyages en Birmanie et regions voisines. Annali del Museo Civico di Storia Naturale Giacomo Doria Genova, 10 (2), 537 - 554." type="journal article" year="1891">Régimbart, 1891</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="0AB410C6FFEF224E0EDF7B3DFCC3524B" author="Ochs" box="[648,796,286,310]" page="20" pageId="5" pageNumber="566" year="1930">
<bibRefCitation id="EF844B26FFEF224E0EDF7B3DFCC3524B" author="Ochs" box="[648,796,286,310]" pageId="5" pageNumber="566" pagination="1 - 39" refId="ref22521" refString="Ochs, G. (1930) Gyrinoidea. Catalogue of Indian Insects, 19, 1 - 39." type="journal article" year="1930">Ochs 1930: 20</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFEF224E0CC07B7CFA435054" pageId="5" pageNumber="566" type="materials_examined">
<paragraph id="8BAA36D7FFEF224E0CC07B7CFDE05207" blockId="5.[151,1437,351,1962]" box="[151,575,351,378]" pageId="5" pageNumber="566">
<materialsCitation id="3B7D3C8AFFEF224E0CC07B7CFDE45207" box="[151,571,351,378]" country="Myanmar" location="Karen Hills" pageId="5" pageNumber="566" specimenCount="1" typeStatus="holotype">
<emphasis id="B961EAC5FFEF224E0CC07B7CFEE95204" bold="true" box="[151,310,351,377]" pageId="5" pageNumber="566">
<typeStatus id="54AE8875FFEF224E0CC07B7CFF0D5204" box="[151,210,351,377]" pageId="5" pageNumber="566">Type</typeStatus>
locality.
</emphasis>
<location id="8ECA600CFFEF224E0D6B7B43FE1E5204" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4:8ECA600CFFEF224E0D6B7B43FE1E5204" box="[316,449,351,378]" country="Myanmar" name="Karen Hills" pageId="5" pageNumber="566">Karen Hills</location>
,
<collectingCountry id="F3027647FFEF224E0D9B7B43FDE45207" box="[460,571,352,378]" name="Myanmar" pageId="5" pageNumber="566">Myanmar</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAA36D7FFEF224E0C907BA0FA4E52BC" blockId="5.[151,1437,351,1962]" pageId="5" pageNumber="566">
<emphasis id="B961EAC5FFEF224E0C907BA0FE2C52E0" bold="true" box="[199,499,387,413]" pageId="5" pageNumber="566">Type material examined.</emphasis>
<typeStatus id="54AE8875FFEF224E0DAC7BA0FD5052E0" box="[507,655,387,413]" pageId="5" pageNumber="566" type="holotype">HOLOTYPE</typeStatus>
female ‘Carin / Asciui Ghecù /
<quantity id="4CED9B32FFEF224E08567BA0FB7E52E3" box="[1025,1185,387,414]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="5" pageNumber="566" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">1200–1400 m</quantity>
. / L.Fea III–88 [beige label, handwritten black ink]’ MSNG
<typeStatus id="54AE8875FFEF224E0E687B84FD7852BC" box="[575,679,423,449]" pageId="5" pageNumber="566" type="holotype">Holotype</typeStatus>
DSCN0243. Additional label information not provided to authors.
</paragraph>
<paragraph id="8BAA36D7FFEF224E0C907BE8FA435054" blockId="5.[151,1437,351,1962]" pageId="5" pageNumber="566">
<emphasis id="B961EAC5FFEF224E0C907BE8FDD75298" bold="true" box="[199,520,459,485]" pageId="5" pageNumber="566">Other material examined.</emphasis>
<materialsCitation id="3B7D3C8AFFEF224E0E427BE8FDC95150" collectingDate="2017-05-15" collectionCode="THNHM" collectorName="B. Suksai &amp; C. Jaitamart &amp; T. Paotajai" country="Thailand" county="Mae Kam Pong Stream" elevation="1011" latitude="18.866196" location="Mae On District" longLatPrecision="1" longitude="99.35261" municipality="Huay Kaew Subdistrict" pageId="5" pageNumber="566" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai">
<emphasis id="B961EAC5FFEF224E0E427BE8FC085298" bold="true" box="[533,983,459,485]" pageId="5" pageNumber="566">
<collectingCountry id="F3027647FFEF224E0E427BE8FD6F5298" box="[533,688,459,485]" name="Thailand" pageId="5" pageNumber="566">THAILAND</collectingCountry>
:
<collectingRegion id="49D1F835FFEF224E0E957BE8FC0D5298" box="[706,978,459,485]" country="Thailand" name="Chiang Mai" pageId="5" pageNumber="566">Chiang Mai Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62CB4E5BFFEF224E0FB37BEFFB205298" box="[996,1279,459,486]" pageId="5" pageNumber="566">Mae Kam Pong Stream</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFEF224E09467BEFFECC5174" pageId="5" pageNumber="566">Huay Kaew Subdistrict</collectingMunicipality>
,
<location id="8ECA600CFFEF224E0D767BD3FE015174" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4:8ECA600CFFEF224E0D767BD3FE015174" box="[289,478,495,522]" country="Thailand" county="Mae Kam Pong Stream" latitude="18.866196" longLatPrecision="1" longitude="99.35261" municipality="Huay Kaew Subdistrict" name="Mae On District" pageId="5" pageNumber="566" stateProvince="Chiang Mai">Mae On District</location>
,
<geoCoordinate id="EE215010FFEF224E0DBC7BCCFD565177" box="[491,649,495,522]" degrees="18" direction="north" minutes="51" orientation="latitude" pageId="5" pageNumber="566" precision="1" seconds="58.3" value="18.866196">18°51’58.3”N</geoCoordinate>
<geoCoordinate id="EE215010FFEF224E0EC57BCCFCF05174" box="[658,815,495,521]" degrees="99" direction="east" minutes="21" orientation="longitude" pageId="5" pageNumber="566" precision="1" seconds="09.4" value="99.35261">99°21’09.4”E</geoCoordinate>
,
<elevation id="0038D1E4FFEF224E0F6A7BCCFC0D5174" box="[829,978,495,522]" metricMagnitude="3" metricUnit="m" metricValue="1.011" pageId="5" pageNumber="566" unit="m" value="1011.0">
<quantity id="4CED9B32FFEF224E0F6A7BCCFC4C5177" box="[829,915,495,522]" metricMagnitude="3" metricUnit="m" metricValue="1.011" pageId="5" pageNumber="566" unit="m" value="1011.0">1011 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFEF224E0FB77BCCFB895177" box="[992,1110,495,522]" pageId="5" pageNumber="566" value="2017-05-15">
<collectingDate id="EFEFE9FFFFEF224E0FB77BCCFB895177" box="[992,1110,495,522]" pageId="5" pageNumber="566" value="2017-05-15">15.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFEF224E08C27BD3FADA5174" box="[1173,1285,495,522]" pageId="5" pageNumber="566">B. Suksai</collectorName>
,
<collectorName id="26E05301FFEF224E09447BCCFA435174" box="[1299,1436,495,521]" pageId="5" pageNumber="566">C. Jaitamart</collectorName>
&amp;
<collectorName id="26E05301FFEF224E0CE17837FEEC5150" box="[182,307,531,558]" pageId="5" pageNumber="566">T. Paotajai</collectorName>
,
<specimenCount id="9D13FD5EFFEF224E0D167830FE515150" box="[321,398,531,557]" pageId="5" pageNumber="566" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFEF224E0DF77830FDD25150" box="[416,525,531,557]" pageId="5" pageNumber="566">THNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D3C8AFFEF224E0E757830FE685108" collectingDate="2017-05-25" collectionCode="KKU" collectorName="P. Sapewisut" country="Thailand" county="Nam Kuen Stream" elevation="1327" latitude="19.122862" location="Mae Chedi Subdistrict" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" pageId="5" pageNumber="566" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Chiang Rai">
<emphasis id="B961EAC5FFEF224E0E757830FCF55150" bold="true" box="[546,810,531,557]" pageId="5" pageNumber="566">
<collectingRegion id="49D1F835FFEF224E0E757830FCFA5150" box="[546,805,531,557]" country="Thailand" name="Chiang Rai" pageId="5" pageNumber="566">Chiang Rai Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62CB4E5BFFEF224E0F647837FBD35150" box="[819,1036,531,558]" pageId="5" pageNumber="566">Nam Kuen Stream</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFEF224E084D7837FACF5153" box="[1050,1296,531,558]" pageId="5" pageNumber="566">Ban Huai Nam Kuen</collectingMunicipality>
,
<location id="8ECA600CFFEF224E09497837FECC512C" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4:8ECA600CFFEF224E09497837FECC512C" country="Thailand" county="Nam Kuen Stream" latitude="19.122862" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" name="Mae Chedi Subdistrict" pageId="5" pageNumber="566" stateProvince="Chiang Rai">Mae Chedi Subdistrict</location>
,
<location id="8ECA600CFFEF224E0D757814FDF7512C" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4:8ECA600CFFEF224E0D757814FDF7512C" box="[290,552,567,594]" country="Thailand" county="Nam Kuen Stream" latitude="19.122862" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" name="Wiang Pa Pao District" pageId="5" pageNumber="566" stateProvince="Chiang Rai">Wiang Pa Pao District</location>
,
<geoCoordinate id="EE215010FFEF224E0E617814FD0A512C" box="[566,725,567,593]" degrees="19" direction="north" minutes="07" orientation="latitude" pageId="5" pageNumber="566" precision="1" seconds="22.3" value="19.122862">19°07’22.3”N</geoCoordinate>
<geoCoordinate id="EE215010FFEF224E0E887814FCA3512C" box="[735,892,567,593]" degrees="99" direction="east" minutes="19" orientation="longitude" pageId="5" pageNumber="566" precision="1" seconds="43.3" value="99.3287">99°19’43.3”E</geoCoordinate>
,
<elevation id="0038D1E4FFEF224E0FDC7814FBFB512C" box="[907,1060,567,594]" metricMagnitude="3" metricUnit="m" metricValue="1.327" pageId="5" pageNumber="566" unit="m" value="1327.0">
<quantity id="4CED9B32FFEF224E0FDC7814FC3B512F" box="[907,996,567,594]" metricMagnitude="3" metricUnit="m" metricValue="1.327" pageId="5" pageNumber="566" unit="m" value="1327.0">1327 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFEF224E08637814FB76512F" box="[1076,1193,567,594]" pageId="5" pageNumber="566" value="2017-05-25">
<collectingDate id="EFEFE9FFFFEF224E08637814FB76512F" box="[1076,1193,567,594]" pageId="5" pageNumber="566" value="2017-05-25">25.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFEF224E08BB781BFA5F512C" box="[1260,1408,567,594]" pageId="5" pageNumber="566">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFEF224E09D97814FF045108" pageId="5" pageNumber="566" type="male">3 males</specimenCount>
,
<specimenCount id="9D13FD5EFFEF224E0CBD7878FE855108" box="[234,346,603,629]" pageId="5" pageNumber="566" type="female">3 females</specimenCount>
(
<collectionCode id="ED04AE12FFEF224E0D3B7878FE705108" box="[364,431,603,629]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="5" pageNumber="566">KKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFEF224E0D91787FFD1451E4" collectingDate="2018-11-11" collectionCode="GTGC, KKU, THNHM, UMC" collectorName="B. Suksai &amp; P. Sapewisut &amp; S. Kullasoot" country="Thailand" county="Nam Kuen Stream" elevation="1327" latitude="19.122862" location="Mae Chedi Subdistrict" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" pageId="5" pageNumber="566" specimenCount="11" specimenCount-female="7" specimenCount-male="4" stateProvince="Chiang Rai">
same locality except for,
<date id="FFAB1017FFEF224E0EB97878FCB05108" box="[750,879,603,629]" pageId="5" pageNumber="566" value="2018-11-11">
<collectingDate id="EFEFE9FFFFEF224E0EB97878FCB05108" box="[750,879,603,629]" pageId="5" pageNumber="566" value="2018-11-11">11.XI.2018</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFEF224E0FE5787FFBFC5108" box="[946,1059,603,630]" pageId="5" pageNumber="566">B. Suksai</collectorName>
,
<collectorName id="26E05301FFEF224E0865787FFB1C5108" box="[1074,1219,603,630]" pageId="5" pageNumber="566">P. Sapewisut</collectorName>
&amp;
<collectorName id="26E05301FFEF224E08B97878FA5F5108" box="[1262,1408,603,629]" pageId="5" pageNumber="566">S. Kullasoot</collectorName>
,
<specimenCount id="9D13FD5EFFEF224E09D97878FF0451E4" pageId="5" pageNumber="566" type="male">4 males</specimenCount>
,
<specimenCount id="9D13FD5EFFEF224E0CB078A3FE8C51E4" box="[231,339,639,666]" pageId="5" pageNumber="566" type="female">7 females</specimenCount>
(
<collectionCode id="ED04AE12FFEF224E0D35785CFE6C51E4" box="[354,435,639,665]" pageId="5" pageNumber="566">GTGC</collectionCode>
,
<collectionCode id="ED04AE12FFEF224E0DE978A3FE2251E7" box="[446,509,640,666]" country="Thailand" httpUri="http://grbio.org/cool/pwjq-e3rs" name="Herbarium, Department of Biology, Khon Kaen University" pageId="5" pageNumber="566">KKU</collectionCode>
,
<collectionCode id="ED04AE12FFEF224E0E5E78A3FDAC51E7" box="[521,627,640,666]" pageId="5" pageNumber="566">THNHM</collectionCode>
,
<collectionCode id="ED04AE12FFEF224E0E29785CFD1C51E4" box="[638,707,639,665]" pageId="5" pageNumber="566">UMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFEF224E0E8178A3FDEA51C0" collectingDate="2019-05-03" collectionCode="THNHM" collectorName="B. Suksai &amp; P. Sapewisut &amp; N. Plangklang" country="Thailand" county="Nam Kuen Stream" elevation="1327" latitude="19.122862" location="Mae Chedi Subdistrict" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" pageId="5" pageNumber="566" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Rai">
same locality except for,
<date id="FFAB1017FFEF224E0FA6785CFB8751E4" box="[1009,1112,639,665]" pageId="5" pageNumber="566" value="2019-05-03">
<collectingDate id="EFEFE9FFFFEF224E0FA6785CFB8751E4" box="[1009,1112,639,665]" pageId="5" pageNumber="566" value="2019-05-03">3.V.2019</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFEF224E08C378A3FADD51E4" box="[1172,1282,639,666]" pageId="5" pageNumber="566">B. Suksai</collectorName>
,
<collectorName id="26E05301FFEF224E095978A3FA4351E4" box="[1294,1436,639,666]" pageId="5" pageNumber="566">P. Sapewisut</collectorName>
&amp;
<collectorName id="26E05301FFEF224E0CE27887FE8651C0" box="[181,345,675,702]" pageId="5" pageNumber="566">N. Plangklang</collectorName>
,
<specimenCount id="9D13FD5EFFEF224E0D327880FE6E51C0" box="[357,433,675,701]" pageId="5" pageNumber="566" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFEF224E0D977880FDF251C0" box="[448,557,675,701]" pageId="5" pageNumber="566">THNHM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B7D3C8AFFEF224E0E167880FC5151C0" box="[577,910,675,701]" collectingDate="2019-05-03" collectionCode="THNHM, GTGC" collectorName="B. Suksai &amp; P. Sapewisut &amp; N. Plangklang" country="Thailand" county="Nam Kuen Stream" elevation="1327" isTailEnumeration="true" latitude="19.122862" location="Mae Chedi Subdistrict" longLatPrecision="1" longitude="99.3287" municipality="Ban Huai Nam Kuen" pageId="5" pageNumber="566" specimenCount="4" specimenCount-female="4" stateProvince="Chiang Rai">
<specimenCount id="9D13FD5EFFEF224E0E167880FD7151C0" box="[577,686,675,701]" pageId="5" pageNumber="566" type="female">4 females</specimenCount>
(
<collectionCode id="ED04AE12FFEF224E0EEA7880FCF751C0" box="[701,808,675,701]" pageId="5" pageNumber="566">THNHM</collectionCode>
,
<collectionCode id="ED04AE12FFEF224E0F637880FC5A51C0" box="[820,901,675,701]" pageId="5" pageNumber="566">GTGC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D3C8AFFEF224E0FCE7880FDDD5078" collectingDate="2017-05-29" collectionCode="THNHM" collectorName="P. Sapewisut" country="Thailand" county="Khun Sathan Natl. Park." elevation="786" latitude="18.252344" location="Mueng Li Subdistrict" longLatPrecision="1" longitude="100.49326" municipality="Ban Mueng Li" pageId="5" pageNumber="566" specimenCount="1" specimenCount-male="1" stateProvince="Nan">
<emphasis id="B961EAC5FFEF224E0FCE7880FB9A51C0" bold="true" box="[921,1093,675,701]" pageId="5" pageNumber="566">
<collectingRegion id="49D1F835FFEF224E0FCE7880FB9F51C0" box="[921,1088,675,701]" country="Thailand" name="Nan" pageId="5" pageNumber="566">Nan Province</collectingRegion>
:
</emphasis>
<collectingCounty id="62CB4E5BFFEF224E081A7880FABC51C0" box="[1101,1379,675,701]" pageId="5" pageNumber="566">Khun Sathan Natl. Park.</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFEF224E09387887FED6519C" pageId="5" pageNumber="566">Ban Mueng Li</collectingMunicipality>
,
<location id="8ECA600CFFEF224E0D4278EBFDD7519C" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4:8ECA600CFFEF224E0D4278EBFDD7519C" box="[277,520,711,738]" country="Thailand" county="Khun Sathan Natl. Park." latitude="18.252344" longLatPrecision="1" longitude="100.49326" municipality="Ban Mueng Li" name="Mueng Li Subdistrict" pageId="5" pageNumber="566" stateProvince="Nan">Mueng Li Subdistrict</location>
,
<location id="8ECA600CFFEF224E0E4378EBFD03519C" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4:8ECA600CFFEF224E0E4378EBFD03519C" box="[532,732,711,738]" country="Thailand" county="Khun Sathan Natl. Park." latitude="18.252344" longLatPrecision="1" longitude="100.49326" municipality="Ban Mueng Li" name="Na Muen District" pageId="5" pageNumber="566" stateProvince="Nan">Na Muen District</location>
,
<geoCoordinate id="EE215010FFEF224E0EBF78E4FC4B519F" box="[744,916,711,738]" degrees="18" direction="north" minutes="15" orientation="latitude" pageId="5" pageNumber="566" precision="1" seconds="08.44" value="18.252344">18°15’08.44”N</geoCoordinate>
<geoCoordinate id="EE215010FFEF224E0FCC78E4FB8A519F" box="[923,1109,711,738]" degrees="100" direction="east" minutes="29" orientation="longitude" pageId="5" pageNumber="566" precision="1" seconds="35.77" value="100.49326">100°29’35.77”E</geoCoordinate>
,
<elevation id="0038D1E4FFEF224E083678E4FB3A519C" box="[1121,1253,711,738]" metricMagnitude="2" metricUnit="m" metricValue="7.859999999999999" pageId="5" pageNumber="566" unit="m" value="786.0">
<quantity id="4CED9B32FFEF224E083678E4FB77519F" box="[1121,1192,711,738]" metricMagnitude="2" metricUnit="m" metricValue="7.859999999999999" pageId="5" pageNumber="566" unit="m" value="786.0">786 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFEF224E08A578E4FAB8519C" box="[1266,1383,711,737]" pageId="5" pageNumber="566" value="2017-05-29">
<collectingDate id="EFEFE9FFFFEF224E08A578E4FAB8519C" box="[1266,1383,711,737]" pageId="5" pageNumber="566" value="2017-05-29">29.V.2017</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFEF224E0CC078CFFEF85078" box="[151,295,747,774]" pageId="5" pageNumber="566">P. Sapewisut</collectorName>
,
<specimenCount id="9D13FD5EFFEF224E0D6578C8FEA25078" box="[306,381,747,773]" pageId="5" pageNumber="566" type="male">1 male</specimenCount>
(
<collectionCode id="ED04AE12FFEF224E0DDB78C8FE265078" box="[396,505,747,773]" pageId="5" pageNumber="566">THNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D3C8AFFEF224E0E5B78CFFA475054" collectingDate="2013-01-03" collectionCode="NHMW" collectorName="K. Rattanachan" country="Thailand" county="Mae Charim Natl. Park." elevation="461" latitude="18.812939" location="Mae Charim Subdistrict" longLatPrecision="1" longitude="101.0695" municipality="Huay Nampangyaekkwar" pageId="5" pageNumber="566" specimenCount="3" specimenCount-female="3" stateProvince="Nan">
<collectingCounty id="62CB4E5BFFEF224E0E5B78CFFCC45078" box="[524,795,747,774]" pageId="5" pageNumber="566">Mae Charim Natl. Park.</collectingCounty>
,
<collectingMunicipality id="6BCEACADFFEF224E0F7178CFFB9A5078" box="[806,1093,747,774]" pageId="5" pageNumber="566">Huay Nampangyaekkwar</collectingMunicipality>
,
<location id="8ECA600CFFEF224E080778CFFA805078" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4:8ECA600CFFEF224E080778CFFA805078" box="[1104,1375,747,774]" country="Thailand" county="Mae Charim Natl. Park." latitude="18.812939" longLatPrecision="1" longitude="101.0695" municipality="Huay Nampangyaekkwar" name="Mae Charim Subdistrict" pageId="5" pageNumber="566" stateProvince="Nan">Mae Charim Subdistrict</location>
,
<location id="8ECA600CFFEF224E093D78CFFE965054" LSID="urn:lsid:plazi:treatment:03BC87C1FFEF224E0CC07AB4FC6C54D4:8ECA600CFFEF224E093D78CFFE965054" country="Thailand" county="Mae Charim Natl. Park." latitude="18.812939" longLatPrecision="1" longitude="101.0695" municipality="Huay Nampangyaekkwar" name="Mae Charim District" pageId="5" pageNumber="566" stateProvince="Nan">Mae Charim District</location>
,
<geoCoordinate id="EE215010FFEF224E0D03792CFDDF5057" box="[340,512,783,810]" degrees="18" direction="north" minutes="48" orientation="latitude" pageId="5" pageNumber="566" precision="1" seconds="46.58" value="18.812939">18°48’46.58”N</geoCoordinate>
<geoCoordinate id="EE215010FFEF224E0E50792CFD1F5054" box="[519,704,783,809]" degrees="101" direction="east" minutes="04" orientation="longitude" pageId="5" pageNumber="566" precision="1" seconds="10.21" value="101.0695">101°04’10.21”E</geoCoordinate>
,
<elevation id="0038D1E4FFEF224E0E9B792CFC8F5054" box="[716,848,783,810]" metricMagnitude="2" metricUnit="m" metricValue="4.61" pageId="5" pageNumber="566" unit="m" value="461.0">
<quantity id="4CED9B32FFEF224E0E9B792CFCCC5057" box="[716,787,783,810]" metricMagnitude="2" metricUnit="m" metricValue="4.61" pageId="5" pageNumber="566" unit="m" value="461.0">461 m</quantity>
a.s.l.
</elevation>
,
<date id="FFAB1017FFEF224E0F0B792CFC635054" box="[860,956,783,809]" pageId="5" pageNumber="566" value="2013-01-03">
<collectingDate id="EFEFE9FFFFEF224E0F0B792CFC635054" box="[860,956,783,809]" pageId="5" pageNumber="566" value="2013-01-03">3.I.2013</collectingDate>
</date>
, leg.
<collectorName id="26E05301FFEF224E0FAF7933FB785054" box="[1016,1191,783,810]" pageId="5" pageNumber="566">K. Rattanachan</collectorName>
,
<specimenCount id="9D13FD5EFFEF224E08E5792CFAC15054" box="[1202,1310,783,809]" pageId="5" pageNumber="566" type="female">3 females</specimenCount>
(
<collectionCode id="ED04AE12FFEF224E097A792CFA505054" box="[1325,1423,783,809]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="566" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFEF224E0C907910FCFE5464" pageId="5" pageNumber="566" type="diagnosis">
<paragraph id="8BAA36D7FFEF224E0C907910FE2F563B" blockId="5.[151,1437,351,1962]" pageId="5" pageNumber="566">
<emphasis id="B961EAC5FFEF224E0C907910FE9D5030" bold="true" box="[199,322,819,845]" pageId="5" pageNumber="566">Diagnosis.</emphasis>
Size: Male: 13.0–14.0 mm, female: 10.0–
<quantity id="4CED9B32FFEF224E0F427910FCA55033" box="[789,890,819,846]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="5" pageNumber="566" unit="mm" value="10.9">10.9 mm</quantity>
; body form in lateral view weakly dorsoventrally convex; elytral glabrous region of male (
<figureCitation id="132E2A52FFEF224E0E317974FD14500C" box="[614,715,855,881]" captionStart="FIGURE 2" captionStartId="6.[151,260,1861,1887]" captionTargetBox="[151,1434,179,1836]" captionTargetId="figure-25@6.[151,1435,179,1836]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042660" httpUri="https://zenodo.org/record/5042660/files/figure.png" pageId="5" pageNumber="566">Figs. 2A</figureCitation>
,
<figureCitation id="132E2A52FFEF224E0E8F7974FD24500F" box="[728,763,855,882]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">5A</figureCitation>
) cordiform, along suture extending ca. 7/9 length of elytra, laterally expanded in anterior 2/3 to considerably wider than pronotal glabrous region, posterior 1/3 strongly acuminate with rounded margins; female elytral glabrous region (
<figureCitation id="132E2A52FFEF224E0F2D79BCFC0150C4" box="[890,990,927,953]" captionStart="FIGURE 3" captionStartId="7.[151,260,1865,1891]" captionTargetBox="[151,1434,182,1841]" captionTargetId="figure-18@7.[151,1435,179,1841]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of females of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 4 mm." figureDoi="http://doi.org/10.5281/zenodo.5042662" httpUri="https://zenodo.org/record/5042662/files/figure.png" pageId="5" pageNumber="566">Figs. 3A</figureCitation>
,
<figureCitation id="132E2A52FFEF224E0FBC79BCFBD450C7" box="[1003,1035,927,954]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">5E</figureCitation>
) dissimilar to male, broad for nearentire length occupying 10/11 elytral width, lateral margins nearly-straight for 3/4 length creating parallel-sided appearance, abruptly acuminate medially in posterior 1/4, posterolateral corners produced into points, posterior margin evenly convex toward suture; in lateral view width of pubescent margin at humeral region nearly equal to that extending posterior; elytral apex of both sexes similar, obliquely truncate without apicolateral sinuation, epipleural angle distinct, weakly rounded; male protibia wedge-shaped (
<figureCitation id="132E2A52FFEF224E0F0F7E70FC6B5710" box="[856,948,1107,1133]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="5" pageNumber="566">Fig. 4A</figureCitation>
), without basal constriction, protibial apex obliquely truncate with or without prominent distolateral angle; male protarsus ca. 3/4 length of protibia (
<figureCitation id="132E2A52FFEF224E09627E54FA4F57EC" box="[1333,1424,1143,1169]" captionStart="FIGURE 4" captionStartId="8.[151,260,707,733]" captionTargetBox="[158,1432,185,677]" captionTargetId="figure-25@8.[152,1436,179,683]" captionTargetPageId="8" captionText="FIGURE 4. Left protibia and protarsus of males of P. landaisi species group: (A) P. apicalis; (B) P. corpulentus; (C) P. garuda sp. nov.; (D) P. landaisi; (E) P. nanensis sp. nov.; (F) P. phetchabunensis sp. nov.; (G) P. salvazai; (H) P. subapicalis. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5042666" httpUri="https://zenodo.org/record/5042666/files/figure.png" pageId="5" pageNumber="566">Fig. 4A</figureCitation>
), protarsomeres 1–4 mostly similar in width; mesoventrite of male with medial pit; aedeagus (
<figureCitation id="132E2A52FFEF224E08D97EB8FADA57CB" box="[1166,1285,1179,1206]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5B–D</figureCitation>
) with median lobe in basal 2/3 weakly tapered, weakly constricted medially, parallel-sided in apical 1/3 before acumination, apex strongly constricted to a needle-like point extending 1/5 length, very weakly constricted in lateral view (
<figureCitation id="132E2A52FFEF224E09387EC0FF64565F" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5C</figureCitation>
); female reproductive tract (
<figureCitation id="132E2A52FFEF224E0E5F7F24FDBC565F" box="[520,611,1287,1314]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5G</figureCitation>
) without additional sclerotized structure between gonocoxae (
<figureCitation id="132E2A52FFEF224E09617F24FA51565F" box="[1334,1422,1287,1314]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5F</figureCitation>
), fertilization duct as in
<figureCitation id="132E2A52FFEF224E0DC47F08FE34563B" box="[403,491,1323,1350]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5H</figureCitation>
.
</paragraph>
<paragraph id="8BAA36D7FFEF224E0C907F6CFCFE5464" blockId="5.[151,1437,351,1962]" pageId="5" pageNumber="566">
<emphasis id="B961EAC5FFEF224E0C907F6CFE1A5614" bold="true" box="[199,453,1359,1385]" pageId="5" pageNumber="566">Differential diagnosis</emphasis>
.
<taxonomicName id="4C154D54FFEF224E0D847F73FDA25614" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[467,637,1359,1385]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFEF224E0D847F73FDA25614" box="[467,637,1359,1385]" italics="true" pageId="5" pageNumber="566">Patrus apicalis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C154D54FFEF224E0F1F7F73FC3C5614" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[840,995,1359,1385]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFEF224E0F1F7F73FC3C5614" box="[840,995,1359,1385]" italics="true" pageId="5" pageNumber="566">P. subapicalis</emphasis>
</taxonomicName>
, as the latter was previously considered to be a subspecies of the former.
<taxonomicName id="4C154D54FFEF224E0DA87F57FDAC56F0" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[511,627,1395,1421]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFEF224E0DA87F57FDAC56F0" box="[511,627,1395,1421]" italics="true" pageId="5" pageNumber="566">P. apicalis</emphasis>
</taxonomicName>
can be distinguished by the mesoventrite of the male having a medial pit, which is absent in both sexes of
<taxonomicName id="4C154D54FFEF224E0DA97FBBFD4656CC" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[510,665,1431,1457]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFEF224E0DA97FBBFD4656CC" box="[510,665,1431,1457]" italics="true" pageId="5" pageNumber="566">P. subapicalis</emphasis>
</taxonomicName>
. Males of these two species can be also distinguished by the shape of the median lobe of the aedeagus, which in
<taxonomicName id="4C154D54FFEF224E0E3C7F9FFD0156A8" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[619,734,1467,1493]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFEF224E0E3C7F9FFD0156A8" box="[619,734,1467,1493]" italics="true" pageId="5" pageNumber="566">P. apicalis</emphasis>
</taxonomicName>
has the apex with a more needle-like point (
<figureCitation id="132E2A52FFEF224E08937F98FAC456AB" box="[1220,1307,1467,1494]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5B</figureCitation>
), compared to
<taxonomicName id="4C154D54FFEF224E0CE37FC3FE905684" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[180,335,1503,1529]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFEF224E0CE37FC3FE905684" box="[180,335,1503,1529]" italics="true" pageId="5" pageNumber="566">P. subapicalis</emphasis>
</taxonomicName>
in which the apex is relatively less constricted and has a triangular tip (
<figureCitation id="132E2A52FFEF224E08397FFCFB0B5684" box="[1134,1236,1503,1529]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="5" pageNumber="566">Fig. 12B</figureCitation>
). Females of both species are similar, but can be separated by the shape of the elytral glabrous region, wherein the lateral margin in
<taxonomicName id="4C154D54FFEF224E09DF7C27FF2F553C" baseAuthorityName="Regimbart" baseAuthorityYear="1891" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFEF224E09DF7C27FF2F553C" italics="true" pageId="5" pageNumber="566">P. apicalis</emphasis>
</taxonomicName>
appears more rounded for 3/4 of its length (
<figureCitation id="132E2A52FFEF224E0EB17C04FCE2553F" box="[742,829,1575,1602]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5E</figureCitation>
), compared to that of
<taxonomicName id="4C154D54FFEF224E086F7C0BFB0B553C" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[1080,1236,1575,1601]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFEF224E086F7C0BFB0B553C" box="[1080,1236,1575,1601]" italics="true" pageId="5" pageNumber="566">P. subapicalis</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A52FFEF224E08B37C04FA97553C" box="[1252,1352,1575,1601]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="5" pageNumber="566">Fig. 12E</figureCitation>
) which has a nearly straight lateral margin that creates a more parallel-sided appearance to the elytral glabrous region. The posterior margin of the elytral glabrous region further differs between the two species, having a greater concavity in
<taxonomicName id="4C154D54FFEF224E0CE17CB7FEF355D0" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[182,300,1683,1709]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFEF224E0CE17CB7FEF355D0" box="[182,300,1683,1709]" italics="true" pageId="5" pageNumber="566">P. apicalis</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A52FFEF224E0D6B7CB0FE4C55D3" box="[316,403,1683,1710]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5E</figureCitation>
) relative to
<taxonomicName id="4C154D54FFEF224E0E487CB7FD6355D0" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" baseAuthorityName="Ochs" baseAuthorityYear="1930" box="[543,700,1683,1709]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="subapicalis">
<emphasis id="B961EAC5FFEF224E0E487CB7FD6355D0" box="[543,700,1683,1709]" italics="true" pageId="5" pageNumber="566">P. subapicalis</emphasis>
</taxonomicName>
(
<figureCitation id="132E2A52FFEF224E0E9B7CB0FCEC55D0" box="[716,819,1683,1709]" captionStart="FIGURE 12" captionStartId="24.[151,260,1869,1895]" captionTargetBox="[162,1426,899,1836]" captionTargetId="figure-324@24.[151,1436,893,1845]" captionTargetPageId="24" captionText="FIGURE 12. Patrus subapicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042688" httpUri="https://zenodo.org/record/5042688/files/figure.png" pageId="5" pageNumber="566">Fig. 12E</figureCitation>
). The female of
<taxonomicName id="4C154D54FFEF224E0FA67CB7FBB855D0" baseAuthorityName="Regimbart" baseAuthorityYear="1891" box="[1009,1127,1683,1709]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="B961EAC5FFEF224E0FA67CB7FBB855D0" box="[1009,1127,1683,1709]" italics="true" pageId="5" pageNumber="566">P. apicalis</emphasis>
</taxonomicName>
is also similar to that of
<taxonomicName id="4C154D54FFEF224E09DF7CB7FF2655AC" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
<emphasis id="B961EAC5FFEF224E09DF7CB7FF2655AC" italics="true" pageId="5" pageNumber="566">P. nanensis</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFEF224E0D567C94FE8355AC" bold="true" box="[257,348,1719,1745]" pageId="5" pageNumber="566">
<taxonomicNameLabel id="A25257BEFFEF224E0D567C94FE8355AC" box="[257,348,1719,1745]" pageId="5" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but can be separated from the latter by the posterior margin of the elytral glabrous region being more rounded in appearance (
<figureCitation id="132E2A52FFEF224E0DB57CF8FDE9558B" box="[482,566,1755,1782]" captionStart="FIGURE 5" captionStartId="8.[151,260,1867,1893]" captionTargetBox="[161,1430,873,1833]" captionTargetId="figure-85@8.[151,1436,863,1843]" captionTargetPageId="8" captionText="FIGURE 5. Patrus apicalis: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042668" httpUri="https://zenodo.org/record/5042668/files/figure.png" pageId="5" pageNumber="566">Fig. 5E</figureCitation>
) compared to
<taxonomicName id="4C154D54FFEF224E0E817CFFFC8C5588" authority="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub, 2021" authorityName="Suksai &amp; Gustafson &amp; Sites &amp; Sangpradub" authorityYear="2021" box="[726,851,1756,1781]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="nanensis" status="sp. nov.">
<emphasis id="B961EAC5FFEF224E0E817CFFFC8C5588" box="[726,851,1756,1781]" italics="true" pageId="5" pageNumber="566">P. nanensis</emphasis>
</taxonomicName>
<emphasis id="B961EAC5FFEF224E0F0D7CF8FC6C5588" bold="true" box="[858,947,1755,1781]" pageId="5" pageNumber="566">
<taxonomicNameLabel id="A25257BEFFEF224E0F0D7CF8FC6C5588" box="[858,947,1755,1781]" pageId="5" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
which has the posterior margin with a more oblique and angular appearance to its convexity (
<figureCitation id="132E2A52FFEF224E0EEA7CDCFCCA5464" box="[701,789,1791,1817]" captionStart="FIGURE 9" captionStartId="18.[151,260,1173,1199]" captionTargetBox="[172,1413,183,1149]" captionTargetId="figure-25@18.[164,1423,178,1158]" captionTargetPageId="18" captionText="FIGURE 9. Patrus nanensis sp. nov.: (A) male pronotum and elytra, dorsal view, scale bar = 4 mm; (B) aedeagus, dorsal view, scale bar = 1 mm; (C) apex of median lobe, lateral view, scale bar = 1 mm; (D) apex of median lobe, ventral view showing operculum, scale bar = 1 mm; (E) female pronotum and elytra, dorsal view, scale bar = 4 mm; (F) gonocoxae, ventral view; (G) female reproductive tract, ventral view, bursal gland not shown, scale bar = 1 mm; (H) fertilization duct, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5042682" httpUri="https://zenodo.org/record/5042682/files/figure.png" pageId="5" pageNumber="566">Fig. 9E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F655CFFEF224E0C907D00FC6C54D4" pageId="5" pageNumber="566" type="distribution">
<paragraph id="8BAA36D7FFEF224E0C907D00FC6C54D4" blockId="5.[151,1437,351,1962]" pageId="5" pageNumber="566">
<emphasis id="B961EAC5FFEF224E0C907D00FEBE5440" bold="true" box="[199,353,1827,1853]" pageId="5" pageNumber="566">Distribution.</emphasis>
Previously known from the Karen Hills region of
<collectingCountry id="F3027647FFEF224E0FF67D07FBD15443" box="[929,1038,1828,1854]" name="Myanmar" pageId="5" pageNumber="566">Myanmar</collectingCountry>
and northern
<collectingCountry id="F3027647FFEF224E08F97D07FB3B5443" box="[1198,1252,1828,1854]" name="Laos" pageId="5" pageNumber="566">Laos</collectingCountry>
(
<bibRefCitation id="EF844B26FFEF224E08A37D00FA505440" author="Peschet, R." box="[1268,1423,1827,1853]" pageId="5" pageNumber="566" pagination="121 - 136" refId="ref22628" refString="Peschet, R. (1923) Fauna entomologique de l'Indochine Gyrinidae. Opuscules de l'Institut Scientifique de l'Indochine Saigon, 6, 121 - 136." type="journal article" year="1923">Peschet 1923</bibRefCitation>
), this species is here recorded from
<collectingCountry id="F3027647FFEF224E0E4E7D64FDA2541C" box="[537,637,1863,1889]" name="Thailand" pageId="5" pageNumber="566">Thailand</collectingCountry>
for the first time. Within
<collectingCountry id="F3027647FFEF224E0FF77D64FBD9541C" box="[928,1030,1863,1889]" name="Thailand" pageId="5" pageNumber="566">Thailand</collectingCountry>
, it can be collected in slow recesses of high elevation streams (
<figureCitation id="132E2A52FFEF224E0D9E7D48FDF054F8" box="[457,559,1899,1925]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="5" pageNumber="566">Fig. 13B</figureCitation>
) across the northwestern mountains (
<figureCitation id="132E2A52FFEF224E0F8C7D48FB9A54F8" box="[987,1093,1899,1925]" captionStart="FIGURE 14" captionStartId="26.[151,260,743,769]" captionTargetBox="[152,1436,179,719]" captionTargetId="figure-25@26.[151,1436,179,719]" captionTargetPageId="26" captionText="FIGURE 14. (A–B) Distribution records of the P. landaisi species group in Thailand." figureDoi="http://doi.org/10.5281/zenodo.5042692" httpUri="https://zenodo.org/record/5042692/files/figure.png" pageId="5" pageNumber="566">Fig. 14A</figureCitation>
). This species has been found together with
<taxonomicName id="4C154D54FFEF224E0D647DB3FE7554D4" baseAuthorityName="Ochs" baseAuthorityYear="1940" box="[307,426,1935,1961]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="salvazai">
<emphasis id="B961EAC5FFEF224E0D647DB3FE7554D4" box="[307,426,1935,1961]" italics="true" pageId="5" pageNumber="566">P. salvazai</emphasis>
</taxonomicName>
in streams in
<collectingRegion id="49D1F835FFEF224E0E107DACFCEE54D4" box="[583,817,1935,1961]" country="Thailand" name="Chiang Rai" pageId="5" pageNumber="566">Chiang Rai Province</collectingRegion>
(
<figureCitation id="132E2A52FFEF224E0F177DACFC7954D4" box="[832,934,1935,1961]" captionStart="FIGURE 13" captionStartId="25.[151,260,1683,1709]" captionTargetBox="[169,1418,181,1656]" captionTargetId="figure-18@25.[169,1418,179,1659]" captionTargetPageId="25" captionText="FIGURE 13. Habitats of P. landaisi species group: (A) stream at Siriphum Waterfall, Doi Inthanon National Park, Chiang Mai Province, habitat of P. garuda sp. nov. and P. subapicalis; (B) Benjamart Suksai in habitat of P. apicalis and P. salvazai at Nam Kuen stream, Chiang Rai Province; (C) shallow stream close to Ban Thai Samakkhi, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (D) Tham Yai Waterfall Phu Kradueng National Park, Loei Province, habitat of P. phetchabunensis sp. nov.; (E) puddle in dry stream bed close to Ban Phithak Thai, Chiang Rai Province, habitat of P. landaisi and P. salvazai; (F) P. landaisi and P. salvazai from puddle in dry stream habitat close to Ban Thai Samakkhi, Chiang Rai Province." figureDoi="http://doi.org/10.5281/zenodo.5042690" httpUri="https://zenodo.org/record/5042690/files/figure.png" pageId="5" pageNumber="566">Fig. 13B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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