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<document id="7211D87CBAB4946DE4D05B187EC202B7" ID-CLB-Dataset="45794" ID-DOI="10.11646/zootaxa.3795.3.10" ID-GBIF-Dataset="33103d18-c977-4091-8b45-fed6896e1e88" ID-ISSN="1175-5326" ID-Zenodo-Dep="227971" ID-ZooBank="84469758-B2C9-4357-B048-4029B2494EB9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460419170663" checkinUser="plazi" docAuthor="Xu, Peng &amp; Li, Xinzheng" docDate="2014" docId="03BC8797FF83CD67FF4DF8FAFD6B1662" docLanguage="en" docName="zt03795p400.pdf" docOrigin="Zootaxa 3795 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thor hainanensis Xu &amp; Li, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="399" masterDocId="FF85FFEFFF83CD62FFDAFFFAFFEA136F" masterDocTitle="A new species of the hippolytid shrimp genus Thor Kingsley, 1878 (Crustacea: Decapoda: Caridea) from Hainan Island, China" masterLastPageNumber="400" masterPageNumber="394" pageNumber="394" updateTime="1698329533363" updateUser="ExternalLinkService">
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<mods:title id="E4E6807B104CFEC4EF00B820379764BA">A new species of the hippolytid shrimp genus Thor Kingsley, 1878 (Crustacea: Decapoda: Caridea) from Hainan Island, China</mods:title>
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<mods:namePart id="5A271C10D347314C7AD32588EAB6C396">Xu, Peng</mods:namePart>
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<mods:namePart id="728865735343C057037A7C17481D9A84">Li, Xinzheng</mods:namePart>
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<treatment id="03BC8797FF83CD67FF4DF8FAFD6B1662" ID-DOI="http://doi.org/10.5281/zenodo.6138680" ID-GBIF-Taxon="119426453" ID-Zenodo-Dep="6138680" LSID="urn:lsid:plazi:treatment:03BC8797FF83CD67FF4DF8FAFD6B1662" httpUri="http://treatment.plazi.org/id/03BC8797FF83CD67FF4DF8FAFD6B1662" lastPageId="5" lastPageNumber="399" pageId="0" pageNumber="394">
<subSubSection id="C30F650AFF83CD62FF4DF8FAFEE01452" pageId="0" pageNumber="394" type="nomenclature">
<paragraph id="8BAA3681FF83CD62FF4DF8FAFE2E1476" blockId="0.[151,452,1792,1853]" box="[151,452,1792,1817]" pageId="0" pageNumber="394">
<heading id="D0E281EDFF83CD62FF4DF8FAFE2E1476" bold="true" box="[151,452,1792,1817]" fontSize="10" level="2" pageId="0" pageNumber="394" reason="2">
<emphasis id="B961EA93FF83CD62FF4DF8FAFE2E1476" bold="true" box="[151,452,1792,1817]" pageId="0" pageNumber="394">
<taxonomicName id="4C154D02FF83CD62FF4DF8FAFE891476" box="[151,355,1792,1817]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="394" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="B961EA93FF83CD62FF4DF8FAFE891476" bold="true" box="[151,355,1792,1817]" italics="true" pageId="0" pageNumber="394">Thor hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257E8FF83CD62FEB0F8FAFE2E1476" box="[362,452,1792,1817]" pageId="0" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA3681FF83CD62FF4DF8DEFEE01452" blockId="0.[151,452,1792,1853]" box="[151,266,1828,1853]" pageId="0" pageNumber="394">
(
<figureCitation id="132E2A04FF83CD62FF45F8DEFEEB1452" box="[159,257,1828,1853]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,250,1955,1977]" captionStartId-1="2.[151,250,1880,1902]" captionStartId-2="3.[151,250,1058,1080]" captionStartId-3="3.[151,250,1945,1967]" captionStartId-4="4.[151,250,857,879]" captionTargetBox-0="[161,1428,1212,1924]" captionTargetBox-1="[210,1380,213,1832]" captionTargetBox-2="[160,1431,201,1036]" captionTargetBox-3="[411,1175,1161,1923]" captionTargetBox-4="[151,1436,193,835]" captionTargetId-0="figure@1.[151,1436,1201,1940]" captionTargetId-1="figure@2.[195,1395,193,1853]" captionTargetId-2="figure@3.[151,1436,193,1049]" captionTargetId-3="figure@3.[411,1175,1161,1924]" captionTargetId-4="figure@4.[151,1436,193,836]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionTargetPageId-4="4" captionText-0="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." captionText-1="FIGURE 2. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm." captionText-2="FIGURE 3. Thor hainanensis sp. nov., paratype, male (cl 1.9 mm). A, right third pereopod, lateral view; B, right fourth pereopod, lateral view; C, endopod of second pleopod, ventral view. Scale bars: 1 mm." captionText-3="FIGURE 4. Thor hainanensis sp. nov., paratype, male (cl 1.9 mm). A, propodus and dactylus of third pereopod, lateral view; B, distal third of propodus and dactylus of third pereopod, lateral view; C, propodus and dactylus of fourth pereopod, lateral view; B, distal third of propodus and dactylus of fourth pereopod, lateral view. Scale bars: A, C (500 µm); B, D (200 µm)." captionText-4="FIGURE 5. Thor hainanensis sp. nov., paratype, ovigerous female (cl 2.0 mm). A, anterior part of body, lateral view; B, key zone indicates the epipod of first pereopod, lateral view. Scale bars: A (1 mm); B (400 µm)." httpUri-0="https://zenodo.org/record/227972/files/figure.png" httpUri-1="https://zenodo.org/record/227973/files/figure.png" httpUri-2="https://zenodo.org/record/227974/files/figure.png" httpUri-3="https://zenodo.org/record/227975/files/figure.png" httpUri-4="https://zenodo.org/record/227976/files/figure.png" pageId="0" pageNumber="394">Figs 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F650AFF83CD63FF4DF896FE431397" lastPageId="1" lastPageNumber="395" pageId="0" pageNumber="394" type="materials_examined">
<paragraph id="8BAA3681FF83CD62FF4DF896FDEA14C6" blockId="0.[151,1437,1900,2033]" pageId="0" pageNumber="394">
<emphasis id="B961EA93FF83CD62FF4DF896FE9714EA" bold="true" box="[151,381,1900,1925]" pageId="0" pageNumber="394">Material examined</emphasis>
.
<typeStatus id="54AE8823FF83CD62FE55F896FE1014EA" box="[399,506,1900,1925]" pageId="0" pageNumber="394" type="holotype">Holotype</typeStatus>
, ovigerous female, MBM136607, cl
<quantity id="4CED9B64FF83CD62FC76F897FC1414EA" box="[940,1022,1901,1925]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="0" pageNumber="394" unit="mm" value="1.8">1.8mm</quantity>
, intertidal, Sanya, Hainan Island,
<date id="FFAB1041FF83CD62FA54F896FF0914C6" pageId="0" pageNumber="394">5 March</date>
, 1997, coll. Xinzheng Li.
</paragraph>
<paragraph id="8BAA3681FF83CD62FF1DF84FFA77149E" blockId="0.[151,1437,1900,2033]" pageId="0" pageNumber="394">
<typeStatus id="54AE8823FF83CD62FF1DF84FFEDD14A2" box="[199,311,1973,1997]" pageId="0" pageNumber="394" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D13FD08FF83CD62FE98F84FFE7A14A2" box="[322,400,1972,1997]" pageId="0" pageNumber="394" type="male">1 male</specimenCount>
, MBM136405, cl
<quantity id="4CED9B64FF83CD62FD85F84FFD5A14A2" box="[607,688,1973,1997]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="0" pageNumber="394" unit="mm" value="1.9">1.9mm</quantity>
, intertidal, Qionghai, Hainan Island,
<date id="FFAB1041FF83CD62FB88F84FFB4F14A2" box="[1106,1189,1972,1997]" pageId="0" pageNumber="394">4 April</date>
, 1992, coll. Xinzheng Li; 5 ovigerous females, MBM136387, cl
<quantity id="4CED9B64FF83CD62FDA0F823FD16149E" box="[634,764,2009,2033]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="0" pageNumber="394" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.72.1mm</quantity>
, intertidal, Lingao, Hainan Island,
<date id="FFAB1041FF83CD62FB51F823FAFD149E" box="[1163,1303,2008,2033]" pageId="0" pageNumber="394">2 December</date>
, 1990, coll.
</paragraph>
<paragraph id="8BAA3681FF82CD63FF4DFF6DFE431397" blockId="1.[151,1437,151,1149]" pageId="1" pageNumber="395">
Huilian Chen; 4 ovigerous females, MBM136589, cl
<quantity id="4CED9B64FF82CD63FD2AFF62FC9913DF" box="[752,883,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="1" pageNumber="395" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.81.9mm</quantity>
, intertidal, Sanya, Hainan Island,
<date id="FFAB1041FF82CD63FB2AFF62FAB813DF" box="[1264,1362,151,176]" pageId="1" pageNumber="395">1 March</date>
, 1997, coll. Xinzheng Li; 5 ovigerous females, MBM136394, cl 1.82.0mm,
<quantity id="4CED9B64FF82CD63FC62FF46FC1013BB" box="[952,1018,188,212]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="1" pageNumber="395" unit="m" value="1.0" valueMax="2.0" valueMin="0.0">02m</quantity>
, Lingshui, Hainan Island,
<date id="FFAB1041FF82CD63FAF2FF46FA7213BA" box="[1320,1432,188,213]" pageId="1" pageNumber="395" value="1926-03">26 March</date>
, 1992, coll. Xinzheng Li.
</paragraph>
</subSubSection>
<subSubSection id="C30F650AFF82CD63FF1DFEFEFD47129A" pageId="1" pageNumber="395" type="diagnosis">
<paragraph id="8BAA3681FF82CD63FF1DFEFEFD47129A" blockId="1.[151,1437,151,1149]" pageId="1" pageNumber="395">
<emphasis id="B961EA93FF82CD63FF1DFEFEFEAB1272" bold="true" box="[199,321,260,285]" pageId="1" pageNumber="395">Diagnosis.</emphasis>
Rostrum with bifid appearance about half length of carapace; carapace with developed antennal tooth and minute pterygostomian tooth; third segment of antennular peduncle with subtriangular dorsal scale. Abdomen rounded dorsally; pleurae of fourth and fifth abdominal somites each with small posteroventral tooth; dorsal surface of telson with 35 pairs of dorsolateral spines. Mandible only with incisor process and molar process. Third maxilliped with exopod and epipod. First pereopod with epipod; carpus on the second pereopod with six-segmented. Third and fourth pereopods prehensile, subchelate in male; second pleopod with appendix masculine about 0.9 length of appendix interna.
</paragraph>
</subSubSection>
<subSubSection id="C30F650AFF82CD67FF1DFE05FDFD12C2" lastPageId="5" lastPageNumber="399" pageId="1" pageNumber="395" type="description">
<paragraph id="8BAA3681FF82CD63FF1DFE05FB9E1177" blockId="1.[151,1437,151,1149]" box="[199,1140,511,536]" pageId="1" pageNumber="395">
<emphasis id="B961EA93FF82CD63FF1DFE05FEBB1177" bold="true" box="[199,337,511,536]" pageId="1" pageNumber="395">Description</emphasis>
.
<emphasis id="B961EA93FF82CD63FE85FE05FDC81177" bold="true" box="[351,546,511,536]" pageId="1" pageNumber="395">
<typeStatus id="54AE8823FF82CD63FE85FE05FE261177" box="[351,460,511,536]" pageId="1" pageNumber="395" type="holotype">Holotype</typeStatus>
female
</emphasis>
. Small hippolytid shrimp without supraorbital spine.
</paragraph>
<paragraph id="8BAA3681FF82CD63FF1DFDDFFD6E11C7" blockId="1.[151,1437,151,1149]" pageId="1" pageNumber="395">
Rostrum (
<figureCitation id="132E2A04FF82CD63FEE1FDDEFE971153" box="[315,381,548,573]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="1" pageNumber="395">Fig. 1</figureCitation>
A, B) about half length of carapace, nearly straight, slightly descending, reaching but not exceeding basal segment of antennular peduncle; dorsal margin with three teeth, posterior one on carapace posterior to margin of orbit; ventral margin armed with only one subterminal tooth, giving rostrum bifid appearance; lateral carina feebly developed.
</paragraph>
<paragraph id="8BAA3681FF82CD63FF1DFD4EFE98107A" blockId="1.[151,1437,151,1149]" pageId="1" pageNumber="395">
Carapace (
<figureCitation id="132E2A04FF82CD63FE9DFD4EFE6411A3" box="[327,398,692,717]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="1" pageNumber="395">Fig. 1</figureCitation>
A) with inferior orbital angle slightly produced, subacute; antennal tooth moderately developed; pterygostomian tooth minute; anterolateral margin between antennal tooth and pterygostomian tooth obscurely angulate.
</paragraph>
<paragraph id="8BAA3681FF82CD63FF1DFCE5FD4410CA" blockId="1.[151,1437,151,1149]" pageId="1" pageNumber="395">
Abdomen (
<figureCitation id="132E2A04FF82CD63FE97FCE5FE791057" box="[333,403,799,824]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="1" pageNumber="395">Fig. 1</figureCitation>
A) rounded dorsally. Pleurae of fourth and fifth abdominal somites each with small posteroventral tooth. Sixth somite 1.9 times as long as fifth mid-dorsal length. Telson (
<figureCitation id="132E2A04FF82CD63FB83FCBEFB761033" box="[1113,1180,836,861]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="1" pageNumber="395">Figs 1</figureCitation>
A, 2B) about 1.2 times longer than sixth abdominal somite; dorsal surface with three pairs of dorsolateral spines and three pairs of terminal spines, lateral pair similar to dorsolateral series.
</paragraph>
<paragraph id="8BAA3681FF82CD63FF1DFC4AFB2810A7" blockId="1.[151,1437,151,1149]" box="[199,1218,943,968]" pageId="1" pageNumber="395">
Eye (
<figureCitation id="132E2A04FF82CD63FED8FC55FEAA10A7" box="[258,320,943,968]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="1" pageNumber="395">Fig. 1</figureCitation>
A) subpyriform, pigmented; cornea with distinct ocellar spot, shorter than stalk.
</paragraph>
<paragraph id="8BAA3681FF82CD63FF1DFC2EFCB41712" blockId="1.[151,1437,151,1149]" pageId="1" pageNumber="395">
Antennular peduncle (
<figureCitation id="132E2A04FF82CD63FE1BFC2EFDEE1083" box="[449,516,980,1005]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="1" pageNumber="395">Figs 1</figureCitation>
A, 2A) reaching distal 0.4 of antennal scale. Basal segment with strong subdistal, ventromesial tooth; sharp stylocerite reaching slightly beyond distal margin of second segment, armed with little indistinct tooth near proximal end of lateral margin; second segment with curved lateral spine reaching beyond mid-length of distal segment; third segment with subtriangular dorsal scale. Outer flagellum stout and heavily setose. Inner flagellum slender, elongate, with 2328 segments.
</paragraph>
<caption id="DF6A6609FF82CD63FF4DF859FEA714B8" httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="1" pageNumber="395" targetBox="[161,1428,1212,1924]" targetPageId="1">
<paragraph id="8BAA3681FF82CD63FF4DF859FEA714B8" blockId="1.[151,1436,1955,2007]" pageId="1" pageNumber="395">
<emphasis id="B961EA93FF82CD63FF4DF859FEF814D7" bold="true" box="[151,274,1955,1977]" pageId="1" pageNumber="395">FIGURE 1.</emphasis>
<taxonomicName id="4C154D02FF82CD63FEC2F859FE2314D6" box="[280,457,1955,1977]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="395" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="B961EA93FF82CD63FEC2F859FE2314D6" box="[280,457,1955,1977]" italics="true" pageId="1" pageNumber="395">Thor hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257E8FF82CD63FE15F859FDF514D7" box="[463,543,1955,1976]" pageId="1" pageNumber="395" rank="species">
<emphasis id="B961EA93FF82CD63FE15F859FDF514D7" bold="true" box="[463,543,1955,1976]" pageId="1" pageNumber="395">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF6A6609FF81CD60FF4DF8A2FD7E1486" httpUri="https://zenodo.org/record/227973/files/figure.png" pageId="2" pageNumber="396" targetBox="[210,1380,213,1832]" targetPageId="2">
<paragraph id="8BAA3681FF81CD60FF4DF8A2FD7E1486" blockId="2.[151,1436,1880,2025]" pageId="2" pageNumber="396">
<emphasis id="B961EA93FF81CD60FF4DF8A2FEFC1402" bold="true" box="[151,278,1880,1902]" pageId="2" pageNumber="396">FIGURE 2.</emphasis>
<taxonomicName id="4C154D02FF81CD60FEC5F8A2FE391401" box="[287,467,1880,1902]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="B961EA93FF81CD60FEC5F8A2FE391401" box="[287,467,1880,1902]" italics="true" pageId="2" pageNumber="396">Thor hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257E8FF81CD60FE07F8A2FDDA1402" box="[477,560,1880,1901]" pageId="2" pageNumber="396" rank="species">
<emphasis id="B961EA93FF81CD60FE07F8A2FDDA1402" bold="true" box="[477,560,1880,1901]" pageId="2" pageNumber="396">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF6A6609FF80CD61FF4DFBD8FBEC1738" httpUri="https://zenodo.org/record/227974/files/figure.png" pageId="3" pageNumber="397" targetBox="[160,1431,201,1036]" targetPageId="3">
<paragraph id="8BAA3681FF80CD61FF4DFBD8FBEC1738" blockId="3.[151,1436,1058,1111]" pageId="3" pageNumber="397">
<emphasis id="B961EA93FF80CD61FF4DFBD8FEFC1758" bold="true" box="[151,278,1058,1080]" pageId="3" pageNumber="397">FIGURE 3.</emphasis>
<taxonomicName id="4C154D02FF80CD61FEFAFBD8FE3E1757" box="[288,468,1058,1080]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="397" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="B961EA93FF80CD61FEFAFBD8FE3E1757" box="[288,468,1058,1080]" italics="true" pageId="3" pageNumber="397">Thor hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257E8FF80CD61FE04FBD9FDD81757" box="[478,562,1059,1080]" pageId="3" pageNumber="397" rank="species">
<emphasis id="B961EA93FF80CD61FE04FBD9FDD81757" bold="true" box="[478,562,1059,1080]" pageId="3" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, male (cl 1.9 mm). A, right third pereopod, lateral view; B, right fourth pereopod, lateral view; C, endopod of second pleopod, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF6A6609FF80CD61FF4DF863FAB71483" httpUri="https://zenodo.org/record/227975/files/figure.png" pageId="3" pageNumber="397" targetBox="[411,1175,1161,1923]" targetPageId="3">
<paragraph id="8BAA3681FF80CD61FF4DF863FAB71483" blockId="3.[151,1436,1945,2028]" pageId="3" pageNumber="397">
<emphasis id="B961EA93FF80CD61FF4DF863FEF914C1" bold="true" box="[151,275,1945,1967]" pageId="3" pageNumber="397">FIGURE 4.</emphasis>
<taxonomicName id="4C154D02FF80CD61FEC0F863FE2614C0" box="[282,460,1945,1967]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="397" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="B961EA93FF80CD61FEC0F863FE2614C0" box="[282,460,1945,1967]" italics="true" pageId="3" pageNumber="397">Thor hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257E8FF80CD61FE08F863FDC914C1" box="[466,547,1945,1966]" pageId="3" pageNumber="397" rank="species">
<emphasis id="B961EA93FF80CD61FE08F863FDC914C1" bold="true" box="[466,547,1945,1966]" pageId="3" pageNumber="397">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, male (cl 1.9 mm). A, propodus and dactylus of third pereopod, lateral view; B, distal third of propodus and dactylus of third pereopod, lateral view; C, propodus and dactylus of fourth pereopod, lateral view; B, distal third of propodus and dactylus of fourth pereopod, lateral view. Scale bars: A, C (500µm); B, D (200µm).
</paragraph>
</caption>
<caption id="DF6A6609FF87CD66FF4DFCA3FBC110E2" httpUri="https://zenodo.org/record/227976/files/figure.png" pageId="4" pageNumber="398" targetBox="[151,1436,193,835]" targetPageId="4">
<paragraph id="8BAA3681FF87CD66FF4DFCA3FBC110E2" blockId="4.[151,1435,857,909]" pageId="4" pageNumber="398">
<emphasis id="B961EA93FF87CD66FF4DFCA3FEF91001" bold="true" box="[151,275,857,879]" pageId="4" pageNumber="398">FIGURE 5.</emphasis>
<taxonomicName id="4C154D02FF87CD66FEC1FCA3FE261000" box="[283,460,857,879]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="398" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="B961EA93FF87CD66FEC1FCA3FE261000" box="[283,460,857,879]" italics="true" pageId="4" pageNumber="398">Thor hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257E8FF87CD66FE09FCA3FDCE1001" box="[467,548,857,878]" pageId="4" pageNumber="398" rank="species">
<emphasis id="B961EA93FF87CD66FE09FCA3FDCE1001" bold="true" box="[467,548,857,878]" pageId="4" pageNumber="398">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, ovigerous female (cl 2.0 mm). A, anterior part of body, lateral view; B, key zone indicates the epipod of first pereopod, lateral view. Scale bars: A (1mm); B (400µm).
</paragraph>
</caption>
<paragraph id="8BAA3681FF87CD66FF1DFC40FA851774" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
Antennal peduncle (
<figureCitation id="132E2A04FF87CD66FE72FC40FE0D10BC" box="[424,487,954,979]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="4" pageNumber="398">Fig. 1</figureCitation>
A) with basicerite bearing strong distal, ventrolateral tooth, dorsodistal angle bluntly produced. Carpocerite overreaching second segment of antennular peduncle. Antennal scale shorter than length of carapace, 2.4 times as long as wide; lateral margin faintly concave, with distolateral tooth exceeded by lamina.
</paragraph>
<paragraph id="8BAA3681FF87CD66FF1DFBDCFD40170D" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
Mouthparts (
<figureCitation id="132E2A04FF87CD66FE83FBDCFE761750" box="[345,412,1062,1087]" captionStart="FIGURE 2" captionStartId="2.[151,250,1880,1902]" captionTargetBox="[210,1380,213,1832]" captionTargetId="figure@2.[195,1395,193,1853]" captionTargetPageId="2" captionText="FIGURE 2. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227973/files/figure.png" pageId="4" pageNumber="398">Fig. 2</figureCitation>
CG) typical of genus. Mandible without palp; incisor process slender, with series of four subequal, tiny, distal teeth; molar process stout.
</paragraph>
<paragraph id="8BAA3681FF87CD66FF1DFB94FEA9179D" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
Third maxilliped (
<figureCitation id="132E2A04FF87CD66FE40FB94FE0A17E8" box="[410,480,1134,1159]" captionStart="FIGURE 2" captionStartId="2.[151,250,1880,1902]" captionTargetBox="[210,1380,213,1832]" captionTargetId="figure@2.[195,1395,193,1853]" captionTargetPageId="2" captionText="FIGURE 2. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227973/files/figure.png" pageId="4" pageNumber="398">Fig. 2</figureCitation>
H) long and slender, somewhat beyond antennal scale; ultimate segment 4.3 times longer than penultimate segment, with corneous spines distally; antepenultimate segment subequal in length to ultimate segment, distoventral margin with small spine. Exopod exceeding middle of antepenultimate segment; epipod present.
</paragraph>
<paragraph id="8BAA3681FF87CD66FF1DFB04FF1916C8" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
First pereopod (
<figureCitation id="132E2A04FF87CD66FEA2FB04FE511678" box="[376,443,1278,1303]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="4" pageNumber="398">Figs 1</figureCitation>
A,
<date id="FFAB1041FF87CD66FE3BFB05FE101678" box="[481,506,1279,1303]" pageId="4" pageNumber="398">2I</date>
) with strap-like, terminally hooked epipod; moderately stout, falling short of distal margin of antennal scale. Chela about 1.2 times longer than carpus, 3.1 times longer than wide; dactylus with bifid tip, slightly longer than half length of palm; palm subcylindrical, 2.1 times as long as wide, fixed finger terminating in single corneous claw. Carpus 3.1 times longer than wide. Merus subequal in length to carpus, 2.3 times as long as wide.
</paragraph>
<paragraph id="8BAA3681FF87CD66FF1DFA48FC6D1680" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
Second pereopod (
<figureCitation id="132E2A04FF87CD66FE78FA48FE0316A4" box="[418,489,1458,1483]" captionStart="FIGURE 2" captionStartId="2.[151,250,1880,1902]" captionTargetBox="[210,1380,213,1832]" captionTargetId="figure@2.[195,1395,193,1853]" captionTargetPageId="2" captionText="FIGURE 2. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227973/files/figure.png" pageId="4" pageNumber="398">Fig. 2</figureCitation>
J) slender and chelate; carpus with six sub-segments, third sub-segment longest. Merus shorter than carpus, 6.3 times longer than width. No epipod.
</paragraph>
<paragraph id="8BAA3681FF87CD66FF1DFA00FB5015CC" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
Third pereopod (
<figureCitation id="132E2A04FF87CD66FE53FA00FE25157C" box="[393,463,1530,1555]" captionStart="FIGURE 2" captionStartId="2.[151,250,1880,1902]" captionTargetBox="[210,1380,213,1832]" captionTargetId="figure@2.[195,1395,193,1853]" captionTargetPageId="2" captionText="FIGURE 2. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227973/files/figure.png" pageId="4" pageNumber="398">Fig. 2</figureCitation>
K) overreaching antennal scale by the length of dactylus; biunguiculate dactylus 0.2 times as long as propodus, about 2.6 times longer than high, armed with three accessory spines along entire length of flexor margin; propodus 10 times as long as wide; flexor margin with about 910 spinules, spinules increasing in size distally; carpus less than half length of propodus, 2.94.2 times longer than wide. Merus about equal to propodus, 5.9 times longer than deep, with prominent lateral spines distally. Ischium unarmed.
</paragraph>
<paragraph id="8BAA3681FF87CD66FF1DF954FDB51460" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
Fourth pereopod (
<figureCitation id="132E2A04FF87CD66FE41F954FE0915A8" box="[411,483,1710,1735]" captionStart="FIGURE 2" captionStartId="2.[151,250,1880,1902]" captionTargetBox="[210,1380,213,1832]" captionTargetId="figure@2.[195,1395,193,1853]" captionTargetPageId="2" captionText="FIGURE 2. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227973/files/figure.png" pageId="4" pageNumber="398">Fig. 2</figureCitation>
L) similar to third pereopod, overreaching antennal scale, biunguiculate dactylus armed with four accessory spines along entire length of flexor margin; merus with one distolateral spine. Fifth pereopod (
<figureCitation id="132E2A04FF87CD66FED4F90CFEBB1460" box="[270,337,1782,1807]" captionStart="FIGURE 2" captionStartId="2.[151,250,1880,1902]" captionTargetBox="[210,1380,213,1832]" captionTargetId="figure@2.[195,1395,193,1853]" captionTargetPageId="2" captionText="FIGURE 2. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227973/files/figure.png" pageId="4" pageNumber="398">Fig. 2</figureCitation>
M) with no meral spine.
</paragraph>
<paragraph id="8BAA3681FF87CD66FF1DF8E0FC691439" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
Pleopods (
<figureCitation id="132E2A04FF87CD66FEE5F8E0FE6A145C" box="[319,384,1818,1843]" captionStart="FIGURE 1" captionStartId="1.[151,250,1955,1977]" captionTargetBox="[161,1428,1212,1924]" captionTargetId="figure@1.[151,1436,1201,1940]" captionTargetPageId="1" captionText="FIGURE 1. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, body, lateral view; B, rostrum, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227972/files/figure.png" pageId="4" pageNumber="398">Fig. 1</figureCitation>
A) without special features. Uropods (
<figureCitation id="132E2A04FF87CD66FCE6F8E0FC68145C" box="[828,898,1818,1843]" captionStart="FIGURE 2" captionStartId="2.[151,250,1880,1902]" captionTargetBox="[210,1380,213,1832]" captionTargetId="figure@2.[195,1395,193,1853]" captionTargetPageId="2" captionText="FIGURE 2. Thor hainanensis sp. nov., holotype, ovigerous female (cl 1.8 mm). A, antennule, dorsal view; B, telson and uropod, dorsal view; C, left mandible, external view; D, left maxillule, external view; E, left maxilla, external view; F, left first maxilliped, external view; G, left second maxilliped, external view; H, left third maxilliped, lateral view; I, left first pereopod, lateral view; J, left second pereopod, lateral view; K, left third pereopod, lateral view; L, left fourth pereopod, lateral view; M, left fifth pereopod, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227973/files/figure.png" pageId="4" pageNumber="398">Fig. 2</figureCitation>
B) longer than telson, fringed with setae; outer branch of uropod with fixed tooth and movable spine along suture.
</paragraph>
<paragraph id="8BAA3681FF87CD66FF1DF898FE1414AC" blockId="4.[151,1437,954,1987]" pageId="4" pageNumber="398">
<emphasis id="B961EA93FF87CD66FF1DF898FE7F1414" bold="true" box="[199,405,1890,1915]" pageId="4" pageNumber="398">
<typeStatus id="54AE8823FF87CD66FF1DF898FEDE1414" box="[199,308,1890,1915]" pageId="4" pageNumber="398" type="paratype">Paratype</typeStatus>
females
</emphasis>
. Very similar to
<typeStatus id="54AE8823FF87CD66FD89F898FD5F1414" box="[595,693,1890,1915]" pageId="4" pageNumber="398" type="holotype">holotype</typeStatus>
female except for following: merus of third pereopod armed with two distolateral spines; merus of fifth pereopod with one distolateral spine; dorsal surface of telson with four or five pairs of dorsolateral spines.
</paragraph>
<paragraph id="8BAA3681FF86CD67FF1DFF6DFE7712E8" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">
<emphasis id="B961EA93FF86CD67FF1DFF6DFE9C13DF" bold="true" box="[199,374,151,176]" pageId="5" pageNumber="399">
<typeStatus id="54AE8823FF86CD67FF1DFF6DFEDE13DF" box="[199,308,151,176]" pageId="5" pageNumber="399" type="paratype">Paratype</typeStatus>
male
</emphasis>
. Third pereopod (
<figureCitation id="132E2A04FF86CD67FD9EFF6DFD6C13DF" box="[580,646,151,176]" captionStart="FIGURE 3" captionStartId="3.[151,250,1058,1080]" captionTargetBox="[160,1431,201,1036]" captionTargetId="figure@3.[151,1436,193,1049]" captionTargetPageId="3" captionText="FIGURE 3. Thor hainanensis sp. nov., paratype, male (cl 1.9 mm). A, right third pereopod, lateral view; B, right fourth pereopod, lateral view; C, endopod of second pleopod, ventral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227974/files/figure.png" pageId="5" pageNumber="399">Fig. 3</figureCitation>
A) prehensile, subchelate. Dactylus (
<figureCitation id="132E2A04FF86CD67FBF3FF6DFB8713DF" box="[1065,1133,151,176]" captionStart="FIGURE 4" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[411,1175,1161,1923]" captionTargetId="figure@3.[411,1175,1161,1924]" captionTargetPageId="3" captionText="FIGURE 4. Thor hainanensis sp. nov., paratype, male (cl 1.9 mm). A, propodus and dactylus of third pereopod, lateral view; B, distal third of propodus and dactylus of third pereopod, lateral view; C, propodus and dactylus of fourth pereopod, lateral view; B, distal third of propodus and dactylus of fourth pereopod, lateral view. Scale bars: A, C (500 µm); B, D (200 µm)." httpUri="https://zenodo.org/record/227975/files/figure.png" pageId="5" pageNumber="399">Fig. 4</figureCitation>
B) with bifid tip and many closely appressed spinules on flexor margin; propodus (
<figureCitation id="132E2A04FF86CD67FCDEFF46FCAD13BB" box="[772,839,188,213]" captionStart="FIGURE 4" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[411,1175,1161,1923]" captionTargetId="figure@3.[411,1175,1161,1924]" captionTargetPageId="3" captionText="FIGURE 4. Thor hainanensis sp. nov., paratype, male (cl 1.9 mm). A, propodus and dactylus of third pereopod, lateral view; B, distal third of propodus and dactylus of third pereopod, lateral view; C, propodus and dactylus of fourth pereopod, lateral view; B, distal third of propodus and dactylus of fourth pereopod, lateral view. Scale bars: A, C (500 µm); B, D (200 µm)." httpUri="https://zenodo.org/record/227975/files/figure.png" pageId="5" pageNumber="399">Fig. 4</figureCitation>
A, B) more than twice as long as dactylus, distal third of flexor margin with densely spinules; carpus less than half as long as propodus; merus slightly shorter than propodus, armed with strong spine near distal end of lateral surface. Fourth pereopod (
<figureCitation id="132E2A04FF86CD67FB5FFEFEFB231273" box="[1157,1225,260,285]" captionStart="FIGURE 3" captionStartId="3.[151,250,1058,1080]" captionTargetBox="[160,1431,201,1036]" captionTargetId="figure@3.[151,1436,193,1049]" captionTargetPageId="3" captionText="FIGURE 3. Thor hainanensis sp. nov., paratype, male (cl 1.9 mm). A, right third pereopod, lateral view; B, right fourth pereopod, lateral view; C, endopod of second pleopod, ventral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227974/files/figure.png" pageId="5" pageNumber="399">Fig. 3</figureCitation>
B) similar to third pereopod, prehensile, subchelate; merus also armed with one distolateral spine. Second pleopod (
<figureCitation id="132E2A04FF86CD67FB24FEDDFAA9122F" box="[1278,1347,295,320]" captionStart="FIGURE 3" captionStartId="3.[151,250,1058,1080]" captionTargetBox="[160,1431,201,1036]" captionTargetId="figure@3.[151,1436,193,1049]" captionTargetPageId="3" captionText="FIGURE 3. Thor hainanensis sp. nov., paratype, male (cl 1.9 mm). A, right third pereopod, lateral view; B, right fourth pereopod, lateral view; C, endopod of second pleopod, ventral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/227974/files/figure.png" pageId="5" pageNumber="399">Fig. 3</figureCitation>
C) with appendix masculine moderately stout, almost 6 times as long as wide, about 0.9 length of appendix interna, bearing about 5 long stiff setae.
</paragraph>
<paragraph id="8BAA3681FF86CD67FF1DFE6EFDFD12C2" blockId="5.[151,1437,151,1293]" box="[199,535,404,429]" pageId="5" pageNumber="399">
<emphasis id="B961EA93FF86CD67FF1DFE6EFE7812C2" bold="true" box="[199,402,404,429]" pageId="5" pageNumber="399">Coloration in life</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30F650AFF86CD67FF1DFE4DFDC1129A" pageId="5" pageNumber="399" type="distribution">
<paragraph id="8BAA3681FF86CD67FF1DFE4DFDC1129A" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">
<emphasis id="B961EA93FF86CD67FF1DFE4DFEB312BF" bold="true" box="[199,345,439,464]" pageId="5" pageNumber="399">Distribution</emphasis>
. Only known from the
<typeStatus id="54AE8823FF86CD67FDBFFE42FD7F12BF" box="[613,661,440,464]" pageId="5" pageNumber="399">type</typeStatus>
locality, Hainan Island. Specimens collected from shallow intertidal coral reef pools at depths of
<quantity id="4CED9B64FF86CD67FE0CFE26FDCC129A" box="[470,550,476,501]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="5" pageNumber="399" unit="m" value="8.5" valueMax="15.0" valueMin="2.0">
2
<specimenCount id="9D13FD08FF86CD67FE2EFE26FDCC129A" box="[500,550,476,501]" pageId="5" pageNumber="399" type="generic">15m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F650AFF86CD67FF1DFE05FB051177" box="[199,1263,511,536]" pageId="5" pageNumber="399" type="etymology">
<paragraph id="8BAA3681FF86CD67FF1DFE05FB051177" blockId="5.[151,1437,151,1293]" box="[199,1263,511,536]" pageId="5" pageNumber="399">
<emphasis id="B961EA93FF86CD67FF1DFE05FEAC1177" bold="true" box="[199,326,511,536]" pageId="5" pageNumber="399">Etymology</emphasis>
. The name is derived from the
<typeStatus id="54AE8823FF86CD67FD7CFDFAFD3C1177" box="[678,726,512,536]" pageId="5" pageNumber="399">type</typeStatus>
locality, Hainan Island, in the South
<collectingCountry id="F3027611FF86CD67FBACFE05FB531177" box="[1142,1209,511,536]" name="China" pageId="5" pageNumber="399">China</collectingCountry>
Sea.
</paragraph>
</subSubSection>
<subSubSection id="C30F650AFF86CD67FF1DFDDEFB0B1713" pageId="5" pageNumber="399" type="discussion">
<paragraph id="8BAA3681FF86CD67FF1DFDDEFE3111EA" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">
<emphasis id="B961EA93FF86CD67FF1DFDDEFEDE1152" bold="true" box="[199,308,548,573]" pageId="5" pageNumber="399">Remarks</emphasis>
. The most important morphological character of the new species is the presence of an epipod (
<figureCitation id="132E2A04FF86CD67FAB5FDDEFF4F110F" captionStart="FIGURE 5" captionStartId="4.[151,250,857,879]" captionTargetBox="[151,1436,193,835]" captionTargetId="figure@4.[151,1436,193,836]" captionTargetPageId="4" captionText="FIGURE 5. Thor hainanensis sp. nov., paratype, ovigerous female (cl 2.0 mm). A, anterior part of body, lateral view; B, key zone indicates the epipod of first pereopod, lateral view. Scale bars: A (1 mm); B (400 µm)." httpUri="https://zenodo.org/record/227976/files/figure.png" pageId="5" pageNumber="399">Fig. 5</figureCitation>
A, B) on the first pereopod. The fourth pereopod (
<figureCitation id="132E2A04FF86CD67FD0FFDBDFCF2110F" box="[725,792,583,608]" captionStart="FIGURE 4" captionStartId="3.[151,250,1945,1967]" captionTargetBox="[411,1175,1161,1923]" captionTargetId="figure@3.[411,1175,1161,1924]" captionTargetPageId="3" captionText="FIGURE 4. Thor hainanensis sp. nov., paratype, male (cl 1.9 mm). A, propodus and dactylus of third pereopod, lateral view; B, distal third of propodus and dactylus of third pereopod, lateral view; C, propodus and dactylus of fourth pereopod, lateral view; B, distal third of propodus and dactylus of fourth pereopod, lateral view. Scale bars: A, C (500 µm); B, D (200 µm)." httpUri="https://zenodo.org/record/227975/files/figure.png" pageId="5" pageNumber="399">Fig. 4</figureCitation>
C, D) of male specimen is prehensile and subchelate, also rare and unique in the genus.
</paragraph>
<paragraph id="8BAA3681FF86CD67FF1DFD75FECC11A2" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">Besides the epipod on the first pereopod, the new species can be easily distinguished from the known species in the genus.</paragraph>
<paragraph id="8BAA3681FF86CD67FF1DFD2DFE6F1034" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">
The new species is very similar to
<taxonomicName id="4C154D02FF86CD67FD8BFD2DFBC2119F" authority="Wicksten &amp; Vargas, 2001" authorityName="Wicksten &amp; Vargas" authorityYear="2001" box="[593,1064,727,752]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="cocoensis">
<emphasis id="B961EA93FF86CD67FD8BFD2DFD16119F" box="[593,764,727,752]" italics="true" pageId="5" pageNumber="399">Thor cocoensis</emphasis>
<bibRefCitation id="EF844B70FF86CD67FCDEFD2DFBC2119F" author="Wicksten" box="[772,1064,727,752]" pageId="5" pageNumber="399" refString="Wicksten, M. K. &amp; Vargas, R. (2001) A new species of Thor Kingsley, 1878 (Crustacea: Decapoda: Caridea: Hippolytidae) from the tropical eastern Pacific. Proceedings of the Biological Society of Washington, 114, 139 - 144." type="journal article" year="2001">Wicksten &amp; Vargas, 2001</bibRefCitation>
</taxonomicName>
. It can be distinguished from the latter by (1) the dorsal margin of rostrum armed with three teeth in
<taxonomicName id="4C154D02FF86CD67FC4CFD07FBD3107A" box="[918,1081,764,789]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B961EA93FF86CD67FC4CFD07FC49107B" box="[918,931,765,788]" italics="true" pageId="5" pageNumber="399">T</emphasis>
.
<emphasis id="B961EA93FF86CD67FC69FD06FBD3107A" box="[947,1081,764,789]" italics="true" pageId="5" pageNumber="399">hainanensis</emphasis>
</taxonomicName>
, however,
<taxonomicName id="4C154D02FF86CD67FB6CFD07FAAA107B" box="[1206,1344,765,788]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="cocoensis">
<emphasis id="B961EA93FF86CD67FB6CFD07FAAA107B" box="[1206,1344,765,788]" italics="true" pageId="5" pageNumber="399">T. cocoensis</emphasis>
</taxonomicName>
bearing only one or two dorsal teeth; (2) the tip of rostrum shows distinctly bifid in
<taxonomicName id="4C154D02FF86CD67FC33FCDBFB601057" box="[1001,1162,799,824]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B961EA93FF86CD67FC33FCDBFC1C1057" box="[1001,1014,801,824]" italics="true" pageId="5" pageNumber="399">T</emphasis>
.
<emphasis id="B961EA93FF86CD67FBDEFCE5FB601057" box="[1028,1162,799,824]" italics="true" pageId="5" pageNumber="399">hainanensis</emphasis>
</taxonomicName>
,
<emphasis id="B961EA93FF86CD67FB43FCDBFB5D1057" box="[1177,1207,801,824]" italics="true" pageId="5" pageNumber="399">vs.</emphasis>
the tip of rostrum is acute in
<taxonomicName id="4C154D02FF86CD67FF2FFCBFFE941033" box="[245,382,837,860]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="cocoensis">
<emphasis id="B961EA93FF86CD67FF2FFCBFFE941033" box="[245,382,837,860]" italics="true" pageId="5" pageNumber="399">T. cocoensis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAA3681FF86CD67FF1DFC9DFD0B10A7" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">
The new species distinguishes from
<taxonomicName id="4C154D02FF86CD67FDADFC9DFC3810EF" authority="Wicksten, 1996" authorityName="Wicksten" authorityYear="1996" box="[631,978,871,896]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="cordelli">
<emphasis id="B961EA93FF86CD67FDADFC9DFCFA10EF" box="[631,784,871,896]" italics="true" pageId="5" pageNumber="399">Thor cordelli</emphasis>
<bibRefCitation id="EF844B70FF86CD67FCC6FC9DFC3810EF" author="Wicksten" box="[796,978,871,896]" pageId="5" pageNumber="399" refString="Wicksten, M. K. (1996) A new species of hippolytid shrimp from Rocas Alijos. In: Schmieder, R. W. (Ed.), Rocas Alijos. Kluwer Academic Publsihers, Dordrecht, pp. 295 - 298. http: // dx. doi. org / 10.1007 / 978 - 94 - 017 - 2917 - 8 _ 24" type="book chapter" year="1996">Wicksten, 1996</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C154D02FF86CD67FC39FC93FB1210EF" authority="Chace, 1972" authorityName="Chace" authorityYear="1972" box="[995,1272,871,896]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="dobkini">
<emphasis id="B961EA93FF86CD67FC39FC93FBB210EF" box="[995,1112,871,896]" italics="true" pageId="5" pageNumber="399">T. dobkini</emphasis>
<bibRefCitation id="EF844B70FF86CD67FBBEFC9DFB1210EF" author="Chace" box="[1124,1272,871,896]" pageId="5" pageNumber="399" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. http: // dx. doi. org / 10.5479 / si. 00810282.98" type="journal article" year="1972">Chace, 1972</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C154D02FF86CD67FAD2FC93FEAC10CB" authority="Kingsley, 1878" authorityName="Kingsley" authorityYear="1878" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="floridanus">
<emphasis id="B961EA93FF86CD67FAD2FC93FA7610EF" box="[1288,1436,871,896]" italics="true" pageId="5" pageNumber="399">T. floridanus</emphasis>
<bibRefCitation id="EF844B70FF86CD67FF4DFC76FEAC10CB" author="Kingsley" box="[151,326,908,933]" pageId="5" pageNumber="399" refString="Kingsley, J. S. (1878) Notes on the North American Caridea in the Museum of the Peabody Academy of Science at Salem, Mass. Proceedings of the Academy of Natural Sciences of Philadelphia, 1878, 89 - 98." type="proceedings paper" year="1878">Kingsley, 1878</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C154D02FF86CD67FE8EFC77FD9D10CB" authority="Chace, 1972" authorityName="Chace" authorityYear="1972" box="[340,631,908,933]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="manningi">
<emphasis id="B961EA93FF86CD67FE8EFC77FE3610CB" box="[340,476,909,932]" italics="true" pageId="5" pageNumber="399">T. manningi</emphasis>
<bibRefCitation id="EF844B70FF86CD67FE3FFC76FD9D10CB" author="Chace" box="[485,631,908,933]" pageId="5" pageNumber="399" refString="Chace, F. A. Jr. (1972) The shrimps of the Smithsonian-Bredin Caribbean expeditions with a summary of the West Indian shallow-water species (Crustacea: Decapoda: Natantia). Smithsonian Contributions to Zoology, 98, 1 - 179. http: // dx. doi. org / 10.5479 / si. 00810282.98" type="journal article" year="1972">Chace, 1972</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C154D02FF86CD67FD5FFC77FC7E10CB" authority="Bruce, 1983" authorityName="Bruce" authorityYear="1983" box="[645,916,908,932]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B961EA93FF86CD67FD5FFC77FD1510CB" box="[645,767,909,932]" italics="true" pageId="5" pageNumber="399">T. spinipes</emphasis>
<bibRefCitation id="EF844B70FF86CD67FCD3FC77FC7E10CB" author="Bruce" box="[777,916,908,932]" pageId="5" pageNumber="399" refString="Bruce, A. J. (1983) Thor spinipes sp. nov., a new hippolytid shrimp from the Cobourg Peninsula, Northern Australia. The Beagle, Records of the Museums and Art Galleries of the Northern Territory Museum of Arts and Sciences, 1, 1 - 10." type="journal article" year="1983">Bruce, 1983</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C154D02FF86CD67FC0AFC77FB0110CA" authority="Boone, 1935" authorityName="Boone" authorityYear="1935" box="[976,1259,908,933]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B961EA93FF86CD67FC0AFC77FBA510CB" box="[976,1103,909,932]" italics="true" pageId="5" pageNumber="399">T. spinosus</emphasis>
<bibRefCitation id="EF844B70FF86CD67FB83FC77FB0110CA" author="Boone" box="[1113,1259,908,933]" pageId="5" pageNumber="399" refString="Boone, L. (1935) Crustacea: Anomura, Macrura, Euphausiacea, Isopoda, Amphipoda and Echinodermata: Asteroidea and Echinoidea. Scientific results of the world cruise of the yacht &quot; Alva, &quot; 1931, William K. Vanderbilt, Commanding. Bulletin of the Vanderbilt Marine Museum, 6, 1 - 264." type="journal article" year="1935">Boone, 1935</bibRefCitation>
</taxonomicName>
by the entirely lacking of a distinct supraorbital spine or its vestige.
</paragraph>
<paragraph id="8BAA3681FF86CD67FF1DFC2EFB0E175B" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">
The telson of
<taxonomicName id="4C154D02FF86CD67FEBEFC2EFD3F1083" authority="Holthuis, 1947" authorityName="Holthuis" authorityYear="1947" box="[356,725,980,1005]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B961EA93FF86CD67FEBEFC2EFDCC1082" box="[356,550,980,1005]" italics="true" pageId="5" pageNumber="399">Thor intermedius</emphasis>
<bibRefCitation id="EF844B70FF86CD67FDF7FC2EFD3F1083" author="Holthuis" box="[557,725,980,1005]" pageId="5" pageNumber="399" refString="Holthuis, L. B. (1947) The Decapoda of the Siboga Expedition. Part IX. The Hippolytidae and Rhynchocinetidae collected by the Siboga and Snellius expeditions with remarks on other species. Siboga Expeditie, 39 a 8, 1 - 100." type="journal article" year="1947">Holthuis, 1947</bibRefCitation>
</taxonomicName>
has only one pair of very minute spinules on the posterior half of the dorsal surface; the stylocerite of
<taxonomicName id="4C154D02FF86CD67FD91FC03FC9E177F" authority="Heller, 1862" authorityName="Heller" authorityYear="1862" box="[587,884,1015,1040]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="paschalis">
<emphasis id="B961EA93FF86CD67FD91FC03FD3A177F" box="[587,720,1015,1040]" italics="true" pageId="5" pageNumber="399">T. paschalis</emphasis>
(
<bibRefCitation id="EF844B70FF86CD67FD05FC0DFC86177F" author="Heller" box="[735,876,1015,1040]" pageId="5" pageNumber="399" refString="Heller, C. (1862) Beitrage zur Crustaceen-Fauna des rothen Meeres. II. Sitzungsberichte der Akademie der Wissenschaften in Wien, 44, 241 - 295." type="journal article" year="1862">Heller, 1862</bibRefCitation>
)
</taxonomicName>
totally lacks the little tooth at basal part of ourter margin (
<bibRefCitation id="EF844B70FF86CD67FF2EFBE6FE4B175B" author="Holthuis" box="[244,417,1052,1077]" pageId="5" pageNumber="399" refString="Holthuis, L. B. (1947) The Decapoda of the Siboga Expedition. Part IX. The Hippolytidae and Rhynchocinetidae collected by the Siboga and Snellius expeditions with remarks on other species. Siboga Expeditie, 39 a 8, 1 - 100." type="journal article" year="1947">Holthuis, 1947</bibRefCitation>
). These morphological differences make a distinction from
<taxonomicName id="4C154D02FF86CD67FBE6FBE7FB37175A" box="[1084,1245,1052,1077]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B961EA93FF86CD67FBE6FBE7FB37175A" box="[1084,1245,1052,1077]" italics="true" pageId="5" pageNumber="399">T. hainanensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAA3681FF86CD67FF1DFBC5FB0B1713" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">
The new species can be distinguished from
<taxonomicName id="4C154D02FF86CD67FD16FBC5FBFD1737" authority="Bruce, 1978" authorityName="Bruce" authorityYear="1978" box="[716,1047,1087,1112]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="marguitae">
<emphasis id="B961EA93FF86CD67FD16FBC5FC951737" box="[716,895,1087,1112]" italics="true" pageId="5" pageNumber="399">Thor marguitae</emphasis>
<bibRefCitation id="EF844B70FF86CD67FC50FBBAFBFD1737" author="Bruce" box="[906,1047,1088,1112]" pageId="5" pageNumber="399" refString="Bruce, A. J. (1978) Thor marguitae sp. nov., a new hippolytid shrimp from Heron Island, Australia. Crustaceana, 35, 160 - 169. http: // dx. doi. org / 10.1163 / 156854078 x 00060" type="journal article" year="1978">Bruce, 1978</bibRefCitation>
</taxonomicName>
by the rostrum with bifid tip. In ovigerous females,
<taxonomicName id="4C154D02FF86CD67FEABFB9FFE151713" box="[369,511,1125,1148]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="marguitae">
<emphasis id="B961EA93FF86CD67FEABFB9FFE151713" box="[369,511,1125,1148]" italics="true" pageId="5" pageNumber="399">T. marguitae</emphasis>
</taxonomicName>
has a more strongly developed rostrum, with typically a trifid tip.
</paragraph>
</subSubSection>
<subSubSection id="C30F650AFF86CD67FF1DFB7DFD6B1662" pageId="5" pageNumber="399" type="reference_group">
<paragraph id="8BAA3681FF86CD67FF1DFB7DFD6B1662" blockId="5.[151,1437,151,1293]" pageId="5" pageNumber="399">
<taxonomicName id="4C154D02FF86CD67FF1DFB7DFE7A17CF" box="[199,400,1159,1184]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="amboinensis">
<emphasis id="B961EA93FF86CD67FF1DFB7DFE7A17CF" box="[199,400,1159,1184]" italics="true" pageId="5" pageNumber="399">Thor amboinensis</emphasis>
</taxonomicName>
(de
<bibRefCitation id="EF844B70FF86CD67FE1BFB73FDD117CF" author="Man" box="[449,571,1160,1184]" pageId="5" pageNumber="399" refString="De Man, J. G. (1888) Bericht uber die von Herrn Dr. J. Brock im indischen Archipel gesammelten Decapoden und Stomatopoden. Archiv fur Naturgeschichte, 53, 215 - 600." type="journal article" year="1888">Man, 1888</bibRefCitation>
) is well-known by associated with all kinds of coelenterates and always has a distally acute simple rostrum, without any evidence of ventral teeth. Apart from the existence of epipod on the first pereopod,
<taxonomicName id="4C154D02FF86CD67FED7FB2BFE441787" box="[269,430,1231,1256]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B961EA93FF86CD67FED7FB2BFE441787" box="[269,430,1231,1256]" italics="true" pageId="5" pageNumber="399">T. hainanensis</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName id="4C154D02FF86CD67FCF2FB2BFC3A1787" box="[808,976,1231,1256]" class="Malacostraca" family="Hippolytidae" genus="Thor" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="399" phylum="Arthropoda" rank="species" species="amboinensis">
<emphasis id="B961EA93FF86CD67FCF2FB2BFC3A1787" box="[808,976,1231,1256]" italics="true" pageId="5" pageNumber="399">T. amboinensis</emphasis>
</taxonomicName>
by its smaller body and bifid tip rostrum (
<bibRefCitation id="EF844B70FF86CD67FF44FB0EFEA01663" author="Holthuis" box="[158,330,1268,1293]" pageId="5" pageNumber="399" refString="Holthuis, L. B. (1947) The Decapoda of the Siboga Expedition. Part IX. The Hippolytidae and Rhynchocinetidae collected by the Siboga and Snellius expeditions with remarks on other species. Siboga Expeditie, 39 a 8, 1 - 100." type="journal article" year="1947">Holthuis, 1947</bibRefCitation>
;
<bibRefCitation id="EF844B70FF86CD67FE8FFB0EFD9E1662" author="Miyake" box="[341,628,1268,1293]" pageId="5" pageNumber="399" refString="Miyake, S. &amp; Hayashi, K. I. (1966) Some hippolytid shrimp living in coral reefs of the West Pacific. Journal of the Faculty of Agriculture, Kyushu University, 14, 143 - 160." type="journal article" year="1966">Miyake &amp; Hayashi, 1966</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>