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<document id="6A989694179A8B78323D6939C0C05D9E" ID-DOI="10.5281/zenodo.175790" ID-GBIF-Dataset="2a6512b7-5bd9-4682-b5db-79c72373b68c" ID-ISSN="1175-5326" ID-Zenodo-Dep="175790" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459820893039" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03B9F24A7C17FF89FF3055B26096237F" docLanguage="en" docName="zt01431p044.pdf" docOrigin="Zootaxa 1431" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hydraena (Hydraenopsis) kodadai Freitag &amp; Jach, sp. n." docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="FF808A327C00FF90FFA75435641C2645" masterDocTitle="Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925" masterLastPageNumber="44" masterPageNumber="1" pageNumber="24" updateTime="1698221773297" updateUser="plazi">
<mods:mods id="971F24AA24AE5795D64F8FC81877488D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F3E4CD66C51E5CF0080AACEFE70ECA84">
<mods:title id="99863427E77547942AC76BB9E2C84B5C">Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925</mods:title>
</mods:titleInfo>
<mods:name id="84CAD4DE8051FCC844D17DF6049C15C6" type="personal">
<mods:role id="1FF6564CA75A9F3667DF530943DFEA30">
<mods:roleTerm id="E6BAE0988564595AC43D70BEB44E960A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02522EB6FBF4840943F3BBE3073B6177">Freitag, Hendrik</mods:namePart>
</mods:name>
<mods:name id="14AA918C097DD93538564294FFF3A3A5" type="personal">
<mods:role id="AAB463D131138559B9D0A669A296D44F">
<mods:roleTerm id="F7327E0A97751423D6BA18FBC3F6FF2A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="889E121F1A97674793A3F97128499447">Jäch, Manfred A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A604F67D641720980C6C86068DB22E98">text</mods:typeOfResource>
<mods:relatedItem id="DFECD0B4DA482ACABAEEC0BFD6E6103B" type="host">
<mods:titleInfo id="2BABBFF55EC45682053B9AB90989C228">
<mods:title id="90AE2005FC995391BA96B5CCFAFB7B5E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="08ADBD6A0399A88013B8906B55A5B1C7">
<mods:date id="B4C85BF0915EACCD2005BC0318167D5A">2007</mods:date>
<mods:detail id="F2F8C0D133FCF47A46C0D22FCE71442F" type="volume">
<mods:number id="306B8A13257F0DD8A86139938DF77CA2">1431</mods:number>
</mods:detail>
<mods:extent id="8EA1FCEE9AA9EC9DB9A06F3D29C430A5" unit="page">
<mods:start id="92649D57C295DFC666D675B92EF353E2">1</mods:start>
<mods:end id="1CFFD0E8602879BE6BC53F65F5C5EC59">44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E13B07D2598064D950F065917C91838E">journal article</mods:classification>
<mods:identifier id="5F14E6A334C1B81CAD054282D07C5471" type="DOI">10.5281/zenodo.175790</mods:identifier>
<mods:identifier id="3228B3DF3CF402D6DE12906C7F5730A1" type="GBIF-Dataset">2a6512b7-5bd9-4682-b5db-79c72373b68c</mods:identifier>
<mods:identifier id="284DDCE2F4DDEF8E6B7D03EEED1925AE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="26DDBD7C90F11143A6F9F4AB6ECE8173" type="Zenodo-Dep">175790</mods:identifier>
</mods:mods>
<treatment id="03B9F24A7C17FF89FF3055B26096237F" ID-DOI="http://doi.org/10.5281/zenodo.6248081" ID-GBIF-Taxon="119355656" ID-Zenodo-Dep="6248081" LSID="urn:lsid:plazi:treatment:03B9F24A7C17FF89FF3055B26096237F" httpUri="http://treatment.plazi.org/id/03B9F24A7C17FF89FF3055B26096237F" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="C30A10D77C17FF87FF3055B2654C2781" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BAF435C7C17FF87FF3055B2675027E4" blockId="23.[151,844,391,452]" box="[151,844,391,417]" pageId="23" pageNumber="24">
<heading id="D0E7F4307C17FF87FF3055B2675027E4" bold="true" box="[151,844,391,417]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B9649F4E7C17FF87FF3055B2675027E4" bold="true" box="[151,844,391,417]" pageId="23" pageNumber="24">
<taxonomicName id="4C1038DF7C17FF87FF3055B266E027E4" ID-CoL="3N3CD" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[151,764,391,417]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="kodadai" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B9649F4E7C17FF87FF3055B2650C27E4" bold="true" box="[151,272,391,417]" italics="true" pageId="23" pageNumber="24">Hydraena</emphasis>
(
<emphasis id="B9649F4E7C17FF87FE8755B265DA27E4" bold="true" box="[288,454,391,417]" italics="true" pageId="23" pageNumber="24">Hydraenopsis</emphasis>
)
<emphasis id="B9649F4E7C17FF87FE7055B2662B27E4" bold="true" box="[471,567,391,417]" italics="true" pageId="23" pageNumber="24">kodadai</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<taxonomicNameLabel id="A25722357C17FF87FCAE55B2675027E4" box="[777,844,391,417]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF435C7C17FF87FF30559F654C2781" blockId="23.[151,844,391,452]" box="[151,336,426,452]" pageId="23" pageNumber="24">(Figs. 9, 21a–h)</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C17FF87FF3055CF67EF23E9" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BAF435C7C17FF87FF3055CF64C02421" blockId="23.[151,1437,506,2012]" pageId="23" pageNumber="24">
<emphasis id="B9649F4E7C17FF87FF3055CF65212451" bold="true" box="[151,317,506,532]" pageId="23" pageNumber="24">
<typeStatus id="54ABFDFE7C17FF87FF3055CF64C82451" box="[151,212,506,532]" pageId="23" pageNumber="24">Type</typeStatus>
locality.
</emphasis>
Small stream in primary forest, ca.
<quantity id="4CE8EEB97C17FF87FD4355CF67322451" box="[740,814,506,532]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="100.0">100 m</quantity>
a.s.l., tributary of Cabayugan River, St. Paul Subterranean River National Park, Cabayugan district,
<quantity id="4CE8EEB97C17FF87FD76561767062479" box="[721,794,546,572]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="23" pageNumber="24" unit="km" value="40.0">40 km</quantity>
NNE of Puerto Princesa City, central Palawan,
<collectingCountry id="F30703CC7C17FF87FAEB561764C52421" name="Philippines" pageId="23" pageNumber="24">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BAF435C7C17FF87FF60564767EF23E9" blockId="23.[151,1437,506,2012]" pageId="23" pageNumber="24">
<emphasis id="B9649F4E7C17FF87FF60564765E624C9" bold="true" box="[199,506,626,652]" pageId="23" pageNumber="24">
<typeStatus id="54ABFDFE7C17FF87FF605647651824C9" box="[199,260,626,652]" pageId="23" pageNumber="24">Type</typeStatus>
material.
<typeStatus id="54ABFDFE7C17FF87FE2F564765E624C9" box="[392,506,626,652]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
ɗ (
<collectionCode id="ED01DB997C17FF87FD8E5647666724C9" box="[553,635,626,652]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
): “
<collectingCountry id="F30703CC7C17FF87FD3C5647673924C9" box="[667,805,626,652]" name="Philippines" pageId="23" pageNumber="24">Philippines</collectingCountry>
, PALAWAN centr. smal stream in primary forest ca.
<quantity id="4CE8EEB97C17FF87FF3056AF64FA24F1" box="[151,230,666,692]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="100.0">100 m</quantity>
, Cabayugan env.,
<date id="FFAE659C7C17FF87FE1856AF665424F1" box="[447,584,666,692]" pageId="23" pageNumber="24" value="1995-12-02">2. 12. 1995</date>
, Ján Kodada lgt.”, terminal parts of abdomen and aedeagus glued separately.
<emphasis id="B9649F4E7C17FF87FF4C56F7656E2499" bold="true" box="[235,370,706,732]" pageId="23" pageNumber="24">
<typeStatus id="54ABFDFE7C17FF87FF4C56F765712499" box="[235,365,706,732]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
:
</emphasis>
2 ΨΨ (1 ex.
<collectionCode id="ED01DB997C17FF87FDBA56F766742499" box="[541,616,706,732]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
, 1 ex.
<collectionCode id="ED01DB997C17FF87FD6656F7670E2499" box="[705,786,706,732]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="23" pageNumber="24">UPLB</collectionCode>
): same locality data as
<typeStatus id="54ABFDFE7C17FF87FB9156F760BC2499" box="[1078,1184,706,732]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
; 1 ɗ (
<collectionCode id="ED01DB997C17FF87FB5656F7615E2499" box="[1265,1346,706,732]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
), 4 ΨΨ (
<collectionCode id="ED01DB997C17FF87FF3856DF64ED2541" box="[159,241,746,772]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
): “
<collectingCountry id="F30703CC7C17FF87FEB756DF65862541" box="[272,410,746,772]" name="Philippines" pageId="23" pageNumber="24">Philippines</collectingCountry>
, PALAWAN centr., Ulangoan river,
<quantity id="4CE8EEB97C17FF87FCEF56DF678F2541" box="[840,915,746,772]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="24" unit="km" value="18.0">18 km</quantity>
NE San Rafael
<date id="FFAE659C7C17FF87FBF456DF60CB2541" box="[1107,1239,746,772]" pageId="23" pageNumber="24" value="1995-12-06">6.XII.1995</date>
, J. Kodada &amp; B. Rigová lgt.”; 2 ΨΨ (
<collectionCode id="ED01DB997C17FF87FE22572765CE2569" box="[389,466,786,812]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
, 1 ex. mutilated): “
<collectionCode id="ED01DB997C17FF87FD13572766E32569" box="[692,767,786,812]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, P. Princesa Sabang, Waterfall (upper,
<quantity id="4CE8EEB97C17FF87FA92572761752569" box="[1333,1385,786,812]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="24" unit="m" value="30.0">30m</quantity>
asl)
<geoCoordinate id="EE24259B7C17FF87FF30570F64EB2511" box="[151,247,826,852]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="10.216666">10°13'N</geoCoordinate>
<geoCoordinate id="EE24259B7C17FF87FEA6570F65702511" box="[257,364,826,852]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="925" value="118.88333">
118°53'
<geoCoordinate id="EE24259B7C17FF87FEFF570F65702511" box="[344,364,826,852]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5555" value="26.3">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C17FF87FED2570F65F12511" box="[373,493,826,852]" pageId="23" pageNumber="24" value="1995-03-26">26.3.1995</date>
, leg. Freitag (19a)”; 1 ɗ (
<collectionCode id="ED01DB997C17FF87FC83570F676A2511" box="[804,886,826,852]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="23" pageNumber="24">UPLB</collectionCode>
): “
<collectionCode id="ED01DB997C17FF87FC31570F67FD2511" box="[918,993,826,852]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.
<geoCoordinate id="EE24259B7C17FF87FF30575765372539" box="[151,299,866,892]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="10.2525">10°15.150'N</geoCoordinate>
<geoCoordinate id="EE24259B7C17FF87FE94575765CD2539" box="[307,465,866,892]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="118.967514">
118°58.051'
<geoCoordinate id="EE24259B7C17FF87FE19575765CD2539" box="[446,465,866,892]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5555" value="17.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C17FF87FE7E5757664D2539" box="[473,593,866,892]" pageId="23" pageNumber="24" value="2001-05-17">17.5.2001</date>
, leg. Freitag (
<date id="FFAE659C7C17FF87FD53575767022539" box="[756,798,866,892]" pageId="23" pageNumber="24" value="1925-01">i25</date>
)”; 1 ɗ (
<collectionCode id="ED01DB997C17FF87FCDD575767DB2539" box="[890,967,866,892]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
, mutilated), 1 Ψ (
<collectionCode id="ED01DB997C17FF87FB33575760FA2539" box="[1172,1254,866,892]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C17FF87FAA25757614C2539" box="[1285,1360,866,892]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.
<geoCoordinate id="EE24259B7C17FF87FD8C57BF66A325E1" box="[555,703,906,932]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="10.2525">10°15.150'N</geoCoordinate>
<geoCoordinate id="EE24259B7C17FF87FD6B57BF677725E1" box="[716,875,906,932]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="118.967514">
118°58.051'
<geoCoordinate id="EE24259B7C17FF87FCFF57BF677725E1" box="[856,875,906,932]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5555" value="17.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C17FF87FCD057BF67EC25E1" box="[887,1008,906,932]" pageId="23" pageNumber="24" value="2001-05-17">17.5.2001</date>
, leg. Freitag (e18)”; 1 ɗ (
<collectionCode id="ED01DB997C17FF87FA9957BF618C25E1" box="[1342,1424,906,932]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C17FF87FF05578764F12589" box="[162,237,946,972]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.
<geoCoordinate id="EE24259B7C17FF87FD12578767562589" box="[693,842,946,972]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="10.2525">10°15.150'N</geoCoordinate>
<geoCoordinate id="EE24259B7C17FF87FCF4578767EE2589" box="[851,1010,946,972]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="118.967514">
118°58.051'
<geoCoordinate id="EE24259B7C17FF87FC78578767EE2589" box="[991,1010,946,972]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5555" value="17.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C17FF87FC5C5787606F2589" box="[1019,1139,946,972]" pageId="23" pageNumber="24" value="2001-05-17">17.5.2001</date>
, leg. Freitag (e26)”; 1 ɗ (WPU): “
<collectionCode id="ED01DB997C17FF87FEAB57EF654B25B1" box="[268,343,986,1012]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.
<geoCoordinate id="EE24259B7C17FF87FC9F57EF67D025B1" box="[824,972,986,1012]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="10.2525">10°15.150'N</geoCoordinate>
<geoCoordinate id="EE24259B7C17FF87FC7D57EF606425B1" box="[986,1144,986,1012]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="118.967514">
118°58.051'
<geoCoordinate id="EE24259B7C17FF87FBC257EF606425B1" box="[1125,1144,986,1012]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="5555" value="17.5">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C17FF87FB2257EF60E225B1" box="[1157,1278,986,1012]" pageId="23" pageNumber="24" value="2001-05-17">17.5.2001</date>
, leg. Freitag (e29)”; 1 ɗ (
<collectionCode id="ED01DB997C17FF87FE9A503765932259" box="[317,399,1026,1052]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C17FF87FE14503765E22259" box="[435,510,1026,1052]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.PR1, 1.Drift
<geoCoordinate id="EE24259B7C17FF87FBD4503760E02259" box="[1139,1276,1026,1052]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="15" value="10.2525">10°15'09''N</geoCoordinate>
<geoCoordinate id="EE24259B7C17FF87FAAE503761802259" box="[1289,1436,1026,1052]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="15" value="118.9675">118°58'03''E</geoCoordinate>
<date id="FFAE659C7C17FF87FF30501F650E2201" box="[151,274,1066,1092]" pageId="23" pageNumber="24" value="2001-05-17">17.V.2001</date>
, leg. Freitag (a48)”; 1 Ψ (
<collectionCode id="ED01DB997C17FF87FDCD501F66A02201" box="[618,700,1066,1092]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C17FF87FD45501F67312201" box="[738,813,1066,1092]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, P. Princesa Panaguman R.PR1, 1.Drift
<geoCoordinate id="EE24259B7C17FF87FF305067653C2229" box="[151,288,1106,1132]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="15" value="10.2525">10°15'09''N</geoCoordinate>
<geoCoordinate id="EE24259B7C17FF87FE8C506765A22229" box="[299,446,1106,1132]" direction="east" orientation="longitude" pageId="23" pageNumber="24" precision="15" value="118.9675">118°58'03''E</geoCoordinate>
<date id="FFAE659C7C17FF87FE6E5067665F2229" box="[457,579,1106,1132]" pageId="23" pageNumber="24" value="2001-05-17">17.V.2001</date>
, leg. Freitag (a49)”; 1 ɗ (
<collectionCode id="ED01DB997C17FF87FC2F506767C62229" box="[904,986,1106,1132]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C17FF87FC5A506760542229" box="[1021,1096,1106,1132]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, P. Princesa SSW Maratarpi Cabayugan R1
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118°49'29’’
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<date id="FFAE659C7C17FF87FCA8504F678922D1" box="[783,917,1146,1172]" pageId="23" pageNumber="24" value="2000-11-12">11.12.2000</date>
, leg. Freitag (a7)”; 1 Ψ (WPU, mutilated): “
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.: Palawan, P. Princesa SSW Martarpi Cabayugan R.1
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118°49'29''
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, leg. Freitag (9)”; 1 ɗ, 1 Ψ (
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): “
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.: Palawan, P. Princesa SSW Maratarpi Cabayugan R1
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118°49'29’’
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<date id="FFAE659C7C17FF87FE9E50C765AD2349" box="[313,433,1266,1292]" pageId="23" pageNumber="24" value="2001-06-17">17.6.2001</date>
, leg. Freitag (e10)”; 1 ɗ (
<collectionCode id="ED01DB997C17FF87FD4050C7672A2349" box="[743,822,1266,1292]" pageId="23" pageNumber="24">IRML</collectionCode>
; aedeagus mutilated): “
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.: Palawan, P. Princesa SSW Martarpi Cabayugan R.1,
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<date id="FFAE659C7C17FF87FCFE512F67C02371" box="[857,988,1306,1332]" pageId="23" pageNumber="24" value="2001-07-01">1.VII.2001</date>
, leg. Freitag (a46)E”; 1 Ψ (
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): “
<collectionCode id="ED01DB997C17FF87FF05517764F12319" box="[162,237,1346,1372]" pageId="23" pageNumber="24">PHIL</collectionCode>
.: Palawan, Taytay; 1/3way highway to Lake
<taxonomicName id="4C1038DF7C17FF87FCA35177676A2319" box="[772,886,1346,1372]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
,
<taxonomicName id="4C1038DF7C17FF87FC24517767EE2319" box="[899,1010,1346,1372]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
Stream trib., prim.forest c.
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asl,
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119°31'
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</geoCoordinate>
<date id="FFAE659C7C17FF87FED6515F65F423C1" box="[369,488,1386,1412]" pageId="23" pageNumber="24" value="1995-04-29">29.4.1995</date>
, leg. Freitag (13b)M”; 1 Ψ (
<collectionCode id="ED01DB997C17FF87FC89515F676723C1" box="[814,891,1386,1412]" pageId="23" pageNumber="24">IRML</collectionCode>
) “
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.: Palawan, P. Princesa SSW Maratarpi Cabayugan R1
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118°49'29’’
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<date id="FFAE659C7C17FF87FD2451A7671423E9" box="[643,776,1426,1452]" pageId="23" pageNumber="24" value="2000-11-16">16.11.2000</date>
, leg. Freitag (e12)”.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C17FF89FF60518F673525EF" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24" type="description">
<paragraph id="8BAF435C7C17FF87FF60518F601D23B9" blockId="23.[151,1437,506,2012]" pageId="23" pageNumber="24">
<emphasis id="B9649F4E7C17FF87FF60518F65432391" bold="true" box="[199,351,1466,1492]" pageId="23" pageNumber="24">Description.</emphasis>
Body
<quantity id="4CE8EEB97C17FF87FE0A518F664B2391" box="[429,599,1466,1492]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="23" pageNumber="24" unit="mm" value="1.375" valueMax="1.45" valueMin="1.3">1.30–1.45 mm</quantity>
long. Habitus as in Fig. 9. Entire body reddish brown, sometimes goldbrown; frons slightly darker brown, legs and maxillary palpi slightly paler.
</paragraph>
<paragraph id="8BAF435C7C17FF87FF60523F610F20A9" blockId="23.[151,1437,506,2012]" pageId="23" pageNumber="24">Pronotum wider than long, broadest at about posterior 0.3, entire pronotum moderately densely punctate; punctures small and shallowly (posteriorly moderately deeply) impressed, interstices glabrous; foveae hardly perceptible, or absent; lateral margins anteriorly convergent, distinctly sinuously convergent to rectangular posterior angles, forming an almost rectangular pronoto-elytral angle; lateral rim denticulate; anterior margins concave, distinctly broader than straight posterior margin; lateral hypomeron approximately as broad as profemur, inner margin (hypomeral carina) slightly biconvex; mesal hypomeron narrow, imperceptible.</paragraph>
<paragraph id="8BAF435C7C17FF87FF6052CF67662199" blockId="23.[151,1437,506,2012]" pageId="23" pageNumber="24">Elytra elongately oval, apex with tiny excision next to sutural keel, with ca. 12 longitudinal, not very regular, unimpressed rows of punctures (seven between suture and shoulder); punctures small and shallowly (basally moderately deeply) impressed, sparsely arranged; interstices and intervals flat, glabrous; lateral gutter of elytra widely explanate, not reaching elytral apex; elytral rim posteriorly finely denticulate; pseudepipleuron anteriorly about as broad as metafemur, reaching posterior 0.2, with indistinct foveae and one inconspicuous row of punctures; epipleuron narrow, short, inconspicuous.</paragraph>
<caption id="DF6F13D47C18FF88FF3052B3611F2085" httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="24" pageNumber="25" targetBox="[274,1322,194,1644]" targetPageId="24">
<paragraph id="8BAF435C7C18FF88FF3052B3611F2085" blockId="24.[151,1436,1670,1728]" pageId="24" pageNumber="25">
<emphasis id="B9649F4E7C18FF88FF3052B3653020DB" bold="true" box="[151,300,1670,1694]" pageId="24" pageNumber="25">FIGURE 21.</emphasis>
<taxonomicName id="4C1038DF7C18FF88FE9F52B2661820DB" box="[312,516,1671,1694]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kodadai" status="sp. nov.">
<emphasis id="B9649F4E7C18FF88FE9F52B2661820DB" box="[312,516,1671,1694]" italics="true" pageId="24" pageNumber="25">Hydraena kodadai</emphasis>
</taxonomicName>
<emphasis id="B9649F4E7C18FF88FDB752B3664D20DB" bold="true" box="[528,593,1670,1694]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="A25722357C18FF88FDB752B3664D20DB" box="[528,593,1670,1694]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
a–c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f–g) spermatheca; h) female tergite X.
</paragraph>
</caption>
<paragraph id="8BAF435C7C18FF88FF6052C164FE21EB" blockId="24.[151,1436,1780,2006]" pageId="24" pageNumber="25">Mesoventrite with well developed longitudinal ridges; meso- and metaventrite, pubescent; abdominal ventrites sparsely pubescent; mesoventral intercoxal process moderately long, very wide; metaventral disc deeply impressed; metaventral plaques distinct, widely separated (wider than inflexed lateral portion of elytra), narrow, glabrous; intercoxal sternite large, approximately as broad as metafemora and as long as broad.</paragraph>
<paragraph id="8BAF435C7C18FF88FF60538960882193" blockId="24.[151,1436,1780,2006]" box="[199,1172,1980,2006]" pageId="24" pageNumber="25">Femora with inner margin straight to concave, outer margin conspicuously convex.</paragraph>
<paragraph id="8BAF435C7C19FF89FF6054AD60262717" blockId="25.[151,1437,152,1338]" pageId="25" pageNumber="26">
Aedeagus (
<figureCitation id="132B5FD97C19FF89FEED54AD65B526F7" box="[330,425,152,178]" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="25" pageNumber="26">Figs. 21</figureCitation>
a–c) slender, gently evenly curved ventrad and dextrad, with one dorsal subapical seta; dorso-ventrally flattened apically, appearing almost symmetric (ventral/dorsal view); phallobase asymmetric. Distal lobe not clearly delimited from main piece, median portion weakly sclerotized. Parameres articulately connected with main piece, moderately long; each paramere with a row of about 10 moderately long setae; left paramere inserted at about basal 0.50, right paramere at basal 0.55 of aedeagus.
</paragraph>
<paragraph id="8BAF435C7C19FF89FF6055556726278F" blockId="25.[151,1437,152,1338]" pageId="25" pageNumber="26">
Gonocoxite (
<figureCitation id="132B5FD97C19FF89FEC5555565A6273F" box="[354,442,352,378]" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="25" pageNumber="26">Fig. 21</figureCitation>
e) subsemioval; apical area of ventral plate almost as long as basal area; subapical tufts curved; apical margin of basal area with median projection; condyles very small; dorsal plate very simple, slightly surpassing outer plate basally, without cavea.
</paragraph>
<paragraph id="8BAF435C7C19FF89FF6055ED616027B7" blockId="25.[151,1437,152,1338]" box="[199,1404,472,498]" pageId="25" pageNumber="26">
Spermatheca (
<figureCitation id="132B5FD97C19FF89FEC955ED65D127B7" box="[366,461,472,498]" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="25" pageNumber="26">
Figs.
<quantity id="4CE8EEB97C19FF89FE1755ED65D127B7" box="[432,461,472,498]" metricMagnitude="-2" metricUnit="kg" metricValue="2.1" pageId="25" pageNumber="26" unit="g" value="21.0">21</quantity>
</figureCitation>
g–f). Proximal portion extremely large, strongly curved; distal portion discoidal.
</paragraph>
<paragraph id="8BAF435C7C19FF89FF60563560BD24A7" blockId="25.[151,1437,152,1338]" pageId="25" pageNumber="26">
Secondary sexual characters. Male usually slightly larger than female. Meso- and metatibia in male slightly curved. Male profemur with (rather inconspicuous) ventro-basal ledge. Male terminal sternite (
<figureCitation id="132B5FD97C19FF89FACA561D64A8242F" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="25" pageNumber="26">Fig. 21</figureCitation>
d) oblongly subhexagonal, apical margin rather straight, base broad, distinctly produced into lateral appendages; spiculum gastrale about twice as long as terminal sternite, almost straight. Male tergite X distinctly excised apically. Female tergite X (
<figureCitation id="132B5FD97C19FF89FD06569566E524FF" box="[673,761,672,698]" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="25" pageNumber="26">Fig. 21</figureCitation>
h) subsemicircular, with rather few long discal trichoid setae; subapical fringe with moderately long trichoid setae; hyaline apical margin entire.
</paragraph>
<paragraph id="8BAF435C7C19FF89FF6056DA673525EF" blockId="25.[151,1437,152,1338]" pageId="25" pageNumber="26">
<emphasis id="B9649F4E7C19FF89FF6056DA65C2254C" bold="true" box="[199,478,751,777]" pageId="25" pageNumber="26">Differential Diagnosis.</emphasis>
<taxonomicName id="4C1038DF7C19FF89FE4256C566DD254C" box="[485,705,752,777]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kodadai">
<emphasis id="B9649F4E7C19FF89FE4256C566DD254C" box="[485,705,752,777]" italics="true" pageId="25" pageNumber="26">Hydraena kodadai</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C1038DF7C19FF89FCE356C567B1254C" box="[836,941,752,777]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="zetteli">
<emphasis id="B9649F4E7C19FF89FCE356C567B1254C" box="[836,941,752,777]" italics="true" pageId="25" pageNumber="26">H. zetteli</emphasis>
</taxonomicName>
in size, colour and general habitus. Externally,
<taxonomicName id="4C1038DF7C19FF89FF79572D657C2574" box="[222,352,792,817]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kodadai">
<emphasis id="B9649F4E7C19FF89FF79572D657C2574" box="[222,352,792,817]" italics="true" pageId="25" pageNumber="26">H. kodadai</emphasis>
</taxonomicName>
can be distinguished from the latter by its narrower explanate elytral gutter, which ends near apical 0.2 of elytra. Furthermore,
<taxonomicName id="4C1038DF7C19FF89FD83577566BA251C" box="[548,678,832,857]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kodadai">
<emphasis id="B9649F4E7C19FF89FD83577566BA251C" box="[548,678,832,857]" italics="true" pageId="25" pageNumber="26">H. kodadai</emphasis>
</taxonomicName>
can easily be recognized by its posteriorly distinctly convergent pronotum, number of elytral striae (usually 12) and its broad, subsymmetric, bilobed and dorso-ventrally flattened aedeagus, and by its extremely large spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C19FF89FF605782679522AF" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BAF435C7C19FF89FF60578261952597" blockId="25.[151,1437,152,1338]" box="[199,1417,951,978]" pageId="25" pageNumber="26">
<emphasis id="B9649F4E7C19FF89FF605782657C2594" bold="true" box="[199,352,951,977]" pageId="25" pageNumber="26">Distribution</emphasis>
(
<figureCitation id="132B5FD97C19FF89FED7578D65DB2597" box="[368,455,952,978]" captionStart="FIGURE 31" captionStartId="42.[151,255,1170,1194]" captionTargetBox="[267,1314,194,1138]" captionTargetId="figure@42.[264,1323,194,1146]" captionTargetPageId="42" captionText="FIGURE 31. Collecting sites and geographical distribution of Hydraena species in Palawan and Busuanga, including enlarged map of St. Paul National Park: Hydraena palawanensis sp. n., H. pseudopalawanensis sp. n., H. spp. ΨΨ cf. palawanensis / pseudopalawanensis, Hydraena sp. 1, H. manguao sp. n., H. hosiwergi sp. n., H. busuanga sp. n., Hydraena sp. 2, Hydraena sp. 3." httpUri="https://zenodo.org/record/175809/files/figure.png" pageId="25" pageNumber="26">Fig. 31</figureCitation>
).
<taxonomicName id="4C1038DF7C19FF89FE7B578D66AB2594" box="[476,695,952,977]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kodadai">
<emphasis id="B9649F4E7C19FF89FE7B578D66AB2594" box="[476,695,952,977]" italics="true" pageId="25" pageNumber="26">Hydraena kodadai</emphasis>
</taxonomicName>
is known from four small stream systems in central Palawan.
</paragraph>
<paragraph id="8BAF435C7C19FF89FF6057EA679522AF" blockId="25.[151,1437,152,1338]" pageId="25" pageNumber="26">
<emphasis id="B9649F4E7C19FF89FF6057EA652C25BC" bold="true" box="[199,304,991,1017]" pageId="25" pageNumber="26">Ecology.</emphasis>
All known macro-habitats are rather undisturbed streams (first to fourth order) surrounded by forest, however, with a remarkable variation in physicochemical conditions. PR1: acidic pH (6.32–7.02), low electrical conductivity (84 µS/cm on average), CR1: alkaline water (pH 8.07–8.45), high conductivity (435 µS/cm on average) due to ultra-basic subsoils (
<bibRefCitation id="EF813EAD7C19FF89FD6A506D67702237" author="Freitag" box="[717,876,1112,1138]" pageId="25" pageNumber="43" refString="Freitag, H. (2004) Composition and longitudinal patterns of aquatic insect emergence in small rivers of Palawan Island, the Philippines. International Revue of Hydrobiology, 89, 375 - 391." type="journal article" year="2004">Freitag 2004</bibRefCitation>
). Besides numerous specimens trapped in drift nets, some were collected repeatedly in gravel substrates under a small fall and in moderately fast flowing sections. Obviously, this species prefers (or at least tolerates) higher velocity. However, it was collected several times by emergence traps positioned over a pool section as well.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C19FF89FF6050C26096237F" pageId="25" pageNumber="26" type="etymology">
<paragraph id="8BAF435C7C19FF89FF6050C26096237F" blockId="25.[151,1437,152,1338]" pageId="25" pageNumber="26">
<emphasis id="B9649F4E7C19FF89FF6050C2654F2354" bold="true" box="[199,339,1271,1297]" pageId="25" pageNumber="26">Etymology.</emphasis>
Named for Dr. Ján Kodada (Comenius University, Bratislava,
<collectingCountry id="F30703CC7C19FF89FB9650CD60802357" box="[1073,1180,1272,1298]" name="Slovakia" pageId="25" pageNumber="26">Slovakia</collectingCountry>
), excellent water beetle specialist, who considerably contributed to the knowledge of the fauna of Palawan.
</paragraph>
</subSubSection>
</treatment>
</document>
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