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<document id="37E97AE8EC7E5B080893E18CC7CA870B" ID-DOI="10.1206/3961.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5370399" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268866948" checkinUser="carolina" docAuthor="Sinclair, Bradley J. &amp; Grimaldi, David A." docDate="2020" docId="03B94C5BFFDC3C5667C037C0F8F97616" docLanguage="en" docName="N3961.pdf" docOrigin="American Museum Novitates 2020 (3961)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3961/3961.1/Cretaceous-Diversity-of-the-Relict-Genus-Alavesia-Waters-and-Arillo/10.1206/3961.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Alavesia latala Sinclair &amp; Grimaldi 2020, new species" docType="treatment" docVersion="4" lastPageNumber="17" masterDocId="FF803423FFCC3C46662D3721FC617321" masterDocTitle="Cretaceous diversity of the relict genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="17" updateTime="1699205599378" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6331D6641FC4A9915AC8E5CB6FE8E8E9">Cretaceous diversity of the relict genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae)</mods:title>
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<mods:name id="5B49D70BC9F3924DA557269F4E092250" type="personal">
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<mods:namePart id="443E31B6C20B4B3EA19CE8476EF7176F">Sinclair, Bradley J.</mods:namePart>
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<mods:namePart id="2BC22C54C4AC4EDCFE1245D7EAF83063">Grimaldi, David A.</mods:namePart>
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<mods:title id="7387F12AF8660BFBA1BE21819EA1121E">American Museum Novitates</mods:title>
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<mods:date id="346A97D56A0C063D9013EFA786AE7E89">2020</mods:date>
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<mods:number id="37DC1100BB91E5D121D74B69E21C6278">2020-10-06</mods:number>
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<mods:url id="BC7DD0C6F44023EE8E43B9CE147011E9">https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3961/3961.1/Cretaceous-Diversity-of-the-Relict-Genus-Alavesia-Waters-and-Arillo/10.1206/3961.1.full</mods:url>
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<treatment id="03B94C5BFFDC3C5667C037C0F8F97616" ID-DOI="http://doi.org/10.5281/zenodo.5462037" ID-Zenodo-Dep="5462037" LSID="urn:lsid:plazi:treatment:03B94C5BFFDC3C5667C037C0F8F97616" httpUri="http://treatment.plazi.org/id/03B94C5BFFDC3C5667C037C0F8F97616" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="C30AAEC6FFDC3C5667C037C0FF5A73DA" box="[493,827,225,252]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BAFFD4DFFDC3C5667C037C0FF5A73DA" blockId="16.[493,827,225,252]" box="[493,827,225,252]" pageId="16" pageNumber="17">
<heading id="D0E74A21FFDC3C5667C037C0FF5A73DA" box="[493,827,225,252]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<emphasis id="B964215FFFDC3C5667C037C0FEC973DA" bold="true" box="[493,680,225,252]" pageId="16" pageNumber="17">
<taxonomicName id="4C1086CEFFDC3C5667C037C0FEC073DD" ID-CoL="89WYJ" authority="Sinclair &amp; Grimaldi, 2020" authorityName="Sinclair &amp; Grimaldi" authorityYear="2020" box="[493,673,225,252]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="latala" status="sp. nov.">
<emphasis id="B964215FFFDC3C5667C037C0FEC073DD" bold="true" box="[493,673,225,252]" italics="true" pageId="16" pageNumber="17">Alavesia latala</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2579C24FFDC3C56649D37C3FF5A73DA" box="[688,827,225,252]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFDC3C56640F3638FF677212" box="[546,774,281,307]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BAFFD4DFFDC3C56640F3638FF677212" blockId="16.[546,774,281,307]" box="[546,774,281,307]" pageId="16" pageNumber="17">
<heading id="D0E74A21FFDC3C56640F3638FF677212" box="[546,774,281,307]" centered="true" fontSize="36" level="3" pageId="16" pageNumber="17" reason="7">
<figureCitation id="132BE1C8FFDC3C56640F3638FEFD7212" box="[546,668,281,307]" captionStart="FIGURE" captionStartId="6.[108,197,1285,1306]" captionTargetBox="[200,1125,232,1256]" captionTargetId="figure-108@6.[108,1212,224,1262]" captionTargetPageId="6" captionText="FIGURE. 1. Antennae of Burmese amber Alavesia species, lateral views. All are males except N, O, P. Some antennae were reversed in direction for optimal comparison; longicornuta and longistylata are slightly reconstructed to account for curvature of the stylus. All are to the same scale. Specimen numbers: A. USNM PAL 726867. B. AMNH Bu-494. C. AMNH Bu-496. D. USNM PAL 726868. E. AMNH Bu-KL2-7. F. AMNH Bu-1476. G. JZC Bu-556 (AMNH). H. USNM PAL 726872. K. USNM PAL 726869. L. AMNH Bu-KL7-18. M. JZC Bu-282. N. Undetermined species. O. A. zigrasi. P. A. zigrasi." figureDoi="http://doi.org/10.5281/zenodo.5370401" httpUri="https://zenodo.org/record/5370401/files/figure.png" pageId="16" pageNumber="17">Figures 1F</figureCitation>
,
<figureCitation id="132BE1C8FFDC3C5664843638FEAB7212" box="[681,714,281,307]" captionStart="FIGURE 2" captionStartId="7.[108,194,1456,1477]" captionTargetBox="[121,1197,229,1434]" captionTargetId="figure-45@7.[108,1212,224,1437]" captionTargetPageId="7" captionText="FIGURE 2. Wings of six species of Alavesia in Burmese amber. All are males, to the same scale. Specimens: A. USNM PAL 726867. B. AMNH Bu-494. C. AMNH Bu-496. D. USNM PAL 726868. E. AMNH Bu-1476. F. JZC Bu-556." figureDoi="http://doi.org/10.5281/zenodo.5370403" httpUri="https://zenodo.org/record/5370403/files/figure.png" pageId="16" pageNumber="17">2E</figureCitation>
,
<figureCitation id="132BE1C8FFDC3C5664FA3638FF677212" box="[727,774,281,307]" captionStart="FIGURE 13" captionStartId="22.[108,194,1544,1565]" captionTargetBox="[108,1212,224,1524]" captionTargetId="figure-50@22.[108,1212,224,1527]" captionTargetPageId="22" captionText="FIGURE 13. Alavesia spp. in Burmese amber. A. A. lanceolata, USNM PAL 726868. B. A. pankowskiorum, USNM PAL 726869. C. A. latala, holotype, AMNH Bu-1476. D. A. longicornuta, holotype, JZC Bu-556" figureDoi="http://doi.org/10.5281/zenodo.5370425" httpUri="https://zenodo.org/record/5370425/files/figure.png" pageId="16" pageNumber="17">13C</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFDC3C5666883670FEA27297" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BAFFD4DFFDC3C5666883670FEA27297" blockId="16.[108,1215,337,1335]" pageId="16" pageNumber="17">
<materialsCitation id="3B78F710FFDC3C5666883670FEDE7297" collectionCode="R" longitude="-2.48" pageId="16" pageNumber="17" specimenCode="R4" specimenCount="1">
DIAGNOSIS: Wing short, broad, L/
<geoCoordinate id="EE249B8AFFDC3C5664143670FEF1724D" box="[569,656,337,364]" degrees="2.15" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-2.15">W 2.15</geoCoordinate>
, apex broadly rounded (fig. 13C);
<specimenCode id="DBB65536FFDC3C5662063673F8297251" box="[1067,1096,338,368]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="17">
R
<subScript id="1794FF08FFDC3C566213367EF8297251" attach="left" box="[1086,1096,351,368]" fontSize="7" pageId="16" pageNumber="17">4</subScript>
</specimenCode>
+5 and M
<subScript id="1794FF08FFDC3C56629E367EF8DD7251" attach="left" box="[1203,1212,351,368]" fontSize="7" pageId="16" pageNumber="17">1</subScript>
parallel for entire length; apex of CuA strongly recurved, anal lobe absent. Postpedicel triangular, L/
<geoCoordinate id="EE249B8AFFDC3C5666FD36BDFD477297" box="[208,294,412,438]" degrees="2.48" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-2.48">W 2.48</geoCoordinate>
, arista 0.54
<emphasis id="B964215FFFDC3C56678136BDFDDC7297" bold="true" box="[428,445,412,438]" pageId="16" pageNumber="17">×</emphasis>
length of postpedicel
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFDC3C56668E36E0FC8C7780" pageId="16" pageNumber="17" type="description">
<paragraph id="8BAFFD4DFFDC3C56668E36E0FFDA7107" blockId="16.[108,1215,337,1335]" pageId="16" pageNumber="17">
DESCRIPTION:
<emphasis id="B964215FFFDC3C56677D36E0FDEB72FA" box="[336,394,449,475]" italics="true" pageId="16" pageNumber="17">Male</emphasis>
.
<emphasis id="B964215FFFDC3C5667B536E0FE7E72FA" bold="true" box="[408,543,449,475]" pageId="16" pageNumber="17">Coloration</emphasis>
not preserved.
<emphasis id="B964215FFFDC3C5664F936E0FF7C72FA" bold="true" box="[724,797,449,475]" pageId="16" pageNumber="17">Head:</emphasis>
Scape short, setose; pedicel slightly larger, quadrate, postpedicel an elongate triangle, lanceolate (fig. 1F), L/
<geoCoordinate id="EE249B8AFFDC3C56658736C6F8617120" box="[938,1024,487,513]" degrees="2.48" direction="west" orientation="longitude" pageId="16" pageNumber="17" precision="555" value="-2.48">W 2.48</geoCoordinate>
; stylus relatively short, thick, 0.54
<emphasis id="B964215FFFDC3C56671E352DFD257107" bold="true" box="[307,324,524,550]" pageId="16" pageNumber="17">×</emphasis>
length of postpedicel, with two minute basal articles.
</paragraph>
<paragraph id="8BAFFD4DFFDC3C56668E3510FF9A719A" blockId="16.[108,1215,337,1335]" pageId="16" pageNumber="17">
<emphasis id="B964215FFFDC3C56668E3510FD63716A" bold="true" box="[163,258,561,587]" pageId="16" pageNumber="17">Thorax:</emphasis>
Macrosetae thickened; acrostichals not fully visible; dorsocentrals ending in stout pair of long prescutellars; postpronotal lobe with 3 short setae; notopleurals not fully visible, but 3 well-developed ones visible; 1 postsutural supraalar seta, 1 postalar seta; 3 pairs scutellar setae, apical pair upturned and crossed, lengths ~2
<emphasis id="B964215FFFDC3C5664EE3580FEB5719A" bold="true" box="[707,724,673,699]" pageId="16" pageNumber="17">×</emphasis>
that of lateral-most pair.
</paragraph>
<paragraph id="8BAFFD4DFFDC3C56668B35E6FCA57057" blockId="16.[108,1215,337,1335]" pageId="16" pageNumber="17">
<emphasis id="B964215FFFDC3C56668B35E6FC8471C0" bold="true" box="[166,229,711,737]" pageId="16" pageNumber="17">Legs:</emphasis>
Forecoxa with dense setae on anterior surface. Forefemur with some fine ventral spinules, but complete distribution not visible; mid and hind femora with spinelike setae near apex only. Tibiae with long apical, ventral seta(e): 1-2-2. Hind tibia with dorsal row of ~6 thicker setae, lengths about half width of tibia; posterior surface with short brush of fine setulae.
</paragraph>
<paragraph id="8BAFFD4DFFDC3C56668E34A0FD607710" blockId="16.[108,1215,337,1335]" pageId="16" pageNumber="17">
<emphasis id="B964215FFFDC3C56668E34A0FC8C70BA" bold="true" box="[163,237,897,923]" pageId="16" pageNumber="17">Wing:</emphasis>
Length
<quantity id="4CE850A8FFDC3C56676134A0FDD270BD" box="[332,435,897,924]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="16" pageNumber="17" unit="mm" value="1.55">1.55 mm</quantity>
(fig. 2E), vein C terminating slightly beyond M
<subScript id="1794FF08FFDC3C5665F534AEFF977081" attach="left" box="[984,1014,911,928]" fontSize="7" pageId="16" pageNumber="17">1+2</subScript>
; Sc nearly reaching costal margin; R
<subScript id="1794FF08FFDC3C5667703495FD1A70E4" attach="left" box="[349,379,948,965]" fontSize="7" pageId="16" pageNumber="17">4+5</subScript>
perfectly parallel to M
<subScript id="1794FF08FFDC3C5664A03495FECA70E4" attach="left" box="[653,683,948,965]" fontSize="7" pageId="16" pageNumber="17">1+2</subScript>
; crossvein r-m at middle of cell dm; apex of M
<subScript id="1794FF08FFDC3C5666AB34FBFCEE70CA" attach="left" box="[134,143,986,1003]" fontSize="7" pageId="16" pageNumber="17">4</subScript>
equidistant between M
<subScript id="1794FF08FFDC3C56678434FBFDA670CA" attach="left" box="[425,455,986,1003]" fontSize="7" pageId="16" pageNumber="17">1+2</subScript>
and CuA+CuP; CuP recurved, in line with bm-cu; apex of cell cua truncate; cell cua shorter than cell bm; vein A well developed but incomplete; anal lobe very narrow.
</paragraph>
<paragraph id="8BAFFD4DFFDC3C566688331DFC8C7780" blockId="16.[108,1215,337,1335]" pageId="16" pageNumber="17">
<emphasis id="B964215FFFDC3C566688331DFD477777" bold="true" box="[165,294,1084,1110]" pageId="16" pageNumber="17">Abdomen:</emphasis>
About same length as thorax; setation mostly not visible. Terminalia mostly not visible, but ventral margins of epandrium with fringe of long setae; surstyli very slender, digitiform.
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFDC3C566688338CF81F7630" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BAFFD4DFFDC3C566688338CF81F7630" blockId="16.[108,1215,337,1335]" pageId="16" pageNumber="17">
<materialsCitation id="3B78F710FFDC3C566688338CF81F7630" collectionCode="TYPE, MATERIAL, AMNH" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED016588FFDC3C566688338CFD0477E7" box="[165,357,1196,1223]" pageId="16" pageNumber="17">TYPE MATERIAL</collectionCode>
:
<typeStatus id="54AB43EFFFDC3C566759338DFD8277E7" box="[372,483,1196,1222]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
, male:
<collectionCode id="ED016588FFDC3C566414338DFEF577E7" box="[569,660,1196,1222]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="17">AMNH</collectionCode>
Bu-1476. The fly is complete but preserved in turbid amber (fig. 13C), which obscures some details (i.e., of genitalia). Despite the preservation, the distinctive wing shape and venation allowed a reliable diagnosis of this species.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30AAEC6FFDC3C566689323DF8F97616" box="[164,1176,1308,1335]" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BAFFD4DFFDC3C566689323DF8F97616" blockId="16.[108,1215,337,1335]" box="[164,1176,1308,1335]" pageId="16" pageNumber="17">
ETYMOLOGY: In reference to the broad wing, from Latin
<emphasis id="B964215FFFDC3C566568323DFF1D7617" box="[837,892,1308,1334]" italics="true" pageId="16" pageNumber="17">latus</emphasis>
(broad) and
<emphasis id="B964215FFFDC3C566234323DF85C7617" box="[1049,1085,1308,1334]" italics="true" pageId="16" pageNumber="17">ala</emphasis>
(wing).
</paragraph>
</subSubSection>
</treatment>
</document>